treatments-xml/data/03/DA/87/03DA87E60F35FFC4FF19B9633140F87E.xml
2024-06-21 12:22:17 +02:00

432 lines
76 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="D2A04418A4A4E708C92C47BC8AA8E332" ID-CLB-Dataset="25918" ID-DOI="10.11646/zootaxa.4659.1.1" ID-GBIF-Dataset="0cbe0b8e-2019-4f5a-8336-2d4d9fcd8bf6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3377239" ID-ZooBank="57309E14-0330-4ED7-BCDA-355EE6618215" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566808762347" checkinUser="plazi" docAuthor="Whitworth, Terry L. &amp; Yusseff-Vanegas, Sohath" docDate="2019" docId="03DA87E60F35FFC4FF19B9633140F87E" docLanguage="en" docName="zootaxa.4659.1.1.pdf" docOrigin="Zootaxa 4659 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Mesembrinellidae Shannon 1926" docType="treatment" docVersion="17" lastPageNumber="18" masterDocId="FFE3FF9E0F3AFFD5FF8EBA27342CFFB3" masterDocTitle="A revision of the genera and species of the Neotropical family Mesembrinellidae (Diptera: Oestroidea)" masterLastPageNumber="146" masterPageNumber="1" pageNumber="16" updateTime="1698752698920" updateUser="ExternalLinkService">
<mods:mods id="7D8561E348C206782B909DBFA4B64C12" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B7CD942C675DC31E3C25FA178B66DF12">
<mods:title id="A88D7206CE365F7BF36DBF4F932D8273">A revision of the genera and species of the Neotropical family Mesembrinellidae (Diptera: Oestroidea)</mods:title>
</mods:titleInfo>
<mods:name id="6E040F15F89033D915344D2D64903C4A" type="personal">
<mods:role id="D63068788EB144C52E03033D02457105">
<mods:roleTerm id="7BD1C8FEFC862D9FD5A940B404054DA2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="477DB00F82109066893F86FE519A6190">Whitworth, Terry L.</mods:namePart>
</mods:name>
<mods:name id="6CD4DA54A43DFD2D5C46FD39C563DEB9" type="personal">
<mods:role id="253B1B7B50176DCB3DFA66FFFCD6847B">
<mods:roleTerm id="7AA5909A957AA4B974FABD3D24DF5087">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E5B530F99AEF83140C7FC01BF08AEFE">Yusseff-Vanegas, Sohath</mods:namePart>
</mods:name>
<mods:typeOfResource id="A8CC3BED459E54F645DF647633D50934">text</mods:typeOfResource>
<mods:relatedItem id="A8C278423E8B8E728658667C85BA40F0" type="host">
<mods:titleInfo id="199AA6CEEB6763F12E49A1909F97DC1E">
<mods:title id="915DF7FBA44EFD3D6874F0FC88AF633E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="78C5512AACAB02DCC4E343713F67BB06">
<mods:date id="4897C4A14419E0AD7883128E1B621008">2019</mods:date>
<mods:detail id="98E7C7B2ED0533B7E1E6AB025EAF4154" type="pubDate">
<mods:number id="B81970EC0396DE494CB01A2C0D2F955B">2019-08-26</mods:number>
</mods:detail>
<mods:detail id="A0C9B067E276BFE0E26782823861EA90" type="volume">
<mods:number id="2EDFE47C1E6F9860149CCC26AD328396">4659</mods:number>
</mods:detail>
<mods:detail id="6582657BD4FD308991B02351589404ED" type="issue">
<mods:number id="70A6A335A5447BD302A47A9DC6DF386D">1</mods:number>
</mods:detail>
<mods:extent id="CE6E3768AE1B0A0865B55348D956B108" unit="page">
<mods:start id="B5A6E05CD705C93CD6BE61F72C477799">1</mods:start>
<mods:end id="5FF7AF50BD55CB18FB974295AFA77FC2">146</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="57ADD677D9D47A7B17EA525BA316C21D">journal article</mods:classification>
<mods:identifier id="4A306AA6D6A570F60140F3157DB57F3D" type="CLB-Dataset">25918</mods:identifier>
<mods:identifier id="297E1CFB9847E5A6C934EF3049415D41" type="DOI">10.11646/zootaxa.4659.1.1</mods:identifier>
<mods:identifier id="984869152FDC8165F91072369F45C805" type="GBIF-Dataset">0cbe0b8e-2019-4f5a-8336-2d4d9fcd8bf6</mods:identifier>
<mods:identifier id="B58EF4AEC82C3E62A74C0DA3C66F3A23" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5DCBE5A8A1FEDB9484A18E5F654FDB56" type="Zenodo-Dep">3377239</mods:identifier>
<mods:identifier id="E115C3DC5FBD56973BB4FB2CDF0C2EF3" type="ZooBank">57309E14-0330-4ED7-BCDA-355EE6618215</mods:identifier>
</mods:mods>
<treatment id="03DA87E60F35FFC4FF19B9633140F87E" ID-GBIF-Taxon="159243665" LSID="urn:lsid:plazi:treatment:03DA87E60F35FFC4FF19B9633140F87E" httpUri="http://treatment.plazi.org/id/03DA87E60F35FFC4FF19B9633140F87E" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="C369657B0F35FFDAFF19B96336BBFCED" box="[151,663,836,862]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BCC36F00F35FFDAFF19B96336BBFCED" blockId="15.[151,663,836,862]" box="[151,663,836,862]" pageId="15" pageNumber="16">
<heading id="D084819C0F35FFDAFF19B96336BBFCED" bold="true" box="[151,663,836,862]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B907EAE20F35FFDAFF19B96336BBFCED" bold="true" box="[151,663,836,862]" pageId="15" pageNumber="16">
Family
<taxonomicName id="4C734D730F35FFDAFF79B96336BBFCED" ID-CoL="89NT9" authority="Shannon, 1926" authorityName="Shannon" authorityYear="1926" box="[247,663,836,862]" class="Insecta" family="Mesembrinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">
Mesembrinellidae
<bibRefCitation id="EFE24B010F35FFDAFE51B96336BBFCED" author="Shannon, R. C." box="[479,663,836,862]" pageId="15" pageNumber="16" pagination="115 - 139" refId="ref109567" refString="Shannon, R. C. (1926) Synopsis of the American Calliphoridae (Diptera). Proceedings of the Entomological Society of Washington, 28, 115 - 139." type="journal article" year="1926">Shannon, 1926</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369657B0F35FFC4FF19B9AC3140F87E" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BCC36F00F35FFDAFF19B9AC3047F808" blockId="15.[151,1437,907,1979]" pageId="15" pageNumber="16">
<emphasis id="B907EAE20F35FFDAFF19B9AC353EFC15" bold="true" box="[151,274,907,934]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
This is a morphologically diverse family with species ranging from
<quantity id="4C8B9B150F35FFDAFBACB9AA3084FC14" box="[1058,1192,908,935]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.7" metricValueMin="0.7" pageId="15" pageNumber="16" unit="mm" value="12.0" valueMax="17.0" valueMin="7.0">7 to 17 mm</quantity>
in body length, sharing many characters with
<taxonomicName id="4C734D730F35FFDAFE30B9973676FC79" authorityName="Brauer &amp; von Bergenstamm" authorityYear="1889" box="[446,602,944,970]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Calliphoridae</taxonomicName>
. Thorax usually subshiny, blue, tan or brown, sometimes with pale to tan tomentose stripes. Abdomen often with shiny blue to brown cuticle with pale tomentum. Anterior thoracic spiracle normally ovate, with a broad oval opening above (
<figureCitation id="13482A750F35FFDAFD41B9DF3738FBA1" box="[719,788,1015,1042]" captionStart="FIGURES 18" captionStartId="16.[151,264,1826,1851]" captionTargetBox="[183,1404,184,1807]" captionTargetId="figure@16.[183,1404,166,1807]" captionTargetPageId="16" captionText="FIGURES 18. Diagnostic characters of Mesembrinellidae and Calliphoridae. 1. Wing of Mesembrinella flavicrura Aldrich, showing veins and cells. 2. Wing of M. currani Guimar„es, showing infuscated anterior margin. 3. Wing of Laneella fusconitida sp. nov., showing dark macula around crossvein r-m. 4. Wing of Calliphora vomitoria (Linnaeus), showing acute bend of vein M. 5. Anterior spiracle of Mesembrinella batesi Aldrich, showing broad oval opening above. 6. Anterior spiracle of M. bolivar (Bonatto), showing long, narrow opening gradually widening above. 7. Meral setae and posterior spiracle of M. bicolor (Fabricius). 8. Last abdominal tergites of M. vogelsangi (Mello), showing setae." httpUri="https://zenodo.org/record/3377241/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
); in a few species the anterior spiracle has a long, narrow opening gradually widening above (
<figureCitation id="13482A750F35FFDAFDA7BE3B3641FB84" box="[553,621,1052,1079]" captionStart="FIGURES 18" captionStartId="16.[151,264,1826,1851]" captionTargetBox="[183,1404,184,1807]" captionTargetId="figure@16.[183,1404,166,1807]" captionTargetPageId="16" captionText="FIGURES 18. Diagnostic characters of Mesembrinellidae and Calliphoridae. 1. Wing of Mesembrinella flavicrura Aldrich, showing veins and cells. 2. Wing of M. currani Guimar„es, showing infuscated anterior margin. 3. Wing of Laneella fusconitida sp. nov., showing dark macula around crossvein r-m. 4. Wing of Calliphora vomitoria (Linnaeus), showing acute bend of vein M. 5. Anterior spiracle of Mesembrinella batesi Aldrich, showing broad oval opening above. 6. Anterior spiracle of M. bolivar (Bonatto), showing long, narrow opening gradually widening above. 7. Meral setae and posterior spiracle of M. bicolor (Fabricius). 8. Last abdominal tergites of M. vogelsangi (Mello), showing setae." httpUri="https://zenodo.org/record/3377241/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
). Posterior thoracic spiracle varying in size from small to large, reniform (= kidney-shaped), with an opening midway along upper margin and a fringe of long, ventrally-directed setae (
<figureCitation id="13482A750F35FFDAFAE1BE67348BFBCC" captionStart="FIGURES 18" captionStartId="16.[151,264,1826,1851]" captionTargetBox="[183,1404,184,1807]" captionTargetId="figure@16.[183,1404,166,1807]" captionTargetPageId="16" captionText="FIGURES 18. Diagnostic characters of Mesembrinellidae and Calliphoridae. 1. Wing of Mesembrinella flavicrura Aldrich, showing veins and cells. 2. Wing of M. currani Guimar„es, showing infuscated anterior margin. 3. Wing of Laneella fusconitida sp. nov., showing dark macula around crossvein r-m. 4. Wing of Calliphora vomitoria (Linnaeus), showing acute bend of vein M. 5. Anterior spiracle of Mesembrinella batesi Aldrich, showing broad oval opening above. 6. Anterior spiracle of M. bolivar (Bonatto), showing long, narrow opening gradually widening above. 7. Meral setae and posterior spiracle of M. bicolor (Fabricius). 8. Last abdominal tergites of M. vogelsangi (Mello), showing setae." httpUri="https://zenodo.org/record/3377241/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
). Most species have a subscutellum ranging from weakly to moderately or prominently developed, different from the closely related
<taxonomicName id="4C734D730F35FFDAFEE2BEAF3628FB11" authorityName="Brauer &amp; von Bergenstamm" authorityYear="1889" box="[364,516,1160,1186]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Calliphoridae</taxonomicName>
where the subscutellum is not developed or at most weak. Row of meral setae in shape of an inverted L, with setae ranging from pale and fine to dark and stout (
<figureCitation id="13482A750F35FFDAFBA6BE8B3042FB74" box="[1064,1134,1196,1223]" captionStart="FIGURES 18" captionStartId="16.[151,264,1826,1851]" captionTargetBox="[183,1404,184,1807]" captionTargetId="figure@16.[183,1404,166,1807]" captionTargetPageId="16" captionText="FIGURES 18. Diagnostic characters of Mesembrinellidae and Calliphoridae. 1. Wing of Mesembrinella flavicrura Aldrich, showing veins and cells. 2. Wing of M. currani Guimar„es, showing infuscated anterior margin. 3. Wing of Laneella fusconitida sp. nov., showing dark macula around crossvein r-m. 4. Wing of Calliphora vomitoria (Linnaeus), showing acute bend of vein M. 5. Anterior spiracle of Mesembrinella batesi Aldrich, showing broad oval opening above. 6. Anterior spiracle of M. bolivar (Bonatto), showing long, narrow opening gradually widening above. 7. Meral setae and posterior spiracle of M. bicolor (Fabricius). 8. Last abdominal tergites of M. vogelsangi (Mello), showing setae." httpUri="https://zenodo.org/record/3377241/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
). Horizontal portion of inverted L usually composed of about 5 or more setae, in some species shorter, composed of only 23 setae or, rarely, absent. Wing with bend of vein M evenly curved (usually forming an obtuse angle,
<figureCitation id="13482A750F35FFDAFBC2BED33083FABD" box="[1100,1199,1268,1294]" captionStart="FIGURES 18" captionStartId="16.[151,264,1826,1851]" captionTargetBox="[183,1404,184,1807]" captionTargetId="figure@16.[183,1404,166,1807]" captionTargetPageId="16" captionText="FIGURES 18. Diagnostic characters of Mesembrinellidae and Calliphoridae. 1. Wing of Mesembrinella flavicrura Aldrich, showing veins and cells. 2. Wing of M. currani Guimar„es, showing infuscated anterior margin. 3. Wing of Laneella fusconitida sp. nov., showing dark macula around crossvein r-m. 4. Wing of Calliphora vomitoria (Linnaeus), showing acute bend of vein M. 5. Anterior spiracle of Mesembrinella batesi Aldrich, showing broad oval opening above. 6. Anterior spiracle of M. bolivar (Bonatto), showing long, narrow opening gradually widening above. 7. Meral setae and posterior spiracle of M. bicolor (Fabricius). 8. Last abdominal tergites of M. vogelsangi (Mello), showing setae." httpUri="https://zenodo.org/record/3377241/files/figure.png" pageId="15" pageNumber="16">Figs 1, 3</figureCitation>
) in most species, vs. normally angulate (at a right or acute angle) in
<taxonomicName id="4C734D730F35FFDAFD10BF3F371AFA81" authorityName="Brauer &amp; von Bergenstamm" authorityYear="1889" box="[670,822,1304,1330]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Calliphoridae</taxonomicName>
(
<figureCitation id="13482A750F35FFDAFCCBBF3F37A5FA81" box="[837,905,1304,1330]" captionStart="FIGURES 18" captionStartId="16.[151,264,1826,1851]" captionTargetBox="[183,1404,184,1807]" captionTargetId="figure@16.[183,1404,166,1807]" captionTargetPageId="16" captionText="FIGURES 18. Diagnostic characters of Mesembrinellidae and Calliphoridae. 1. Wing of Mesembrinella flavicrura Aldrich, showing veins and cells. 2. Wing of M. currani Guimar„es, showing infuscated anterior margin. 3. Wing of Laneella fusconitida sp. nov., showing dark macula around crossvein r-m. 4. Wing of Calliphora vomitoria (Linnaeus), showing acute bend of vein M. 5. Anterior spiracle of Mesembrinella batesi Aldrich, showing broad oval opening above. 6. Anterior spiracle of M. bolivar (Bonatto), showing long, narrow opening gradually widening above. 7. Meral setae and posterior spiracle of M. bicolor (Fabricius). 8. Last abdominal tergites of M. vogelsangi (Mello), showing setae." httpUri="https://zenodo.org/record/3377241/files/figure.png" pageId="15" pageNumber="16">Fig. 4</figureCitation>
). Frons width typically much narrower in males than in females, with frons to head ratios often providing valuable species-specific characters. Female frons broader, with width ranges often overlapping between species. Female with cruciate (crossed) setae just anterior to ocellar triangle. Male terminalia: most species with a roughly rectangular or square-shaped epandrium and medium-sized surstylus and cercus, adjacent at base (
<figureCitation id="13482A750F35FFDAFDC4BF8F368DFA71" box="[586,673,1447,1474]" captionStart="FIGURES 1520" captionStartId="18.[151,264,1973,1998]" captionTargetBox="[189,1314,166,1947]" captionTargetId="figure@18.[172,1416,166,1954]" captionTargetPageId="18" captionText="FIGURES 1520. Left lateral and posterior views of epandrium, cerci and surstyli of Laneella spp. 1516. L. fusconitida sp. nov. 1718. L. fuscosquamata sp. nov. 1920. L. nigripes Guimar„es." httpUri="https://zenodo.org/record/3377245/files/figure.png" pageId="15" pageNumber="16">Figs 15</figureCitation>
,
<figureCitation id="13482A750F35FFDAFD23BF8036E7FA71" box="[685,715,1447,1474]" captionStart="FIGURES 5356" captionStartId="29.[151,264,1486,1511]" captionTargetBox="[151,1421,181,1446]" captionTargetId="figure@29.[151,1436,181,1466]" captionTargetPageId="29" captionText="FIGURES 5356. Left lateral and posterior views of epandrium, cerci and surstyli of Mesembrinella spp. 5354. M. vogelsangi (Mello). 5556. M. zurquiensis sp. nov." httpUri="https://zenodo.org/record/3377259/files/figure.png" pageId="15" pageNumber="16">55</figureCitation>
,
<figureCitation id="13482A750F35FFDAFD56BF8F36DAFA71" box="[728,758,1448,1474]" captionStart="FIGURES 6368" captionStartId="31.[151,264,1980,2005]" captionTargetBox="[247,1305,166,1957]" captionTargetId="figure@31.[246,1341,166,1960]" captionTargetPageId="31" captionText="FIGURES 6368. Left lateral and posterior views of epandrium, cerci and surstyli of Mesembrinella spp. 6364. M. bicolor (Fabricius). 6566. M. brunnipes Surcouf. 6768. M. bullata sp. nov." httpUri="https://zenodo.org/record/3377263/files/figure.png" pageId="15" pageNumber="16">63</figureCitation>
,
<figureCitation id="13482A750F35FFDAFC8CBF8F370DFA71" box="[770,801,1448,1474]" captionStart="FIGURES 8792" captionStartId="36.[151,264,1965,1990]" captionTargetBox="[241,1313,166,1934]" captionTargetId="figure@36.[241,1347,166,1945]" captionTargetPageId="36" captionText="FIGURES 8792. Left lateral and posterior views of epandrium, cerci and surstyli of Mesembrinella spp. 8788. M. semihyalina Mello. 8990. M. serrata sp. nov. 9192. M. socors (Walker)." httpUri="https://zenodo.org/record/3377271/files/figure.png" pageId="15" pageNumber="16">87</figureCitation>
); in the
<taxonomicName id="4C734D730F35FFDAFCF0BF8E37E3FA71" authorityName="Aldrich." authorityYear="1925" box="[894,975,1449,1474]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="spicata">
<emphasis id="B907EAE20F35FFDAFCF0BF8E37E3FA71" box="[894,975,1449,1474]" italics="true" pageId="15" pageNumber="16">spicata</emphasis>
</taxonomicName>
-group the epandrium is elongate and the surstylus and cercus are tiny (
<figureCitation id="13482A750F35FFDAFE6BBFEB3644FA55" box="[485,616,1484,1510]" captionStart="FIGURES 2732" captionStartId="21.[151,264,1970,1995]" captionTargetBox="[175,1412,166,1905]" captionTargetId="figure@21.[175,1412,166,1949]" captionTargetPageId="21" captionText="FIGURES 2732. Left lateral and posterior views of epandrium, cerci and surstyli of Mesembrinella spp. 2728. M. mexicana sp. nov. 2930. M. spicata Aldrich. 3132. M. epandrioaurantia sp. nov." httpUri="https://zenodo.org/record/3377249/files/figure.png" pageId="15" pageNumber="16">Figs 27, 29</figureCitation>
); in the
<taxonomicName id="4C734D730F35FFDAFD4ABFEB3739FA55" authorityName="Bonatto in Bonatto &amp; Marinoni" authorityYear="2005" box="[708,789,1484,1510]" class="Insecta" family="Calliphoridae" genus="Giovanella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bolivar">
<emphasis id="B907EAE20F35FFDAFD4ABFEB3739FA55" box="[708,789,1484,1510]" italics="true" pageId="15" pageNumber="16">bolivar</emphasis>
</taxonomicName>
-group there is a narrow neck between the epandrium and the (swollen) base of the cercus, with surstylus and cercus widely separated in lateral view (
<figureCitation id="13482A750F35FFDAFB1EBFD730CAF9B9" box="[1168,1254,1520,1546]" captionStart="FIGURES 2732" captionStartId="21.[151,264,1970,1995]" captionTargetBox="[175,1412,166,1905]" captionTargetId="figure@21.[175,1412,166,1949]" captionTargetPageId="21" captionText="FIGURES 2732. Left lateral and posterior views of epandrium, cerci and surstyli of Mesembrinella spp. 2728. M. mexicana sp. nov. 2930. M. spicata Aldrich. 3132. M. epandrioaurantia sp. nov." httpUri="https://zenodo.org/record/3377249/files/figure.png" pageId="15" pageNumber="16">Figs 31</figureCitation>
,
<figureCitation id="13482A750F35FFDAFB7FBFD7313CF9B9" box="[1265,1296,1520,1546]" captionStart="FIGURES 3338" captionStartId="22.[151,264,1966,1991]" captionTargetBox="[233,1318,166,1942]" captionTargetId="figure@22.[233,1354,166,1946]" captionTargetPageId="22" captionText="FIGURES 3338. Left lateral and posterior views of epandrium, cerci and surstyli of Mesembrinella spp. 3334. M. woodorum sp. nov. 3536. M. aeneiventris (Wiedemann). 3738. M. bolivar (Bonatto)." httpUri="https://zenodo.org/record/3377251/files/figure.png" pageId="15" pageNumber="16">33</figureCitation>
); in the
<taxonomicName id="4C734D730F35FFDAFAE7BFD634E3F99D" authorityName="Guimar" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B907EAE20F35FFDAFAE7BFD634E3F99D" italics="true" pageId="15" pageNumber="16">anomala</emphasis>
</taxonomicName>
-group the epandrium is reduced and the surstylus is enlarged (
<figureCitation id="13482A750F35FFDAFC1CBC3337CBF99D" box="[914,999,1556,1582]" captionStart="FIGURES 99102" captionStartId="38.[151,264,1461,1486]" captionTargetBox="[155,1405,181,1431]" captionTargetId="figure@38.[151,1435,181,1441]" captionTargetPageId="38" captionText="FIGURES 99102. Left lateral and posterior views of epandrium, cerci and surstyli of Mesembrinellidae. 99100. Mesembrinella anomala (Guimar„es). 101102. Souzalopesiella facialis (Aldrich)." httpUri="https://zenodo.org/record/3377275/files/figure.png" pageId="15" pageNumber="16">Fig. 99</figureCitation>
). In most species the phallus is slender with epiphallus of various lengths and shapes, from long and slender (
<figureCitation id="13482A750F35FFDAFC3ABC1F3039F9E1" box="[948,1045,1591,1618]" captionStart="FIGURES 105114" captionStartId="43.[151,264,1934,1959]" captionTargetBox="[206,1388,177,1906]" captionTargetId="figure@43.[199,1388,166,1915]" captionTargetPageId="43" captionText="FIGURES 105114. Left lateral and dorsal views of phallus of Laneella spp. 105106. L. fusconitida sp. nov. 107108. L. fuscosquamata sp. nov. 109110. L. nigripes Guimar„es. 111112. L. perisi (Mariluis). 113114. L. purpurea sp. nov." httpUri="https://zenodo.org/record/3377279/files/figure.png" pageId="15" pageNumber="16">Fig. 105</figureCitation>
) to short and blunt (
<figureCitation id="13482A750F35FFDAFB70BC1F314EF9E1" box="[1278,1378,1591,1618]" captionStart="FIGURES 163172" captionStartId="51.[151,264,1927,1952]" captionTargetBox="[202,1386,175,1898]" captionTargetId="figure@51.[202,1386,166,1908]" captionTargetPageId="51" captionText="FIGURES 163172. Left lateral and dorsal views of phallus of Mesembrinella spp. 163164. M. cyaneicincta (Surcouf). 165166. M. flavicrura Aldrich. 167168. M. longicercus sp. nov. 169170. M. peregrina Aldrich. 171172. M. pictipennis Aldrich." httpUri="https://zenodo.org/record/3377291/files/figure.png" pageId="15" pageNumber="16">Fig. 165</figureCitation>
); hypophallic lobes often of unique shape from rounded (
<figureCitation id="13482A750F35FFDAFD67BC7B3765F9C4" box="[745,841,1628,1655]" captionStart="FIGURES 105114" captionStartId="43.[151,264,1934,1959]" captionTargetBox="[206,1388,177,1906]" captionTargetId="figure@43.[199,1388,166,1915]" captionTargetPageId="43" captionText="FIGURES 105114. Left lateral and dorsal views of phallus of Laneella spp. 105106. L. fusconitida sp. nov. 107108. L. fuscosquamata sp. nov. 109110. L. nigripes Guimar„es. 111112. L. perisi (Mariluis). 113114. L. purpurea sp. nov." httpUri="https://zenodo.org/record/3377279/files/figure.png" pageId="15" pageNumber="16">Fig. 106</figureCitation>
) to oval (
<figureCitation id="13482A750F35FFDAFC3ABC7B303DF9C5" box="[948,1041,1628,1654]" captionStart="FIGURES 105114" captionStartId="43.[151,264,1934,1959]" captionTargetBox="[206,1388,177,1906]" captionTargetId="figure@43.[199,1388,166,1915]" captionTargetPageId="43" captionText="FIGURES 105114. Left lateral and dorsal views of phallus of Laneella spp. 105106. L. fusconitida sp. nov. 107108. L. fuscosquamata sp. nov. 109110. L. nigripes Guimar„es. 111112. L. perisi (Mariluis). 113114. L. purpurea sp. nov." httpUri="https://zenodo.org/record/3377279/files/figure.png" pageId="15" pageNumber="16">Fig. 111</figureCitation>
) or rectangular (
<figureCitation id="13482A750F35FFDAFB42BC7B3102F9C5" box="[1228,1326,1628,1654]" captionStart="FIGURES 173182" captionStartId="52.[151,264,1935,1960]" captionTargetBox="[199,1388,175,1895]" captionTargetId="figure@52.[199,1388,166,1915]" captionTargetPageId="52" captionText="FIGURES 173182. Left lateral and dorsal views of phallus of Mesembrinella spp. 173174. M. quadrilineata (Fabricius). 175176. M. randa (Walker). 177178. M. semihyalina Mello. 179180. M. serrata sp. nov. 181182. M. socors (Walker)." httpUri="https://zenodo.org/record/3377293/files/figure.png" pageId="15" pageNumber="16">Fig. 182</figureCitation>
); in a few species the phallus is highly modified (
<figureCitation id="13482A750F35FFDAFDC3BCA736C7F929" box="[589,747,1664,1690]" captionStart="FIGURES 115124" captionStartId="44.[151,264,1938,1963]" captionTargetBox="[204,1390,171,1917]" captionTargetId="figure@44.[198,1390,166,1919]" captionTargetPageId="44" captionText="FIGURES 115124. Left lateral and dorsal views of phallus of Mesembrinella spp. 115116. M. latifrons (Mello). 117118. M. mexicana sp. nov. 119120. M. spicata Aldrich. 121122. M. epandrioaurantia sp. nov. 123124. M. woodorum sp. nov." httpUri="https://zenodo.org/record/3377281/files/figure.png" pageId="15" pageNumber="16">Figs 117120</figureCitation>
). In males of most species ST1 is broad, ST24 are gradually narrowing (e.g.,
<figureCitation id="13482A750F35FFDAFEDEBC833582F90D" box="[336,430,1700,1726]" captionStart="FIGURES 239251" captionStartId="69.[151,264,1864,1889]" captionTargetBox="[223,1364,166,1844]" captionTargetId="figure@69.[223,1364,166,1845]" captionTargetPageId="69" captionText="FIGURES 239251. Male sternites of Lanella spp. 239. L. fusconitida sp. nov. 240. L. fuscosquamata sp. nov. 241. L. nigripes Guimar„es 242. L. perisi (Mariluis). 243. L. purpurea sp. nov. 244. M. latifrons (Mello). 245. M. mexicana sp. nov. 246. M. spicata Aldrich. 247. M. epandrioaurantia sp. nov. 248. M. woodorum sp. nov. 249. M. aeneiventris (Wiedemann). 250. M. decrepita Séguy. 251. M. lara (Bonatto) (ST5 only)." httpUri="https://zenodo.org/record/3377313/files/figure.png" pageId="15" pageNumber="16">Fig. 239</figureCitation>
) and ST5 is usually broad and bilobed, sometimes modified (
<figureCitation id="13482A750F35FFDAFBD6BC833148F90D" box="[1112,1380,1699,1726]" captionStart="FIGURES 239251" captionStartId="69.[151,264,1864,1889]" captionTargetBox="[223,1364,166,1844]" captionTargetId="figure@69.[223,1364,166,1845]" captionTargetPageId="69" captionText="FIGURES 239251. Male sternites of Lanella spp. 239. L. fusconitida sp. nov. 240. L. fuscosquamata sp. nov. 241. L. nigripes Guimar„es 242. L. perisi (Mariluis). 243. L. purpurea sp. nov. 244. M. latifrons (Mello). 245. M. mexicana sp. nov. 246. M. spicata Aldrich. 247. M. epandrioaurantia sp. nov. 248. M. woodorum sp. nov. 249. M. aeneiventris (Wiedemann). 250. M. decrepita Séguy. 251. M. lara (Bonatto) (ST5 only)." httpUri="https://zenodo.org/record/3377313/files/figure.png" pageId="15" pageNumber="16">Figs 239, 245, 247, 249</figureCitation>
). Female with ST15 oval to nearly square or rectangular (
<figureCitation id="13482A750F35FFDAFD77BCEF37B4F951" box="[761,920,1735,1762]" captionStart-0="FIGURES 411423" captionStart-1="FIGURES 424438" captionStart-2="FIGURES 439453" captionStart-3="FIGURE 454" captionStartId-0="91.[151,264,1864,1889]" captionStartId-1="92.[151,264,1856,1881]" captionStartId-2="93.[151,264,1860,1885]" captionStartId-3="94.[151,250,1945,1970]" captionTargetBox-0="[223,1324,166,1843]" captionTargetBox-1="[232,1362,166,1835]" captionTargetBox-2="[224,1360,166,1832]" captionTargetId-0="figure@91.[223,1364,166,1844]" captionTargetId-1="figure@92.[226,1362,166,1837]" captionTargetId-2="figure@93.[224,1362,166,1841]" captionTargetId-3="figure@94.[335,1253,164,1926]" captionTargetPageId-0="91" captionTargetPageId-1="92" captionTargetPageId-2="93" captionTargetPageId-3="94" captionText-0="FIGURES 411423. Female sternites of Mesembrinellidae. 411. Laneella fusconitida sp. nov. (LACM). 412. L. fuscosquamata sp. nov. 413. L. nigripes Guimar„es. 414. L. perisi (Mariluis). 415. L. purpurea sp. nov. 416. Mesembrinella latifrons (Mello). 417. M. guaramacalensis sp. nov. 418. M. mexicana sp. nov. 419. M. spicata Aldrich. 420. M. epandrioaurantia sp. nov. 421. M. aeneiventris (Wiedemann). 422. M. decrepita Séguy. 423. M. lara (Bonatto)." captionText-1="FIGURES 424438. Female sternites of Mesembrinella spp. 424. M. nigrocoerulea sp. nov. 425. M. purpurata Aldrich. 426. M. semiflava Aldrich. 427. M. uniseta Aldrich. 428. M. violacea sp. nov. 429. M. vogelsangi (Mello). 430. M. zurquiensis sp. nov. 431. M. apollinaris Séguy. 432. M. batesi Aldrich. 433. M. bellardiana Aldrich. 434. M. benoisti (Séguy). 435. M. bicolor (Fabricius). 436. M. brunnipes Surcouf. 437. M. bullata sp. nov. 438. M. chantryi sp. nov." captionText-2="FIGURES 439453. Female sternites of Mesembrinella spp. 439. M. currani Guimar„es. 440. M. cyaneicincta (Surcouf). 441. M. flavicrura Aldrich. 442. M. longicercus sp. nov. 443. M. peregrina Aldrich. 444. M. pictipennis Aldrich. 445. M. quadrilineata (Fabricius). 446. M. randa (Walker). 447. M. semihyalina Mello. 448. M. serrata sp. nov. 449. M. socors (Walker). 450. M. townsendi Guimar„es. 451. M. umbrosa Aldrich. 452. M. velasquezae sp. nov. 453. M. anomala (Guimar„es)." captionText-3="FIGURE 454. Female sternites of Souzalopesiella facialis (Aldrich)." figureDoi-3="http://doi.org/10.5281/zenodo.3445960" httpUri-0="https://zenodo.org/record/3377347/files/figure.png" httpUri-1="https://zenodo.org/record/3377349/files/figure.png" httpUri-2="https://zenodo.org/record/3377351/files/figure.png" httpUri-3="https://zenodo.org/record/3445960/files/figure.png" pageId="15" pageNumber="16">Figs 411454</figureCitation>
). Female postabdomen not telescopic; dorsum of ovipositor with large T6, ranging from a flattened (e.g.,
<figureCitation id="13482A750F35FFDAFCAABCCB37AFF8B5" box="[804,899,1772,1798]" captionStart="FIGURES 282287" captionStartId="74.[151,264,1931,1956]" captionTargetBox="[157,1373,189,1911]" captionTargetId="figure@74.[151,1435,166,1911]" captionTargetPageId="74" captionText="FIGURES 282287. Female terminalia (T6T8, epiproct, cerci) of Mesembrinellidae, dorsal view. 282. Laneella fusconitida sp. nov. 283. L. fuscosquamata sp. nov. 284. L. nigripes Guimar„es. 285. L. perisi (Mariluis). 286. L. purpurea sp. nov. 287. M. latifrons (Mello)." httpUri="https://zenodo.org/record/3377321/files/figure.png" pageId="15" pageNumber="16">Fig. 282</figureCitation>
) to a regular U or V shape (e.g.,
<figureCitation id="13482A750F35FFDAFB61BCCB317CF8B5" box="[1263,1360,1772,1798]" captionStart="FIGURES 282287" captionStartId="74.[151,264,1931,1956]" captionTargetBox="[157,1373,189,1911]" captionTargetId="figure@74.[151,1435,166,1911]" captionTargetPageId="74" captionText="FIGURES 282287. Female terminalia (T6T8, epiproct, cerci) of Mesembrinellidae, dorsal view. 282. Laneella fusconitida sp. nov. 283. L. fuscosquamata sp. nov. 284. L. nigripes Guimar„es. 285. L. perisi (Mariluis). 286. L. purpurea sp. nov. 287. M. latifrons (Mello)." httpUri="https://zenodo.org/record/3377321/files/figure.png" pageId="15" pageNumber="16">Fig. 287</figureCitation>
), often with distinctive spiracles; T7 smaller and either bilobed (
<figureCitation id="13482A750F35FFDAFC9DBD37375FF899" box="[787,883,1808,1834]" captionStart="FIGURES 282287" captionStartId="74.[151,264,1931,1956]" captionTargetBox="[157,1373,189,1911]" captionTargetId="figure@74.[151,1435,166,1911]" captionTargetPageId="74" captionText="FIGURES 282287. Female terminalia (T6T8, epiproct, cerci) of Mesembrinellidae, dorsal view. 282. Laneella fusconitida sp. nov. 283. L. fuscosquamata sp. nov. 284. L. nigripes Guimar„es. 285. L. perisi (Mariluis). 286. L. purpurea sp. nov. 287. M. latifrons (Mello)." httpUri="https://zenodo.org/record/3377321/files/figure.png" pageId="15" pageNumber="16">Fig. 282</figureCitation>
) or divided (
<figureCitation id="13482A750F35FFDAFB8CBD373048F899" box="[1026,1124,1808,1834]" captionStart="FIGURES 288293" captionStartId="75.[151,264,1939,1964]" captionTargetBox="[197,1389,196,1908]" captionTargetId="figure@75.[197,1389,166,1920]" captionTargetPageId="75" captionText="FIGURES 288293. Female terminalia (T6T8, epiproct, cerci) of Mesembrinella spp., dorsal view. 288. M. guaramacalensis sp. nov. 289. M. mexicana sp. nov. 290. M. spicata Aldrich. 291. M. epandrioaurantia sp. nov. 292. M. aeneiventris (Wiedemann). 293. M. decrepita Séguy." httpUri="https://zenodo.org/record/3377323/files/figure.png" pageId="15" pageNumber="16">Fig. 289</figureCitation>
); T8 much reduced, divided (
<figureCitation id="13482A750F35FFDAFF2EBD13352CF8FD" box="[160,256,1844,1870]" captionStart="FIGURES 282287" captionStartId="74.[151,264,1931,1956]" captionTargetBox="[157,1373,189,1911]" captionTargetId="figure@74.[151,1435,166,1911]" captionTargetPageId="74" captionText="FIGURES 282287. Female terminalia (T6T8, epiproct, cerci) of Mesembrinellidae, dorsal view. 282. Laneella fusconitida sp. nov. 283. L. fuscosquamata sp. nov. 284. L. nigripes Guimar„es. 285. L. perisi (Mariluis). 286. L. purpurea sp. nov. 287. M. latifrons (Mello)." httpUri="https://zenodo.org/record/3377321/files/figure.png" pageId="15" pageNumber="16">Fig. 284</figureCitation>
) or continuous (
<figureCitation id="13482A750F35FFDAFE37BD133630F8FD" box="[441,540,1844,1870]" captionStart="FIGURES 300305" captionStartId="77.[151,264,1942,1967]" captionTargetBox="[151,1435,176,1912]" captionTargetId="figure@77.[151,1435,166,1922]" captionTargetPageId="77" captionText="FIGURES 300305. Female terminalia (T6T8, epiproct, cerci) of Mesembrinella spp., dorsal view. 300. M. vogelsangi (Mello). 301. M. zurquiensis sp. nov. 302. M. apollinaris Séguy. 303. M. batesi Aldrich. 304. M. bellardiana Aldrich. 305. M. benoisti (Séguy)." httpUri="https://zenodo.org/record/3377327/files/figure.png" pageId="15" pageNumber="16">Fig. 303</figureCitation>
); epiproct and cerci as in
<figureCitation id="13482A750F35FFDAFCB0BD1337F7F8FD" box="[830,987,1843,1870]" captionStart-0="FIGURES 282287" captionStart-1="FIGURES 288293" captionStart-2="FIGURES 294299" captionStart-3="FIGURES 300305" captionStart-4="FIGURES 306311" captionStart-5="FIGURES 312317" captionStart-6="FIGURES 318323" captionStart-7="FIGURES 324325" captionStartId-0="74.[151,264,1931,1956]" captionStartId-1="75.[151,264,1939,1964]" captionStartId-2="76.[151,264,1942,1967]" captionStartId-3="77.[151,264,1942,1967]" captionStartId-4="80.[151,264,1931,1956]" captionStartId-5="81.[151,264,1935,1960]" captionStartId-6="82.[151,264,1931,1956]" captionStartId-7="83.[151,264,769,794]" captionTargetBox-0="[157,1373,189,1911]" captionTargetBox-1="[197,1389,196,1908]" captionTargetBox-2="[151,1435,192,1902]" captionTargetBox-3="[151,1435,176,1912]" captionTargetBox-4="[207,1369,169,1911]" captionTargetBox-5="[151,1431,176,1907]" captionTargetBox-6="[151,1435,166,1897]" captionTargetBox-7="[160,1434,181,747]" captionTargetId-0="figure@74.[151,1435,166,1911]" captionTargetId-1="figure@75.[197,1389,166,1920]" captionTargetId-2="figure@76.[151,1435,166,1923]" captionTargetId-3="figure@77.[151,1435,166,1922]" captionTargetId-4="figure@80.[200,1386,166,1911]" captionTargetId-5="figure@81.[151,1435,166,1915]" captionTargetId-6="figure@82.[151,1435,166,1911]" captionTargetId-7="figure@83.[151,1435,181,749]" captionTargetPageId-0="74" captionTargetPageId-1="75" captionTargetPageId-2="76" captionTargetPageId-3="77" captionTargetPageId-4="80" captionTargetPageId-5="81" captionTargetPageId-6="82" captionTargetPageId-7="83" captionText-0="FIGURES 282287. Female terminalia (T6T8, epiproct, cerci) of Mesembrinellidae, dorsal view. 282. Laneella fusconitida sp. nov. 283. L. fuscosquamata sp. nov. 284. L. nigripes Guimar„es. 285. L. perisi (Mariluis). 286. L. purpurea sp. nov. 287. M. latifrons (Mello)." captionText-1="FIGURES 288293. Female terminalia (T6T8, epiproct, cerci) of Mesembrinella spp., dorsal view. 288. M. guaramacalensis sp. nov. 289. M. mexicana sp. nov. 290. M. spicata Aldrich. 291. M. epandrioaurantia sp. nov. 292. M. aeneiventris (Wiedemann). 293. M. decrepita Séguy." captionText-2="FIGURES 294299. Female terminalia (T6T8, epiproct, cerci) of Mesembrinella spp., dorsal view. 294. M. lara (Bonatto). 295. M. nigrocoerulea sp. nov. 296. M. purpurata Aldrich. 297. M. semiflava Aldrich. 298. M. uniseta Aldrich. 299. M. violacea sp. nov." captionText-3="FIGURES 300305. Female terminalia (T6T8, epiproct, cerci) of Mesembrinella spp., dorsal view. 300. M. vogelsangi (Mello). 301. M. zurquiensis sp. nov. 302. M. apollinaris Séguy. 303. M. batesi Aldrich. 304. M. bellardiana Aldrich. 305. M. benoisti (Séguy)." captionText-4="FIGURES 306311. Female terminalia (T6T8, epiproct, cerci) of Mesembrinella spp., dorsal view. 306. M. bicolor (Fabricius). 307. M. brunnipes Surcouf 308. M. bullata sp. nov. 309. M. chantryi sp. nov. 310. M. currani Guimar„es. 311. M. cyaneicincta (Surcouf)." captionText-5="FIGURES 312317. Female terminalia (T6T8, epiproct, cerci) of Mesembrinella spp., dorsal view. 312. M. flavicrura Aldrich. 313. M. longicercus sp. nov. 314. M. peregrina Aldrich. 315. M. pictipennis Aldrich. 316. M. quadrilineata (Fabricius). 317. M. randa (Walker)." captionText-6="FIGURES 318323. Female terminalia (T6T8, epiproct, cerci) of Mesembrinella spp., dorsal view. 318. M. semihyalina Mello. 319. M. serrata sp. nov. 320. M. socors (Walker). 321. M. townsendi Guimar„es. 322. M. umbrosa Aldrich. 323. M. velasquezae sp. nov." captionText-7="FIGURES 324325. Female terminalia (T6T8, epiproct, cerci) of Mesembrinellidae, dorsal view. 324. Mesembrinella anomala (Guimar„es). 325. Souzalopesiella facialis (Aldrich)." httpUri-0="https://zenodo.org/record/3377321/files/figure.png" httpUri-1="https://zenodo.org/record/3377323/files/figure.png" httpUri-2="https://zenodo.org/record/3377325/files/figure.png" httpUri-3="https://zenodo.org/record/3377327/files/figure.png" httpUri-4="https://zenodo.org/record/3377329/files/figure.png" httpUri-5="https://zenodo.org/record/3377331/files/figure.png" httpUri-6="https://zenodo.org/record/3377335/files/figure.png" httpUri-7="https://zenodo.org/record/3377337/files/figure.png" pageId="15" pageNumber="16">Figs 282325</figureCitation>
. Venter of ovipositor composed of ST6, ST7, a greatly reduced ST8, and hypoproct (
<figureCitation id="13482A750F35FFDAFD1FBD7F36D9F8C0" box="[657,757,1880,1907]" captionStart="FIGURES 326349" captionStartId="85.[151,264,1757,1782]" captionTargetBox="[263,1328,166,1735]" captionTargetId="figure@85.[260,1328,166,1737]" captionTargetPageId="85" captionText="FIGURES 326349. ST68 and hypoproct of Mesembrinellidae. 326. Laneella fusconitida sp. nov. 327. L. fuscosquamata sp. nov. 328. L. nigripes Guimar„es. 329. L. perisi (Mariluis). 330. L. purpurea sp. nov. 331. Mesembrinella latifrons (Mello). 332. M. guaramacalensis sp. nov. 333. M. mexicana sp. nov. 334. M. spicata Aldrich. 335. M. epandrioaurantia sp. nov. 336. M. aeneiventris (Wiedemann). 337. M. decrepita Séguy. 338. M. lara (Bonatto). 339. M. nigrocoerulea sp. nov. 340. M. purpurata Aldrich. 341. M. semiflava Aldrich. 342. M. uniseta Aldrich. 343. M. violacea sp. nov. 344. M. vogelsangi (Mello). 345. M. zurquiensis sp. nov. 346. M. apollinaris Séguy. 347. M. batesi Aldrich. 348. M. bellardiana Aldrich. 349. M. benoisti (Séguy)." httpUri="https://zenodo.org/record/3377339/files/figure.png" pageId="15" pageNumber="16">Fig. 326</figureCitation>
). Spermathecae (three in number) of three distinctive
<typeStatus id="54C888520F35FFDAFAD7BD7E31B4F8C0" box="[1369,1432,1881,1907]" pageId="15" pageNumber="16">types</typeStatus>
: filiform (
<figureCitation id="13482A750F35FFDAFF73BD5B35B0F825" box="[253,412,1915,1942]" captionStart="FIGURES 370400" captionStartId="87.[151,264,1727,1752]" captionTargetBox="[274,1318,166,1707]" captionTargetId="figure@87.[270,1318,166,1707]" captionTargetPageId="87" captionText="FIGURES 370400. Spermathecae of Mesembrinellidae. 370. L. fusconitida sp. nov. 371. L. fuscosquamata sp. nov. 372. L. nigripes Guimar„es. 373. L. perisi (Mariluis). 374. L. purpurea sp. nov. 375. Mesembrinella guaramacalensis sp. nov. 376. M. mexicana sp. nov. 377. M. spicata Aldrich. 378. M. aeneiventris (Wiedemann). 379. M. decrepita Séguy. 380. M. lara (Bonatto). 381. M. nigrocoerulea sp. nov. 382. M. purpurata Aldrich. 383. M. semiflava Aldrich. 384. M. uniseta Aldrich. 385. M. vogelsangi (Mello). 386. M. zurquiensis sp. nov. 387. M. apollinaris Séguy. 388. M. batesi Aldrich. 389. M. bellardiana Aldrich. 390. M. benoisti (Séguy). 391. M. bicolor (Fabricius). 392. M. brunnipes Surcouf. 393. M. bullata sp. nov. 394. M. chantryi sp. nov. 395. M. currani Guimar„es. 396. M. cyaneicincta (Surcouf). 397. M. flavicrura Aldrich. 398. M. longicercus sp. nov. 399. M. peregrina Aldrich. 400. M. pictipennis Aldrich." httpUri="https://zenodo.org/record/3377343/files/figure.png" pageId="15" pageNumber="16">Figs 375396</figureCitation>
; most species), tuberform (
<figureCitation id="13482A750F35FFDAFD45BD5B3745F825" box="[715,873,1916,1942]" captionStart="FIGURES 370400" captionStartId="87.[151,264,1727,1752]" captionTargetBox="[274,1318,166,1707]" captionTargetId="figure@87.[270,1318,166,1707]" captionTargetPageId="87" captionText="FIGURES 370400. Spermathecae of Mesembrinellidae. 370. L. fusconitida sp. nov. 371. L. fuscosquamata sp. nov. 372. L. nigripes Guimar„es. 373. L. perisi (Mariluis). 374. L. purpurea sp. nov. 375. Mesembrinella guaramacalensis sp. nov. 376. M. mexicana sp. nov. 377. M. spicata Aldrich. 378. M. aeneiventris (Wiedemann). 379. M. decrepita Séguy. 380. M. lara (Bonatto). 381. M. nigrocoerulea sp. nov. 382. M. purpurata Aldrich. 383. M. semiflava Aldrich. 384. M. uniseta Aldrich. 385. M. vogelsangi (Mello). 386. M. zurquiensis sp. nov. 387. M. apollinaris Séguy. 388. M. batesi Aldrich. 389. M. bellardiana Aldrich. 390. M. benoisti (Séguy). 391. M. bicolor (Fabricius). 392. M. brunnipes Surcouf. 393. M. bullata sp. nov. 394. M. chantryi sp. nov. 395. M. currani Guimar„es. 396. M. cyaneicincta (Surcouf). 397. M. flavicrura Aldrich. 398. M. longicercus sp. nov. 399. M. peregrina Aldrich. 400. M. pictipennis Aldrich." httpUri="https://zenodo.org/record/3377343/files/figure.png" pageId="15" pageNumber="16">Figs 370374</figureCitation>
; genus
<taxonomicName id="4C734D730F35FFDAFC33BD5B3033F825" authorityName=", Laneellinae" box="[957,1055,1916,1942]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F35FFDAFC33BD5B3033F825" box="[957,1055,1916,1942]" italics="true" pageId="15" pageNumber="16">Laneella</emphasis>
</taxonomicName>
, though significantly shorter in
<taxonomicName id="4C734D730F35FFDAFA08BD5A3513F809" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fuscosquamata">
<emphasis id="B907EAE20F35FFDAFA08BD5A3513F809" italics="true" pageId="15" pageNumber="16">L. fuscosquamata</emphasis>
</taxonomicName>
), or bulbous (
<figureCitation id="13482A750F35FFDAFE52BD87366EF809" box="[476,578,1952,1978]" captionStart="FIGURES 370400" captionStartId="87.[151,264,1727,1752]" captionTargetBox="[274,1318,166,1707]" captionTargetId="figure@87.[270,1318,166,1707]" captionTargetPageId="87" captionText="FIGURES 370400. Spermathecae of Mesembrinellidae. 370. L. fusconitida sp. nov. 371. L. fuscosquamata sp. nov. 372. L. nigripes Guimar„es. 373. L. perisi (Mariluis). 374. L. purpurea sp. nov. 375. Mesembrinella guaramacalensis sp. nov. 376. M. mexicana sp. nov. 377. M. spicata Aldrich. 378. M. aeneiventris (Wiedemann). 379. M. decrepita Séguy. 380. M. lara (Bonatto). 381. M. nigrocoerulea sp. nov. 382. M. purpurata Aldrich. 383. M. semiflava Aldrich. 384. M. uniseta Aldrich. 385. M. vogelsangi (Mello). 386. M. zurquiensis sp. nov. 387. M. apollinaris Séguy. 388. M. batesi Aldrich. 389. M. bellardiana Aldrich. 390. M. benoisti (Séguy). 391. M. bicolor (Fabricius). 392. M. brunnipes Surcouf. 393. M. bullata sp. nov. 394. M. chantryi sp. nov. 395. M. currani Guimar„es. 396. M. cyaneicincta (Surcouf). 397. M. flavicrura Aldrich. 398. M. longicercus sp. nov. 399. M. peregrina Aldrich. 400. M. pictipennis Aldrich." httpUri="https://zenodo.org/record/3377343/files/figure.png" pageId="15" pageNumber="16">Figs 397</figureCitation>
,
<figureCitation id="13482A750F35FFDAFDC3BD873656F809" box="[589,634,1952,1978]" captionStart="FIGURES 401410" captionStartId="89.[151,264,1699,1724]" captionTargetBox="[154,1435,1032,1669]" captionTargetId="figure@89.[151,1435,1029,1669]" captionTargetPageId="89" captionText="FIGURES 401410. Spermathecae of Mesembrinellidae. 401. Mesembrinella quadrilineata (Fabricius). 402. M. randa (Walker). 403. M. semihyalina Mello. 404. M. serrata sp. nov. 405. M. socors (Walker). 406. M. townsendi Guimar„es. 407. M. umbrosa Aldrich. 408. M. velasquezae sp. nov. 409. M. anomala (Guimar„es). 410. Souzalopesiella facialis (Aldrich)." httpUri="https://zenodo.org/record/3377345/files/figure.png" pageId="15" pageNumber="16">410</figureCitation>
;
<taxonomicName id="4C734D730F35FFDAFD08BD863737F809" authorityName="Aldrich." authorityYear="1925" box="[646,795,1952,1978]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="flavicrura">
<emphasis id="B907EAE20F35FFDAFD08BD863737F809" box="[646,795,1952,1978]" italics="true" pageId="15" pageNumber="16">M. flavicrura</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D730F35FFDAFCDCBD873076F809" baseAuthorityName="Aldrich" baseAuthorityYear="1922" box="[850,1114,1952,1978]" class="Insecta" family="Mesembrinellidae" genus="Souzalopesiella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="facialis">
<emphasis id="B907EAE20F35FFDAFCDCBD873076F809" box="[850,1114,1952,1978]" italics="true" pageId="15" pageNumber="16">Souzalopesiella facialis</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF0C66780F2AFFC5FF19BD0537AFF85D" httpUri="https://zenodo.org/record/3377241/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,264,1826,1851]" targetBox="[183,1404,184,1807]" targetPageId="16">
<paragraph id="8BCC36F00F2AFFC5FF19BD0537AFF85D" blockId="16.[151,1437,1826,2031]" pageId="16" pageNumber="17">
<emphasis id="B907EAE20F2AFFC5FF19BD053510F888" bold="true" box="[151,316,1826,1851]" pageId="16" pageNumber="17">FIGURES 18.</emphasis>
Diagnostic characters of
<taxonomicName id="4C734D730F2AFFC5FDCABD0536D7F888" authorityName="Shannon" authorityYear="1926" box="[580,763,1826,1851]" class="Insecta" family="Mesembrinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Mesembrinellidae</taxonomicName>
and
<taxonomicName id="4C734D730F2AFFC5FCA1BD053790F888" authorityName="Brauer &amp; von Bergenstamm" authorityYear="1889" box="[815,956,1826,1851]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Calliphoridae</taxonomicName>
.
<emphasis id="B907EAE20F2AFFC5FC49BD0537F6F888" bold="true" box="[967,986,1826,1851]" pageId="16" pageNumber="17">1.</emphasis>
Wing of
<taxonomicName id="4C734D730F2AFFC5FBB3BD0431B5F888" authority="Aldrich" authorityName="Aldrich." authorityYear="1925" box="[1085,1433,1826,1851]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="flavicrura">
<emphasis id="B907EAE20F2AFFC5FBB3BD04316EF888" box="[1085,1346,1827,1851]" italics="true" pageId="16" pageNumber="17">Mesembrinella flavicrura</emphasis>
Aldrich
</taxonomicName>
, showing veins and cells.
<emphasis id="B907EAE20F2AFFC5FE1CBD613589F8EC" bold="true" box="[402,421,1862,1887]" pageId="16" pageNumber="17">2.</emphasis>
Wing of
<taxonomicName id="4C734D730F2AFFC5FD8FBD6F36EAF8EC" authority="Guimar" authorityName="Guimaraes" authorityYear="1977" box="[513,710,1862,1887]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="B907EAE20F2AFFC5FD8FBD6F3642F8EC" box="[513,622,1864,1887]" italics="true" pageId="16" pageNumber="17">M. currani</emphasis>
Guimar
</taxonomicName>
„es, showing infuscated anterior margin.
<emphasis id="B907EAE20F2AFFC5FBD0BD61305DF8EC" bold="true" box="[1118,1137,1862,1887]" pageId="16" pageNumber="17">3.</emphasis>
Wing of
<taxonomicName id="4C734D730F2AFFC5FB43BD6031B1F8EC" authority="Whitworth &amp; Yusseff-Vanegas, 2019" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" box="[1229,1437,1863,1887]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="fusconitida" status="sp. nov.">
<emphasis id="B907EAE20F2AFFC5FB43BD6031B1F8EC" box="[1229,1437,1863,1887]" italics="true" pageId="16" pageNumber="17">Laneella fusconitida</emphasis>
</taxonomicName>
<emphasis id="B907EAE20F2AFFC5FF19BD4C34C4F830" bold="true" box="[151,232,1899,1923]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A23457990F2AFFC5FF19BD4C34C4F830" box="[151,232,1899,1923]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, showing dark macula around crossvein r-m.
<emphasis id="B907EAE20F2AFFC5FD37BD4D36E0F830" bold="true" box="[697,716,1898,1923]" pageId="16" pageNumber="17">4.</emphasis>
Wing of
<taxonomicName id="4C734D730F2AFFC5FCA5BD4C3056F830" authority="(Linnaeus)" authorityName="Linnaeus" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[811,1146,1898,1923]" class="Insecta" family="Calliphoridae" genus="Calliphora" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="vomitoria">
<emphasis id="B907EAE20F2AFFC5FCA5BD4C3028F830" box="[811,1028,1899,1923]" italics="true" pageId="16" pageNumber="17">Calliphora vomitoria</emphasis>
(Linnaeus)
</taxonomicName>
, showing acute bend of vein M.
<emphasis id="B907EAE20F2AFFC5FF35BDA834E1F814" bold="true" box="[187,205,1935,1959]" pageId="16" pageNumber="17">5.</emphasis>
Anterior spiracle of
<taxonomicName id="4C734D730F2AFFC5FE2EBDA836FDF814" authority="Aldrich" authorityName="Aldrich." authorityYear="1922" box="[416,721,1934,1959]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="B907EAE20F2AFFC5FE2EBDA83650F814" box="[416,636,1935,1959]" italics="true" pageId="16" pageNumber="17">Mesembrinella batesi</emphasis>
Aldrich
</taxonomicName>
, showing broad oval opening above.
<emphasis id="B907EAE20F2AFFC5FBC5BDA93072F814" bold="true" box="[1099,1118,1934,1959]" pageId="16" pageNumber="17">6.</emphasis>
Anterior spiracle of
<taxonomicName id="4C734D730F2AFFC5FABFBDB734D7F878" authority="(Bonatto)" authorityName="Bonatto" baseAuthorityName="Bonatto" class="Insecta" family="Calliphoridae" genus="Giovanella" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bolivar">
<emphasis id="B907EAE20F2AFFC5FABFBDB731B0F814" box="[1329,1436,1935,1959]" italics="true" pageId="16" pageNumber="17">M. bolivar</emphasis>
(Bonatto)
</taxonomicName>
, showing long, narrow opening gradually widening above.
<emphasis id="B907EAE20F2AFFC5FCDABD94374BF878" bold="true" box="[852,871,1971,1995]" pageId="16" pageNumber="17">7.</emphasis>
Meral setae and posterior spiracle of
<taxonomicName id="4C734D730F2AFFC5FB6BBD9334E0F85C" authority="(Fabricius)" authorityName="Fabricius" baseAuthorityName="Fabricius" baseAuthorityYear="1805" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B907EAE20F2AFFC5FB6BBD933163F878" box="[1253,1359,1971,1995]" italics="true" pageId="16" pageNumber="17">M. bicolor</emphasis>
(Fabricius)
</taxonomicName>
.
<emphasis id="B907EAE20F2AFFC5FF5BBDF134C4F85C" bold="true" box="[213,232,2006,2031]" pageId="16" pageNumber="17">8.</emphasis>
Last abdominal tergites of
<taxonomicName id="4C734D730F2AFFC5FE70BDFF36CFF85C" authority="(Mello)" authorityName="Mello" baseAuthorityName="Mello" baseAuthorityYear="1967" box="[510,739,2006,2031]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="vogelsangi">
<emphasis id="B907EAE20F2AFFC5FE70BDFF36A1F85C" box="[510,653,2007,2031]" italics="true" pageId="16" pageNumber="17">M. vogelsangi</emphasis>
(Mello)
</taxonomicName>
, showing setae.
</paragraph>
</caption>
<caption id="DF0C66780F2BFFC4FF19BE2B3605FB06" httpUri="https://zenodo.org/record/3377243/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,264,1036,1061]" targetBox="[155,1435,181,1017]" targetPageId="17">
<paragraph id="8BCC36F00F2BFFC4FF19BE2B3605FB06" blockId="17.[151,1437,1036,1205]" pageId="17" pageNumber="18">
<emphasis id="B907EAE20F2BFFC4FF19BE2B3564FB96" bold="true" box="[151,328,1036,1061]" pageId="17" pageNumber="18">FIGURES 914.</emphasis>
Diagnostic characters of
<taxonomicName id="4C734D730F2BFFC4FDC3BE2B372BFB96" authorityName="Shannon" authorityYear="1926" box="[589,775,1036,1061]" class="Insecta" family="Mesembrinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Mesembrinellidae</taxonomicName>
.
<emphasis id="B907EAE20F2BFFC4FC9FBE2B3708FB96" bold="true" box="[785,804,1036,1061]" pageId="17" pageNumber="18">9.</emphasis>
Eggs of
<taxonomicName id="4C734D730F2BFFC4FC0FBE2A3054FB96" authority="Whitworth &amp; Yusseff-Vanegas, 2019" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" box="[897,1144,1037,1061]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fuscosquamata" status="sp. nov.">
<emphasis id="B907EAE20F2BFFC4FC0FBE2A3054FB96" box="[897,1144,1037,1061]" italics="true" pageId="17" pageNumber="18">Laneella fuscosquamata</emphasis>
</taxonomicName>
<emphasis id="B907EAE20F2BFFC4FBF1BE2A30DBFB96" bold="true" box="[1151,1271,1036,1061]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23457990F2BFFC4FBF1BE2A30FCFB96" box="[1151,1232,1037,1061]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
10.
</emphasis>
<taxonomicName id="4C734D730F2BFFC4FB70BE2934EBFBFA" authority="(Mariluis)" authorityName="Mariluis" baseAuthorityName="Mariluis" baseAuthorityYear="1987" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="perisi">
<emphasis id="B907EAE20F2BFFC4FB70BE29317FFB96" box="[1278,1363,1038,1061]" italics="true" pageId="17" pageNumber="18">L. perisi</emphasis>
(Mariluis)
</taxonomicName>
, ventral view of tarsal claws, showing pale basal half.
<emphasis id="B907EAE20F2BFFC4FD71BE173732FBFA" bold="true" box="[767,798,1072,1097]" pageId="17" pageNumber="18">11.</emphasis>
<taxonomicName id="4C734D730F2BFFC4FCABBE15379FFBFA" authority="Whitworth &amp; Yusseff-Vanegas, 2019" authorityName="Whitworth &amp; Yusseff-Vanegas" authorityYear="2019" box="[805,947,1073,1097]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fusconitida" status="sp. nov.">
<emphasis id="B907EAE20F2BFFC4FCABBE15379FFBFA" box="[805,947,1073,1097]" italics="true" pageId="17" pageNumber="18">L. fusconitida</emphasis>
</taxonomicName>
<emphasis id="B907EAE20F2BFFC4FC34BE163020FBFA" bold="true" box="[954,1036,1073,1097]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A23457990F2BFFC4FC34BE163020FBFA" box="[954,1036,1073,1097]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ventral view of tarsal claws, showing darker base.
<emphasis id="B907EAE20F2BFFC4FE96BE733514FBDE" bold="true" box="[280,312,1108,1133]" pageId="17" pageNumber="18">12.</emphasis>
<taxonomicName id="4C734D730F2BFFC4FEB0BE71362CFBDE" authority="Aldrich" authorityName="Aldrich." authorityYear="1925" box="[318,512,1108,1133]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="spicata">
<emphasis id="B907EAE20F2BFFC4FEB0BE713585FBDE" box="[318,425,1110,1133]" italics="true" pageId="17" pageNumber="18">M. spicata</emphasis>
Aldrich
</taxonomicName>
, ventral view of male sternites, showing long projections on rear margin of ST5.
<emphasis id="B907EAE20F2BFFC4FAB6BE733174FBDE" bold="true" box="[1336,1368,1108,1133]" pageId="17" pageNumber="18">13.</emphasis>
<taxonomicName id="4C734D730F2BFFC4FAD0BE713568FB22" authority="(Fabricius)" authorityName="Fabricius" baseAuthorityName="Fabricius" baseAuthorityYear="1805" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B907EAE20F2BFFC4FAD0BE7134E0FB22" italics="true" pageId="17" pageNumber="18">M. bicolor</emphasis>
(Fabricius)
</taxonomicName>
, posterior view of female occiput, showing median occipital sclerite.
<emphasis id="B907EAE20F2BFFC4FB8DBE5F300EFB22" bold="true" box="[1027,1058,1144,1169]" pageId="17" pageNumber="18">14.</emphasis>
<taxonomicName id="4C734D730F2BFFC4FBA6BE5D30C2FB22" authority="(Walker)" authorityName="Walker" baseAuthorityName="Walker" baseAuthorityYear="1861" box="[1064,1262,1144,1169]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="socors">
<emphasis id="B907EAE20F2BFFC4FBA6BE5D30A0FB22" box="[1064,1164,1146,1169]" italics="true" pageId="17" pageNumber="18">M. socors</emphasis>
(Walker)
</taxonomicName>
, upper and lower calypters, showing disc, rim and fringe.
</paragraph>
</caption>
<paragraph id="8BCC36F00F2BFFC4FF19BEC430E2F9D5" blockId="17.[151,1437,1251,1998]" pageId="17" pageNumber="18">
<emphasis id="B907EAE20F2BFFC4FF19BEC4364CFB4D" bold="true" box="[151,608,1251,1278]" pageId="17" pageNumber="18">
Status of genera in
<taxonomicName id="4C734D730F2BFFC4FE0CBEC43677FB4D" authorityName="Shannon" authorityYear="1926" box="[386,603,1251,1278]" class="Insecta" family="Mesembrinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Mesembrinellidae</taxonomicName>
.
</emphasis>
Several studies have questioned the validity of most of the genera in the family (
<bibRefCitation id="EFE24B010F2BFFC4FF7EBF2F35C5FA91" author="Vargas, J. &amp; Wood, D. M." box="[240,489,1287,1314]" pageId="17" pageNumber="18" pagination="1297 - 1304" refId="ref109841" refString="Vargas, J. &amp; Wood, D. M. (2009) Calliphoridae (blow flies). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, N. E. &amp; Zumbado, M. A. (Eds.), Manual of Central American Diptera. Vol. 2. NRC Research Press, Ottawa, pp. 1297 - 1304." type="book chapter" year="2009">
<collectingRegion id="49B7F8120F2BFFC4FF7EBF2F3512FA91" box="[240,318,1288,1314]" country="Venezuela" name="Vargas" pageId="17" pageNumber="18">Vargas</collectingRegion>
&amp; Wood 2009
</bibRefCitation>
;
<bibRefCitation id="EFE24B010F2BFFC4FE78BF20365DFA91" author="Moll, P." box="[502,625,1287,1314]" pageId="17" pageNumber="18" refId="ref109099" refString="Moll, P. (2014) Analise cladistica e biogeografica de Mesembrinellidae (Diptera, Oestroidea). Departamento de Zoologia, Instituto de Biociencias da Unversidade de S &quot; o Paulo, Sao Paulo, 214 pp. [Unpublished M. Sc. thesis]" type="book" year="2014">Moll 2014</bibRefCitation>
;
<bibRefCitation id="EFE24B010F2BFFC4FDF3BF203749FA91" author="Marinho, M. A. T. &amp; Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Azeredo-Espin, A. M. L. de &amp; Amorim, D." box="[637,869,1287,1314]" pageId="17" pageNumber="18" pagination="134 - 152" refId="ref108821" refString="Marinho, M. A. T., Wolff, M., Ramos-Pastrana, Y., Azeredo-Espin, A. M. L. de &amp; Amorim, D. d. S. (2017) The first phylogenetic study of Mesembrinellidae (Diptera: Oestroidea) based on molecular data: clades and congruence with morphological characters. Cladistics, 33 (2), 134 - 152. https: // doi. org / 10.1111 / cla. 1215 7" type="journal article" year="2017">
Marinho
<emphasis id="B907EAE20F2BFFC4FD68BF2F3735FA91" box="[742,793,1288,1314]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. 2017
</bibRefCitation>
). It has even been argued that only the genus
<taxonomicName id="4C734D730F2BFFC4FAE1BF2F3537FAF5" authorityName="Giglio-Tos" authorityYear="1893" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FAE1BF2F3537FAF5" italics="true" pageId="17" pageNumber="18">Mesembrinella</emphasis>
</taxonomicName>
should be retained as valid (
<bibRefCitation id="EFE24B010F2BFFC4FDD2BF0C371BFAF5" author="Cerretti, P. &amp; Stireman, J. O. III. &amp; Pape, T. &amp; O'Hara, J. E. &amp; Marinho, M. A. T. &amp; Rognes, K. &amp; Grimaldi, D. A." box="[604,823,1323,1350]" pageId="17" pageNumber="18" pagination="0182101" refId="ref107732" refString="Cerretti, P., Stireman, J. O. III., Pape, T., O'Hara, J. E., Marinho, M. A. T., Rognes, K. &amp; Grimaldi, D. A. (2017) First fossil of an oestroid fly (Diptera: Calyptratae: Oestroidea) and the dating of oestroid divergences. PLoS ONE, 12 (8), e 0182101. https: // doi. org / 10.1371 / journal. pone. 018210 1" type="journal article" year="2017">
Cerretti
<emphasis id="B907EAE20F2BFFC4FD35BF0B36DFFAF5" box="[699,755,1324,1350]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2017
</bibRefCitation>
). We accept Marinhos synonymy of
<taxonomicName id="4C734D730F2BFFC4FB56BF0B31B0FAF5" authorityName="Townsend" authorityYear="1918" box="[1240,1436,1324,1350]" class="Insecta" family="Calliphoridae" genus="Eumesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FB56BF0B31B0FAF5" box="[1240,1436,1324,1350]" italics="true" pageId="17" pageNumber="18">Eumesembrinella</emphasis>
</taxonomicName>
with
<taxonomicName id="4C734D730F2BFFC4FF5FBF77365AFAD9" authority="(Marinho et al. 2017)" authorityName="Marinho" authorityYear="2017" baseAuthorityName="Marinho et al." baseAuthorityYear="2017" box="[209,630,1359,1386]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FF5FBF773555FAD9" box="[209,377,1360,1386]" italics="true" pageId="17" pageNumber="18">Mesembrinella</emphasis>
(
<bibRefCitation id="EFE24B010F2BFFC4FE06BF683643FAD9" author="Marinho, M. A. T. &amp; Wolff, M. &amp; Ramos-Pastrana, Y. &amp; Azeredo-Espin, A. M. L. de &amp; Amorim, D." box="[392,623,1359,1386]" pageId="17" pageNumber="18" pagination="134 - 152" refId="ref108821" refString="Marinho, M. A. T., Wolff, M., Ramos-Pastrana, Y., Azeredo-Espin, A. M. L. de &amp; Amorim, D. d. S. (2017) The first phylogenetic study of Mesembrinellidae (Diptera: Oestroidea) based on molecular data: clades and congruence with morphological characters. Cladistics, 33 (2), 134 - 152. https: // doi. org / 10.1111 / cla. 1215 7" type="journal article" year="2017">
Marinho
<emphasis id="B907EAE20F2BFFC4FE7FBF77360FFAD9" box="[497,547,1360,1386]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. 2017
</bibRefCitation>
)
</taxonomicName>
. Based on our study of specimens of most previously described species and of the fifteen new species described herein, we retain three subfamilies, Laneellinae,
<taxonomicName id="4C734D730F2BFFC4FB2EBF543146FA3E" authorityName="Shannon" authorityYear="1926" box="[1184,1386,1395,1421]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Mesembrinellinae">Mesembrinellinae</taxonomicName>
and Souzalopesiellinae, and three genera,
<taxonomicName id="4C734D730F2BFFC4FDC5BFBF3681FA01" authorityName=", Laneellinae" box="[587,685,1432,1458]" class="Insecta" family="Calliphoridae" genus="Laneella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FDC5BFBF3681FA01" box="[587,685,1432,1458]" italics="true" pageId="17" pageNumber="18">Laneella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C734D730F2BFFC4FD31BFBF374BFA01" baseAuthorityName="Marinho et al." baseAuthorityYear="2017" box="[703,871,1432,1458]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FD31BFBF374BFA01" box="[703,871,1432,1458]" italics="true" pageId="17" pageNumber="18">Mesembrinella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D730F2BFFC4FC2ABFBF3078FA01" authorityName=", Souzalopesiellinae" box="[932,1108,1432,1458]" class="Insecta" family="Mesembrinellidae" genus="Souzalopesiella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FC2ABFBF3078FA01" box="[932,1108,1432,1458]" italics="true" pageId="17" pageNumber="18">Souzalopesiella</emphasis>
</taxonomicName>
. We synonymize the genera
<taxonomicName id="4C734D730F2BFFC4FF19BF9B3519FA65" authorityName="Mello" authorityYear="1967" box="[151,309,1468,1494]" class="Insecta" family="Calliphoridae" genus="Albuquerquea" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FF19BF9B3519FA65" box="[151,309,1468,1494]" italics="true" pageId="17" pageNumber="18">Albuquerquea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C734D730F2BFFC4FECBBF9B35EDFA65" authorityName="Bonatto in Bonatto &amp; Marinoni" authorityYear="2005" box="[325,449,1468,1494]" class="Insecta" family="Calliphoridae" genus="Giovanella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FECBBF9B35EDFA65" box="[325,449,1468,1494]" italics="true" pageId="17" pageNumber="18">Giovanella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C734D730F2BFFC4FE5FBF9B3674FA65" authorityName="Bonatto in Bonatto &amp; Marinoni" authorityYear="2005" box="[465,600,1468,1494]" class="Insecta" family="Calliphoridae" genus="Henriquella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FE5FBF9B3674FA65" box="[465,600,1468,1494]" italics="true" pageId="17" pageNumber="18">Henriquella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C734D730F2BFFC4FDE6BF9B3731FA66" authorityName="Townsend" authorityYear="1931" box="[616,797,1468,1493]" class="Insecta" family="Calliphoridae" genus="Huascaromusca" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FDE6BF9B3731FA66" box="[616,797,1468,1493]" italics="true" pageId="17" pageNumber="18">Huascaromusca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D730F2BFFC4FCD6BF9B37D2FA65" authorityName="Guimar" box="[856,1022,1468,1494]" class="Insecta" family="Tachinidae" genus="Thompsoniella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FCD6BF9B37D2FA65" box="[856,1022,1468,1494]" italics="true" pageId="17" pageNumber="18">Thompsoniella</emphasis>
</taxonomicName>
with
<taxonomicName id="4C734D730F2BFFC4FBCCBF9B30C6FA65" baseAuthorityName="Marinho et al." baseAuthorityYear="2017" box="[1090,1258,1468,1494]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FBCCBF9B30C6FA65" box="[1090,1258,1468,1494]" italics="true" pageId="17" pageNumber="18">Mesembrinella</emphasis>
</taxonomicName>
, and species of
<taxonomicName id="4C734D730F2BFFC4FF19BFC73513FA49" baseAuthorityName="Marinho et al." baseAuthorityYear="2017" box="[151,319,1504,1530]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAE20F2BFFC4FF19BFC73513FA49" box="[151,319,1504,1530]" italics="true" pageId="17" pageNumber="18">Mesembrinella</emphasis>
</taxonomicName>
are classified within six groups: the
<taxonomicName id="4C734D730F2BFFC4FD57BFC73770FA49" authorityName="Mello" authorityYear="1967" box="[729,860,1504,1530]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B907EAE20F2BFFC4FD57BFC736DCFA4A" box="[729,752,1504,1529]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="B907EAE20F2BFFC4FD70BFC73770FA49" box="[766,860,1504,1530]" italics="true" pageId="17" pageNumber="18">latifrons</emphasis>
</taxonomicName>
-group, the
<taxonomicName id="4C734D730F2BFFC4FC53BFC7307FFA4A" authorityName="Aldrich." authorityYear="1925" box="[989,1107,1504,1530]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="spicata">
<emphasis id="B907EAE20F2BFFC4FC53BFC737D8FA4A" box="[989,1012,1504,1529]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="B907EAE20F2BFFC4FB8CBFC7307FFA4A" box="[1026,1107,1504,1529]" italics="true" pageId="17" pageNumber="18">spicata</emphasis>
</taxonomicName>
-group, the
<taxonomicName id="4C734D730F2BFFC4FB5BBFC73167FA49" baseAuthorityName="Bonatto" box="[1237,1355,1504,1530]" class="Insecta" family="Calliphoridae" genus="Giovanella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bolivar">
<emphasis id="B907EAE20F2BFFC4FB5BBFC730C0FA4A" box="[1237,1260,1504,1529]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="B907EAE20F2BFFC4FB74BFC73167FA49" box="[1274,1355,1504,1530]" italics="true" pageId="17" pageNumber="18">bolivar</emphasis>
</taxonomicName>
-group, the
<taxonomicName id="4C734D730F2BFFC4FF4FBC23355CF9AE" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[193,368,1540,1566]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="aeneiventris">
<emphasis id="B907EAE20F2BFFC4FF4FBC2334F4F9AE" box="[193,216,1540,1565]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="B907EAE20F2BFFC4FF69BC23355CF9AE" box="[231,368,1540,1565]" italics="true" pageId="17" pageNumber="18">aeneiventris</emphasis>
</taxonomicName>
-group, the
<taxonomicName id="4C734D730F2BFFC4FE7CBC233645F9AD" baseAuthorityName="Fabricius" baseAuthorityYear="1805" box="[498,617,1540,1566]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B907EAE20F2BFFC4FE7CBC233625F9AE" box="[498,521,1540,1565]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="B907EAE20F2BFFC4FD96BC233645F9AD" box="[536,617,1540,1566]" italics="true" pageId="17" pageNumber="18">bicolor</emphasis>
</taxonomicName>
-group, and the
<taxonomicName id="4C734D730F2BFFC4FC95BC23378FF9AD" authorityName="Guimar" box="[795,931,1540,1566]" class="Insecta" family="Calliphoridae" genus="Mesembrinella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B907EAE20F2BFFC4FC95BC23371EF9AE" box="[795,818,1540,1565]" italics="true" pageId="17" pageNumber="18">M</emphasis>
.
<emphasis id="B907EAE20F2BFFC4FCCFBC23378FF9AD" box="[833,931,1540,1566]" italics="true" pageId="17" pageNumber="18">anomala</emphasis>
</taxonomicName>
-group. These groupings help us discuss species in a logical manner, but they are tentative and we are aware that future phylogenetic studies may suggest further changes in the classification. Characters of each species-group are discussed in detail further on.
</paragraph>
<paragraph id="8BCC36F00F2BFFC4FF49BC573753F836" blockId="17.[151,1437,1251,1998]" pageId="17" pageNumber="18">
<emphasis id="B907EAE20F2BFFC4FF49BC573517F939" bold="true" box="[199,315,1648,1674]" pageId="17" pageNumber="18">Remarks.</emphasis>
The detailed key below can be used to separate
<taxonomicName id="4C734D730F2BFFC4FCEEBC483006F93A" authorityName="Shannon" authorityYear="1926" box="[864,1066,1647,1673]" class="Insecta" family="Mesembrinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Mesembrinellidae</taxonomicName>
from Neotropical
<taxonomicName id="4C734D730F2BFFC4FB73BC4831B5F93A" authorityName="Brauer &amp; von Bergenstamm" authorityYear="1889" box="[1277,1433,1647,1673]" class="Insecta" family="Calliphoridae" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Calliphoridae</taxonomicName>
. The reniform posterior spiracle readily distinguishes mesembrinellids from calliphorids, sarcophagids and tachinids. Other features of sarcophagids are the three conspicuous black stripes on a gray- to gold-pruinose thorax and the checkered abdomen. Sarcophagids typically also have two larger and two smaller notopleural setae on the notopleuron, whereas mesembrinellids have only two stout notopleural setae. Tachinids have a prominently developed subscutellum, rare in mesembrinellids, and normally they have a bare arista (except in Dexiini), which is setose in mesembrinellids. Other similar flies include muscids, which sometimes have a posterior spiracle similar to mesembrinellids but lack meral setae, always present in mesembrinellids.
</paragraph>
<paragraph id="8BCC36F00F2BFFC4FF49BDA83140F87E" blockId="17.[151,1437,1251,1998]" pageId="17" pageNumber="18">
The keys to genera and species of
<taxonomicName id="4C734D730F2BFFC4FDD3BDA8370BF81A" authorityName="Shannon" authorityYear="1926" box="[605,807,1935,1961]" class="Insecta" family="Mesembrinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Mesembrinellidae</taxonomicName>
provided herein were developed following Guimar„es (1977), and are the first comprehensive taxonomic keys for the family
<taxonomicName id="4C734D730F2BFFC4FC21BD943055F87E" authorityName="Shannon" authorityYear="1926" box="[943,1145,1971,1997]" class="Insecta" family="Mesembrinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Mesembrinellidae</taxonomicName>
published since then.
</paragraph>
</subSubSection>
</treatment>
</document>