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<mods:title id="AEF5F3568607F559328E173C0ED986D3">Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)</mods:title>
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<treatment id="03DA87C4FF9C1772FF4C6FBFAE60FD25" ID-DOI="http://doi.org/10.5281/zenodo.6109860" ID-GBIF-Taxon="119596525" ID-Zenodo-Dep="6109860" LSID="urn:lsid:plazi:treatment:03DA87C4FF9C1772FF4C6FBFAE60FD25" httpUri="http://treatment.plazi.org/id/03DA87C4FF9C1772FF4C6FBFAE60FD25" lastPageId="21" lastPageNumber="132" pageId="20" pageNumber="131">
<subSubSection id="C3696559FF9C1773FF4C6FBFABE8FA75" box="[151,500,1502,1528]" pageId="20" pageNumber="131" type="nomenclature">
<paragraph id="8BCC36D2FF9C1773FF4C6FBFABE8FA75" blockId="20.[151,500,1502,1528]" box="[151,500,1502,1528]" pageId="20" pageNumber="131">
<heading id="D08481BEFF9C1773FF4C6FBFABE8FA75" bold="true" box="[151,500,1502,1528]" fontSize="11" level="1" pageId="20" pageNumber="131" reason="1">
<emphasis id="B907EAC0FF9C1773FF4C6FBFABE8FA75" bold="true" box="[151,500,1502,1528]" pageId="20" pageNumber="131">
Genus
<taxonomicName id="4C734D51FF9C1773FF356FBFABE8FA75" ID-CoL="35GS" authority="Kramer, 1875" authorityName="Kramer" authorityYear="1875" box="[238,500,1502,1528]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9C1773FF356FBFAB5DFA75" bold="true" box="[238,321,1502,1528]" italics="true" pageId="20" pageNumber="131">Aturus</emphasis>
Kramer, 1875
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF9C1773FF4C6C47A87AF89F" pageId="20" pageNumber="131" type="reference_group">
<paragraph id="8BCC36D2FF9C1773FF4C6C47A829F9F5" blockId="20.[151,907,1573,1810]" pageId="20" pageNumber="131">
<taxonomicName id="4C734D51FF9C1773FF4C6C47AB70F9B6" ID-CoL="35GS" authority="Cook, 1974" authorityName="Cook" authorityYear="1974" box="[151,364,1573,1595]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9C1773FF4C6C47AAC6F9B6" box="[151,218,1574,1595]" italics="true" pageId="20" pageNumber="131">Aturus</emphasis>
:
<bibRefCitation id="EFE24B23FF9C1773FF336C44AB70F9B6" author="Cook" box="[232,364,1573,1595]" pageId="20" pageNumber="131" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook, 1974a</bibRefCitation>
</taxonomicName>
, pp. 358359, figs. 1558, 1559,
<date id="FFCD1012FF9C1773FD696C44A93BF9B6" box="[690,807,1573,1595]" pageId="20" pageNumber="131" value="1561" valueMax="1564">15611564</date>
.
<taxonomicName id="4C734D51FF9C1773FF4C6C25AB7CF9D7" ID-CoL="35GS" authority="Cook, 1980" authorityName="Cook" authorityYear="1980" box="[151,352,1604,1626]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9C1773FF4C6C25AAC6F9D4" box="[151,218,1604,1625]" italics="true" pageId="20" pageNumber="131">Aturus</emphasis>
:
<bibRefCitation id="EFE24B23FF9C1773FF336C25AB7CF9D7" author="Cook" box="[232,352,1604,1626]" pageId="20" pageNumber="131" refString="Cook, D. R. (1980) Studies on Neotropical water mites. Memoirs of the American Entomological Institute, 31, 1 - 645." type="journal article" year="1980">Cook, 1980</bibRefCitation>
</taxonomicName>
, pp. 292293, figs. 1508,
<date id="FFCD1012FF9C1773FDB36C25A8C0F9D7" box="[616,732,1604,1626]" pageId="20" pageNumber="131" value="1511" valueMax="1516">15111516</date>
.
<taxonomicName id="4C734D51FF9C1773FF4C6C02AB7AF9F5" ID-CoL="35GS" authority="Smith, 1984" authorityName="Smith" authorityYear="1984" box="[151,358,1634,1656]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9C1773FF4C6C02AAC6F9F5" box="[151,218,1635,1656]" italics="true" pageId="20" pageNumber="131">Aturus</emphasis>
: Smith, 1984
</taxonomicName>
, p. 321, figs. 6372.
</paragraph>
<paragraph id="8BCC36D2FF9C1773FF4C6CE3A839F91A" blockId="20.[151,907,1573,1810]" box="[151,549,1665,1687]" pageId="20" pageNumber="131">
<taxonomicName id="4C734D51FF9C1773FF4C6CE3AB89F91A" ID-CoL="35GS" authority="K.O. Viets, 1987" authorityName="K.O. Viets" authorityYear="1987" box="[151,405,1665,1687]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9C1773FF4C6CE3AAC6F91A" box="[151,218,1666,1687]" italics="true" pageId="20" pageNumber="131">Aturus</emphasis>
:
<bibRefCitation id="EFE24B23FF9C1773FF336CE0AB89F91A" author="Viets" box="[232,405,1665,1687]" pageId="20" pageNumber="131" refString="Viets, K. O. (1987) Die Milben des Sußwassers (Hydrachnellae und Halacaridae [part.], Acari). 2. Katalog. Sonderbande des Naturwissenschaftlichen Vereins in Hamburg, 8, 1 - 1012." type="journal article" year="1987">K.O. Viets, 1987</bibRefCitation>
</taxonomicName>
, pp. 164177.
</paragraph>
<paragraph id="8BCC36D2FF9C1773FF4C6CC1A87AF89F" blockId="20.[151,907,1573,1810]" pageId="20" pageNumber="131">
Aturinae:
<bibRefCitation id="EFE24B23FF9C1773FF266CC1ABCEF93B" author="Smith" box="[253,466,1696,1718]" pageId="20" pageNumber="131" refString="Smith, I. M. &amp; Cook, D. R. (1991) Water mites. Chapter 16. In: Thorp, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. Academic Press, San Diego, CA, pp. 523 - 592. [total page number 911 pp.]" type="book chapter" year="1991">Smith &amp; Cook, 1991</bibRefCitation>
, pp. 553, 577, figs. 16.133, 16.136, 16.268.
<taxonomicName id="4C734D51FF9C1773FF4C6CDEAB83F959" ID-CoL="35GS" authority="Smith et al., 2001" authorityName="Smith et al." authorityYear="2001" box="[151,415,1726,1749]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9C1773FF4C6CDEAAC6F959" box="[151,218,1727,1748]" italics="true" pageId="20" pageNumber="131">Aturus</emphasis>
:
<bibRefCitation id="EFE24B23FF9C1773FF336CDFAB83F959" author="Smith" box="[232,415,1726,1749]" pageId="20" pageNumber="131" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2001) Water mites and other arachnids. Chapter 16. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 2 nd Edition. Academic Press, Waltham, Massachusetts, pp. 551 - 659. [1056 pp.] http: // dx. doi. org / 10.1016 / B 978 - 012690647 - 9 / 50017 - X" type="book chapter" year="2001">
Smith
<emphasis id="B907EAC0FF9C1773FEF06CA1AB4BF959" box="[299,343,1726,1749]" italics="true" pageId="20" pageNumber="131">et al</emphasis>
., 2001
</bibRefCitation>
</taxonomicName>
, pp. 580, 613, figs. 168170, 377.
<taxonomicName id="4C734D51FF9C1773FF4C6CBFAB83F97E" ID-CoL="35GS" authority="Smith et al., 2010" authorityName="Smith et al." authorityYear="2010" box="[151,415,1757,1779]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9C1773FF4C6CBFAAC6F97E" box="[151,218,1758,1779]" italics="true" pageId="20" pageNumber="131">Aturus</emphasis>
:
<bibRefCitation id="EFE24B23FF9C1773FF336CBCAB83F97E" author="Smith" box="[232,415,1757,1779]" pageId="20" pageNumber="131" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2010) Water mites and other arachnids. Chapter 15. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 3 rd Edition. Academic Press, Waltham, Massachusetts, pp. 485 - 586. [total page number: 1021 pp.] http: // dx. doi. org / 10.1016 / B 978 - 0 - 12 - 374855 - 3.00015 - 7" type="book chapter" year="2010">
Smith
<emphasis id="B907EAC0FF9C1773FEF06CBFAB4BF97E" box="[299,343,1757,1779]" italics="true" pageId="20" pageNumber="131">et al</emphasis>
., 2010
</bibRefCitation>
</taxonomicName>
, pp. 523, 554, figs. 15 16815.170, 15.378.
<taxonomicName id="4C734D51FF9C1773FF4C6C9DAB97F89F" ID-CoL="35GS" authority="Gerecke, 2014" authorityName="Gerecke" authorityYear="2014" box="[151,395,1788,1810]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9C1773FF4C6C9DAAC6F89C" box="[151,218,1788,1809]" italics="true" pageId="20" pageNumber="131">Aturus</emphasis>
:
<bibRefCitation id="EFE24B23FF9C1773FF336C9DAB97F89F" author="Gerecke" box="[232,395,1788,1810]" pageId="20" pageNumber="131" refString="Gerecke, R. (2014 b) Studies on European species of the water mite family Aturidae (Acari: Hydrachnidia). Zootaxa, 3841 (1), 1 - 46. http: // dx. doi. org / 10.11646 / zootaxa. 3841.1.1" type="journal article" year="2014" yearSuffix="b">Gerecke, 2014b</bibRefCitation>
</taxonomicName>
, pp. 122, figs. 115.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF9C1772FF4C6D5FA8AFFE78" lastPageId="21" lastPageNumber="132" pageId="20" pageNumber="131" type="diagnosis">
<paragraph id="8BCC36D2FF9C1773FF4C6D5FAAF0F8F6" blockId="20.[151,1436,1854,2023]" pageId="20" pageNumber="131">
<emphasis id="B907EAC0FF9C1773FF4C6D5FAB43F8DB" bold="true" box="[151,351,1854,1879]" pageId="20" pageNumber="131">
Diagnosis.
<emphasis id="B907EAC0FF9C1773FEC06D5EAB43F8DB" bold="true" box="[283,351,1855,1878]" italics="true" pageId="20" pageNumber="131">Larva</emphasis>
</emphasis>
(modified from Smith 1984): Character states of subfamily Aturinae (only genus with known larvae).
</paragraph>
<paragraph id="8BCC36D2FF9C1772FF1C6DE7A8AFFE78" blockId="20.[151,1436,1854,2023]" lastBlockId="21.[151,1437,151,680]" lastPageId="21" lastPageNumber="132" pageId="20" pageNumber="131">
<emphasis id="B907EAC0FF9C1773FF1C6DE7AB0EF812" bold="true" box="[199,274,1926,1951]" italics="true" pageId="20" pageNumber="131">Adults</emphasis>
(modified from
<bibRefCitation id="EFE24B23FF9C1773FE036DE7A879F812" author="Cook" box="[472,613,1926,1951]" pageId="20" pageNumber="131" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook 1974a</bibRefCitation>
): Idiosoma elliptical in shape in males, elongate elliptical in shape in females. Idiosomal sclerites with surfaces smooth or finely reticulate. Dorsal and ventral shields separate from one another or fused posteriorly in males, separate from one another in females. Dorsal shield bearing four pairs of glandularia; lacking or bearing one or more pairs of well-defined longitudinal ridges in males, lacking ridges in females; posterior region of shield variously modified and bearing modified setae in males, unmodified in females. Dorsal furrow bearing four pairs of glandularia, one pair of setae, and three or four pairs of lyrifissures arranged on four pairs of platelets. Fourth coxal plate lacking coxoglandularium II and lacking small projection associated with opening for insertion of fourth leg. Ventral shield bearing two pairs of glandularia posteriorly. Genital field terminal and bearing ten to many pairs of acetabula; excretory pore incorporated into dorsal shield; acetabular plates fused with ventral shield with suture lines obliterated. Third leg of males distal segments bearing or lacking modified setae. Fourth leg of males with distal segments moderately to highly modified and bearing highly modified setae. Pedipalp with tibia moderately long and slender, typically lacking a ventral projection and bearing two or three sessile, slender setae distoventrally.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF9D1772FF1C6B9EA96CFDB1" pageId="21" pageNumber="132" type="materials_examined">
<paragraph id="8BCC36D2FF9D1772FF1C6B9EA874FD9A" blockId="21.[151,1437,151,680]" box="[199,616,511,536]" pageId="21" pageNumber="132">
<emphasis id="B907EAC0FF9D1772FF1C6B9EAB7EFD95" bold="true" box="[199,354,511,536]" pageId="21" pageNumber="132">
<typeStatus id="54C88870FF9D1772FF1C6B9EAB1EFD95" box="[199,258,511,536]" pageId="21" pageNumber="132">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C734D51FF9D1772FEB26861A874FD9A" authority="Kramer." authorityName="Kramer." box="[361,616,511,536]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="132" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="B907EAC0FF9D1772FEB26861A819FD95" box="[361,517,511,536]" italics="true" pageId="21" pageNumber="132">Aturus scaber</emphasis>
Kramer.
</taxonomicName>
</paragraph>
<paragraph id="8BCC36D2FF9D1772FF1C6845A96CFDB1" blockId="21.[151,1437,151,680]" box="[199,880,548,573]" pageId="21" pageNumber="132">
<emphasis id="B907EAC0FF9D1772FF1C6845AB8FFDB0" bold="true" box="[199,403,548,573]" pageId="21" pageNumber="132">Species included.</emphasis>
Many species listed by
<bibRefCitation id="EFE24B23FF9D1772FD7B6845A971FDB1" author="Viets" box="[672,877,548,573]" pageId="21" pageNumber="132" refString="Viets, K. O. (1987) Die Milben des Sußwassers (Hydrachnellae und Halacaridae [part.], Acari). 2. Katalog. Sonderbande des Naturwissenschaftlichen Vereins in Hamburg, 8, 1 - 1012." type="journal article" year="1987">K.O. Viets (1987)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF9D1772FF1C6826AF18FDED" box="[199,1284,583,608]" pageId="21" pageNumber="132" type="distribution">
<paragraph id="8BCC36D2FF9D1772FF1C6826AF18FDED" blockId="21.[151,1437,151,680]" box="[199,1284,583,608]" pageId="21" pageNumber="132">
<emphasis id="B907EAC0FF9D1772FF1C6826AB7CFDED" bold="true" box="[199,352,583,608]" pageId="21" pageNumber="132">Distribution.</emphasis>
Holarctic (Eurasia, North
<collectingCountry id="F3647642FF9D1772FD516826A8F3FDED" box="[650,751,583,608]" name="United States of America" pageId="21" pageNumber="132">America</collectingCountry>
), Africa,
<collectingCountry id="F3647642FF9D1772FC8C6826A98FFDED" box="[855,915,583,608]" name="India" pageId="21" pageNumber="132">India</collectingCountry>
, Southeast Asia, South
<collectingCountry id="F3647642FF9D1772FB476826AF1CFDED" box="[1180,1280,583,608]" name="United States of America" pageId="21" pageNumber="132">America</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF9D1772FF1C680DAE60FD25" pageId="21" pageNumber="132" type="discussion">
<paragraph id="8BCC36D2FF9D1772FF1C680DAE60FD25" blockId="21.[151,1437,151,680]" pageId="21" pageNumber="132">
<emphasis id="B907EAC0FF9D1772FF1C680DAB50FD08" bold="true" box="[199,332,620,645]" pageId="21" pageNumber="132">Discussion.</emphasis>
<bibRefCitation id="EFE24B23FF9D1772FE8F680DABECFD09" author="Cook" box="[340,496,620,645]" pageId="21" pageNumber="132" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook (1974a)</bibRefCitation>
considered
<taxonomicName id="4C734D51FF9D1772FDA2680DA8D8FD0E" box="[633,708,620,643]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF9D1772FDA2680DA8D8FD0E" box="[633,708,620,643]" italics="true" pageId="21" pageNumber="132">Aturus</emphasis>
</taxonomicName>
to be a distinct genus. This treatment was followed by
<bibRefCitation id="EFE24B23FF9D1772FAE0680DAB34FD25" author="Smith" pageId="21" pageNumber="132" refString="Smith, I. M. &amp; Cook, D. R. (1991) Water mites. Chapter 16. In: Thorp, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. Academic Press, San Diego, CA, pp. 523 - 592. [total page number 911 pp.]" type="book chapter" year="1991">Smith &amp; Cook (1991)</bibRefCitation>
,
<bibRefCitation id="EFE24B23FF9D1772FEE868EEA81CFD25" author="Smith" box="[307,512,655,680]" pageId="21" pageNumber="132" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2001) Water mites and other arachnids. Chapter 16. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 2 nd Edition. Academic Press, Waltham, Massachusetts, pp. 551 - 659. [1056 pp.] http: // dx. doi. org / 10.1016 / B 978 - 012690647 - 9 / 50017 - X" type="book chapter" year="2001">
Smith
<emphasis id="B907EAC0FF9D1772FEA668F0ABB1FD25" box="[381,429,655,680]" italics="true" pageId="21" pageNumber="132">et al</emphasis>
. (2001
</bibRefCitation>
, 2010), and
<bibRefCitation id="EFE24B23FF9D1772FD5168EEA95BFD25" author="Gerecke" box="[650,839,655,680]" pageId="21" pageNumber="132" refString="Gerecke, R. (2014 b) Studies on European species of the water mite family Aturidae (Acari: Hydrachnidia). Zootaxa, 3841 (1), 1 - 46. http: // dx. doi. org / 10.11646 / zootaxa. 3841.1.1" type="journal article" year="2014" yearSuffix="b">Gerecke (2014b)</bibRefCitation>
and is followed by us here.
</paragraph>
</subSubSection>
</treatment>
</document>