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<mods:title id="80934849BCF939D3D79A45DBB07D1D8D">Stabilisation of some names of European butterflies (Lepidoptera: Pieridae) in their prevailing usage</mods:title>
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<mods:affiliation id="D83AD16A38539E2F22165AC30D973875">Department of Life Sciences and Systems Biology, University of Turin, Italy. &amp; francesca. barbero @ unito. it; http: // orcid. org / 0000 - 0003 - 2667 - 0435</mods:affiliation>
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<mods:affiliation id="E14A0B6F203C67499948C04B305FC718">Department of Life Sciences and Systems Biology, University of Turin, Italy. &amp; Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw, Poland.</mods:affiliation>
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<treatment id="03DA87B0FF808C3FBFE5D596FAE1F975" ID-DOI="http://doi.org/10.5281/zenodo.3854383" ID-GBIF-Taxon="164281503" ID-Zenodo-Dep="3854383" LSID="urn:lsid:plazi:treatment:03DA87B0FF808C3FBFE5D596FAE1F975" httpUri="http://treatment.plazi.org/id/03DA87B0FF808C3FBFE5D596FAE1F975" lastPageNumber="391" pageId="4" pageNumber="391">
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<paragraph id="8BCC36A6FF808C3FBFE5D596FD90FD45" blockId="4.[151,512,657,684]" box="[151,512,657,684]" pageId="4" pageNumber="391">
<heading id="D08481CAFF808C3FBFE5D596FD90FD45" bold="true" box="[151,512,657,684]" fontSize="11" level="1" pageId="4" pageNumber="391" reason="1">
<taxonomicName id="4C734D25FF808C3FBFE5D596FD90FD45" ID-CoL="3BT39" authority="Butler, 1869" authorityName="Butler" authorityYear="1869" box="[151,512,657,684]" class="Insecta" family="Pieridae" genus="Euchloe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="crameri">
<emphasis id="B907EAB4FF808C3FBFE5D596FD90FD45" bold="true" box="[151,512,657,684]" pageId="4" pageNumber="391">
<emphasis id="B907EAB4FF808C3FBFE5D596FEF3FD45" bold="true" box="[151,355,657,684]" italics="true" pageId="4" pageNumber="391">Euchloe crameri</emphasis>
<bibRefCitation id="EFE24B57FF808C3FBE1BD596FD90FD45" author="Butler, A. G." box="[361,512,657,683]" pageId="4" pageNumber="391" pagination="270 - 273" refId="ref5399" refString="Butler, A. G. (1869) Descriptions of species of Lepidoptera, confounded with others described by Linnaeus and Fabricius. The Entomologist's monthly Magazine, 5, 270 - 273." type="journal article" year="1869">Butler, 1869</bibRefCitation>
</emphasis>
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<paragraph id="8BCC36A6FF808C3FBFE5D5DEFDFEFCD5" blockId="4.[151,1437,728,1691]" pageId="4" pageNumber="391">
(Entomol. month. Mag., 5 (59): 271) was published as a new name for the taxon that Cramer [recte Stoll], [1782] (Uitl. Kapell., 4: 225, Pl. 397, Figs. A, B) misidentified as
<taxonomicName id="4C734D25FF808C3FBC5ED5FAFBE3FCF9" authority="Linnaeus, 1767" authorityName="Linnaeus. Therefore" authorityYear="1767" box="[812,1139,765,791]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="belia">
<emphasis id="B907EAB4FF808C3FBC5ED5FAFC2EFCF9" box="[812,958,765,791]" italics="true" pageId="4" pageNumber="391">Papilio belia</emphasis>
<bibRefCitation id="EFE24B57FF808C3FBCB7D5FAFBE3FCF9" author="Linnaeus, C." box="[965,1139,765,791]" pageId="4" pageNumber="391" pagination="533 - 1327" refId="ref7097" refString="Linnaeus, C. (1767) Caroli Linnaei Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 1 (2). 12 th Edition. Laurentii Salvii, Holmiae, 795 pp. [pp. 533 - 1327 + xxxvii (Index, Appendix)] https: // doi. org / 10.5962 / bhl. title. 157601" type="book chapter" year="1767">Linnaeus, 1767</bibRefCitation>
</taxonomicName>
(Syst. Nat., ed. XII, 1 (2): 761, no. 84. Currently
<taxonomicName id="4C734D25FF808C3FBEE7D426FDCDFCD5" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[405,605,801,827]" class="Insecta" family="Pieridae" genus="Anthocharis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="belia">
<emphasis id="B907EAB4FF808C3FBEE7D426FDCDFCD5" box="[405,605,801,827]" italics="true" pageId="4" pageNumber="391">Anthocharis belia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C369652DFF808C3FBFB5D442FAE1F975" pageId="4" pageNumber="391" type="description">
<paragraph id="8BCC36A6FF808C3FBFB5D442FB19FC25" blockId="4.[151,1437,728,1691]" pageId="4" pageNumber="391">
The latter name was extensively used even in relatively recent times, Stoll declaredly did not mean to describe a new species, since he supposed that the specimen depicted was a female of
<taxonomicName id="4C734D25FF808C3FBB66D46EFA08FC6D" authority="Linnaeus. Therefore" authorityName="Linnaeus. Therefore" authorityYear="1767" box="[1044,1432,873,899]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="belia">
<emphasis id="B907EAB4FF808C3FBB66D46EFB38FC6D" box="[1044,1192,873,899]" italics="true" pageId="4" pageNumber="391">Papilio belia</emphasis>
Linnaeus. Therefore
</taxonomicName>
,
<taxonomicName id="4C734D25FF808C3FBFD3D48AFEA5FC49" authorityName="Linnaeus. Therefore" authorityYear="1767" box="[161,309,909,935]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="belia">
<emphasis id="B907EAB4FF808C3FBFD3D48AFEA5FC49" box="[161,309,909,935]" italics="true" pageId="4" pageNumber="391">Papilio belia</emphasis>
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Stoll has no nomenclatural status and is, accordingly, unavailable. No potential Stolls specimen identified as
<taxonomicName id="4C734D25FF808C3FBE43D4B6FE53FC25" authorityName="Linnaeus. Therefore" authorityYear="1767" box="[305,451,945,971]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="belia">
<emphasis id="B907EAB4FF808C3FBE43D4B6FE53FC25" box="[305,451,945,971]" italics="true" pageId="4" pageNumber="391">Papilio belia</emphasis>
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survives in NHMUK (
<bibRefCitation id="EFE24B57FF808C3FBDB8D4B6FC31FC25" author="Chainey, J. E." box="[714,929,944,971]" pageId="4" pageNumber="391" pagination="283 - 337" refId="ref5434" refString="Chainey, J. E. (2005) The species of Papilionidae and Pieridae (Lepidoptera) described by Cramer and Stoll and their putative type material in the Natural History Museum in London. Zoological Journal of the Linnean Society, 145 (3), 283 - 337. https: // doi. org / 10.1111 / j. 1096 - 3642.2005.00184. x" type="journal article" year="2005">Chainey 2005: 316</bibRefCitation>
and D. Lees
<emphasis id="B907EAB4FF808C3FBB44D4B5FBE9FC25" box="[1078,1145,945,971]" italics="true" pageId="4" pageNumber="391">in litt.</emphasis>
).
</paragraph>
<paragraph id="8BCC36A6FF808C3FBFB5D4D2FB96FB05" blockId="4.[151,1437,728,1691]" pageId="4" pageNumber="391">
Stoll rather unequivocally stated that his specimen had been collected at Smyrna. However, the same species (i.e.
<taxonomicName id="4C734D25FF808C3FBFB8D4FEFD82FBFA" authority="Linnaeus, 1767" authorityName="Linnaeus. Therefore" authorityYear="1767" box="[202,530,1017,1044]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="belia">
<emphasis id="B907EAB4FF808C3FBFB8D4FEFECCFBFD" box="[202,348,1017,1043]" italics="true" pageId="4" pageNumber="391">Papilio belia</emphasis>
<bibRefCitation id="EFE24B57FF808C3FBE16D4FEFD82FBFA" author="Linnaeus, C." box="[356,530,1017,1044]" pageId="4" pageNumber="391" pagination="533 - 1327" refId="ref7097" refString="Linnaeus, C. (1767) Caroli Linnaei Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 1 (2). 12 th Edition. Laurentii Salvii, Holmiae, 795 pp. [pp. 533 - 1327 + xxxvii (Index, Appendix)] https: // doi. org / 10.5962 / bhl. title. 157601" type="book chapter" year="1767">Linnaeus, 1767</bibRefCitation>
</taxonomicName>
) was said to occur also on the Coast of Barbaria (i.e. North Africa), in Provence and in Languedoc (“Zy is te Smirna gevangen, doch wordt ook op de Kust van Barbaryen, als mede in het zuidelyke gedeelte van Frankryk, in Provence in Languedocq gevonden”—and “Elle a été prise à Smyrne, mais on la trouve pareillement sur la Côte de Barbarie &amp; dans la partie méridionale de la
<collectingCountry id="F3647636FF808C3FBCCED362FB9BFB91" box="[956,1035,1125,1151]" name="France" pageId="4" pageNumber="391">France</collectingCountry>
, Provence &amp; au Languedoc”, in the facing French text). However, several authors assumed that the
<typeStatus id="54C88804FF808C3FBC06D38EFC34FB4D" box="[884,932,1161,1187]" pageId="4" pageNumber="391">type</typeStatus>
locality of
<taxonomicName id="4C734D25FF808C3FBB5ED38EFA10FB4D" authority="Butler, 1869" authorityName="Butler" authorityYear="1869" box="[1068,1408,1161,1187]" class="Insecta" family="Pieridae" genus="Euchloe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="crameri">
<emphasis id="B907EAB4FF808C3FBB5ED38EFB7AFB4D" box="[1068,1258,1161,1187]" italics="true" pageId="4" pageNumber="391">Euchloe crameri</emphasis>
<bibRefCitation id="EFE24B57FF808C3FBB81D38EFA10FB4D" author="Butler, A. G." box="[1267,1408,1161,1187]" pageId="4" pageNumber="391" pagination="270 - 273" refId="ref5399" refString="Butler, A. G. (1869) Descriptions of species of Lepidoptera, confounded with others described by Linnaeus and Fabricius. The Entomologist's monthly Magazine, 5, 270 - 273." type="journal article" year="1869">Butler, 1869</bibRefCitation>
</taxonomicName>
is “
<collectingRegion id="49B7F844FF808C3FBFD3D3AAFF6DFB29" box="[161,253,1197,1223]" country="Turkey" name="Izmir" pageId="4" pageNumber="391">Smyrna</collectingRegion>
” (
<bibRefCitation id="EFE24B57FF808C3FBE67D3AAFE57FB29" author="Hemming, F. A." box="[277,455,1197,1223]" pageId="4" pageNumber="391" pagination="493 - 504" refId="ref6087" refString="Hemming, F. A. (1931) New material regarding the dates of the plates of the Papiliones in Jacob Hubner's &quot; Sammlung europaischer Schmetterlinge &quot;, with notes on the synonymy and type localities of certain species described therein. Transactions of the entomological Society of London, 79 (3), 493 - 504. https: // doi. org / 10.1111 / j. 1365 - 2311.1931. tb 00713. x" type="journal article" year="1931">Hemming 1931</bibRefCitation>
,
<bibRefCitation id="EFE24B57FF808C3FBEA1D3AAFDE7FB29" author="Bernardi, G." box="[467,631,1197,1223]" pageId="4" pageNumber="391" pagination="1 - 24" refId="ref5209" refString="Bernardi, G. (1947) La nomenclature de quelques formes europeennes de l' Euchloe ausonia Hb. (= belia Auct.). Miscellanea entomologica; Revue entomologique internationale, 44 (1), 1 - 24." type="journal article" year="1947">Bernardi 1947</bibRefCitation>
), viz. a site in an area of West
<collectingCountry id="F3647636FF808C3FBCB4D3AAFB86FB29" box="[966,1046,1197,1223]" name="Turkey" pageId="4" pageNumber="391">Turkey</collectingCountry>
where only the species now generally identified as
<taxonomicName id="4C734D25FF808C3FBE2BD3D6FD85FB05" authorityName="Hubner" authorityYear="1804" box="[345,533,1233,1259]" class="Insecta" family="Pieridae" genus="Euchloe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="ausonia">
<emphasis id="B907EAB4FF808C3FBE2BD3D6FD85FB05" box="[345,533,1233,1259]" italics="true" pageId="4" pageNumber="391">Euchloe ausonia</emphasis>
</taxonomicName>
is known to occur (
<bibRefCitation id="EFE24B57FF808C3FBD83D3D6FC6AFB05" author="Hesselbarth, G. &amp; van Oorschot, H. &amp; Wagener, S." box="[753,1018,1232,1259]" pageId="4" pageNumber="391" pagination="1 - 847" refId="ref6289" refString="Hesselbarth, G., van Oorschot, H. &amp; Wagener, S. (1995) Die Tagfalter der Turkei. 3 Vols. Wagener, Bocholt, 2001 pp. [pp. 1 - 754 + 755 - 1154 + 1 - 847]" type="book chapter" year="1995">
Hesselbarth
<emphasis id="B907EAB4FF808C3FBC0DD3D5FC27FB05" box="[895,951,1233,1259]" italics="true" pageId="4" pageNumber="391">et al.</emphasis>
1995
</bibRefCitation>
).
</paragraph>
<paragraph id="8BCC36A6FF808C3FBFB5D3F2FE4FFAB9" blockId="4.[151,1437,728,1691]" pageId="4" pageNumber="391">
a) Accordingly,
<bibRefCitation id="EFE24B57FF808C3FBE0ED3F2FDB6FAE1" author="Higgins, L. G." box="[380,550,1268,1295]" pageId="4" pageNumber="391" refId="ref6364" refString="Higgins, L. G. (1975) The classification of European butterflies. Collins, London, 320 pp., 60 pls." type="book" year="1975">Higgins (1975)</bibRefCitation>
considered the name
<taxonomicName id="4C734D25FF808C3FBC6FD3F2FBB7FAE1" authority="Butler" authorityName="Butler" authorityYear="1869" box="[797,1063,1269,1295]" class="Insecta" family="Pieridae" genus="Euchloe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="crameri">
<emphasis id="B907EAB4FF808C3FBC6FD3F2FC49FAE1" box="[797,985,1269,1295]" italics="true" pageId="4" pageNumber="391">Euchloe crameri</emphasis>
Butler
</taxonomicName>
the correct name for the eastern dappled white, while this name was deemed a synonym or a subspecies of
<taxonomicName id="4C734D25FF808C3FBCA9D21EFB06FADD" authorityName="Hubner" authorityYear="1804" box="[987,1174,1305,1331]" class="Insecta" family="Pieridae" genus="Euchloe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="ausonia">
<emphasis id="B907EAB4FF808C3FBCA9D21EFB06FADD" box="[987,1174,1305,1331]" italics="true" pageId="4" pageNumber="391">Euchloe ausonia</emphasis>
</taxonomicName>
by a number of authors (
<bibRefCitation id="EFE24B57FF808C3FBFEDD23AFE0CFAB9" author="Higgins, L. G. &amp; Riley, N. D." box="[159,412,1341,1367]" pageId="4" pageNumber="391" refId="ref6420" refString="Higgins, L. G. &amp; Riley, N. D. (1970) A field guide to the Butterflies of Britain and Europe. 1 st Edition. Collins, London, 380 pp., 60 pls." type="book" year="1970">Higgins &amp; Riley 1970</bibRefCitation>
, etc.).
</paragraph>
<paragraph id="8BCC36A6FF808C3FBFB5D266FE5BF9BD" blockId="4.[151,1437,728,1691]" pageId="4" pageNumber="391">
b)
<bibRefCitation id="EFE24B57FF808C3FBF95D266FE12FA92" author="Butler, A. G." box="[231,386,1377,1404]" pageId="4" pageNumber="391" pagination="270 - 273" refId="ref5399" refString="Butler, A. G. (1869) Descriptions of species of Lepidoptera, confounded with others described by Linnaeus and Fabricius. The Entomologist's monthly Magazine, 5, 270 - 273." type="journal article" year="1869">Butler (1869)</bibRefCitation>
, however, had decided to select “S. Europe (obtained by Herr J.J. Becker)”, as type locality of his new taxon.
<bibRefCitation id="EFE24B57FF808C3FBE14D282FDFEFA71" author="Rothschild, W." box="[358,622,1413,1439]" pageId="4" pageNumber="391" pagination="301 - 305" refId="ref7431" refString="Rothschild, W. (1914) A preliminary account of the Lepidopteran Fauna of Guelt-es-Stel, Central Algeria, Novitates Zoologicae, 21, 301 - 305. https: // doi. org / 10.5962 / bhl. part. 16331" type="journal article" year="1914">Rothschild (1914: 302)</bibRefCitation>
managed to find in the British Museum the specimen that he thought to be Butlers
<typeStatus id="54C88804FF808C3FBE4BD2AEFE0BFA2D" box="[313,411,1449,1475]" pageId="4" pageNumber="391" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C734D25FF808C3FBEB1D2AEFDEFFA2D" authorityName="Butler" authorityYear="1869" box="[451,639,1449,1475]" class="Insecta" family="Pieridae" genus="Euchloe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="crameri">
<emphasis id="B907EAB4FF808C3FBEB1D2AEFDEFFA2D" box="[451,639,1449,1475]" italics="true" pageId="4" pageNumber="391">Euchloe crameri</emphasis>
</taxonomicName>
, interpreted as such a specimen collected in
<collectingCountry id="F3647636FF808C3FBB09D2AEFB2BFA2D" box="[1147,1211,1449,1475]" name="Spain" pageId="4" pageNumber="391">Spain</collectingCountry>
and used this name to identify the western dappled white.
<bibRefCitation id="EFE24B57FF808C3FBD12D2CAFC8AFA09" author="Oberthur, C." box="[608,794,1485,1511]" pageId="4" pageNumber="391" pagination="51 - 61" refId="ref7207" refString="Oberthur, C. (1914) Anthocharis crameri Butler. Etudes de Lepidopterologie comparee, 10, 51 - 61. [texted]" type="journal article" year="1914">Oberthür (1914)</bibRefCitation>
, perhaps independently, reached a similar conclusion, to which also concurred
<bibRefCitation id="EFE24B57FF808C3FBEE2D2F6FDADF9E5" author="Bernardi, G." box="[400,573,1520,1547]" pageId="4" pageNumber="391" pagination="66 - 103" refId="ref5169" refString="Bernardi, G. (1945) Etudes sur le genre Euchloe Hb. (Lep. Pieridae). Miscellanea entomologica; Revue entomologique internationale, 42 (6), 66 - 103, pls. IV-VI." type="journal article" year="1945">Bernardi (1945</bibRefCitation>
,
<bibRefCitation id="EFE24B57FF808C3FBD38D2F6FD16F9E5" author="Bernardi, G." box="[586,646,1521,1547]" pageId="4" pageNumber="391" pagination="1 - 24" refId="ref5209" refString="Bernardi, G. (1947) La nomenclature de quelques formes europeennes de l' Euchloe ausonia Hb. (= belia Auct.). Miscellanea entomologica; Revue entomologique internationale, 44 (1), 1 - 24." type="journal article" year="1947">1947</bibRefCitation>
), etc. Actually, Butlers name was based on three
<typeStatus id="54C88804FF808C3FBBC8D2F6FA8FF9E5" box="[1210,1311,1521,1547]" pageId="4" pageNumber="391" type="syntype">syntypes</typeStatus>
, i.e. Stolls lost specimen from “Smyrna”, and two specimens, declaredly collected in “
<collectingCountry id="F3647636FF808C3FBC9AD112FBBDF9C1" box="[1000,1069,1557,1583]" name="Spain" pageId="4" pageNumber="391">Spain</collectingCountry>
” (D. Lees,
<emphasis id="B907EAB4FF808C3FBBDFD111FB61F9C1" box="[1197,1265,1557,1583]" italics="true" pageId="4" pageNumber="391">in litt.</emphasis>
) (see
<figureCitation id="13482A23FF808C3FBA44D112FA1DF9C1" box="[1334,1421,1557,1583]" captionStart="FIGURE 1" captionStartId="2.[151,250,1718,1743]" captionTargetBox="[160,1421,191,1687]" captionTargetId="figure@2.[151,1436,179,1691]" captionTargetPageId="2" captionText="FIGURE 1. (A). Lectotype (♂) of Pieris daplidice race nitida Verity, 1908: (d) dorsal (v) ventral (MZUF). (B). Neotype (♀) of [Papilio] ausonia Hübner, [1804]: (d) dorsal (v) ventral) (MZUF). (C). Centre: Original Freyers sealed box containing the Neotype (♂) of Pontia simplonia Freyer, 1829 (bottom right), together with a female of the same species (top left); Left: electronically extracted enlargement of the Neotype (♂) (d) dorsal (JME). (D). Lectotype (♂) of Euchloe crameri Butler, 1869 as designated by Rothschild (1914) (NHMUK). (E). Lectotype (♂) of Euchloe ausonia var. esperi Kirby, 1871 (5) (NMI). Labels relative to the illustrated specimens are shown on the right of each row." figureDoi="http://doi.org/10.5281/zenodo.3842458" httpUri="https://zenodo.org/record/3842458/files/figure.png" pageId="4" pageNumber="391">Fig. 1D</figureCitation>
), still extant in the NHMUK.
</paragraph>
<paragraph id="8BCC36A6FF808C3FBFB5D15AFAE1F975" blockId="4.[151,1437,728,1691]" pageId="4" pageNumber="391">
<materialsCitation id="3B1B3CFBFF808C3FBFB5D15AFAFCF975" ID-GBIF-Occurrence="2625370302" collectionCode="NHMUK" country="Spain" location="Spain" pageId="4" pageNumber="391" specimenCount="1" typeStatus="lectotype">
Rothschild, although wrongly supposing that Butlers name was based on a single specimen, can be deemed as having designated one of the
<collectionCode id="ED62AE63FF808C3FBE92D186FDDAF975" box="[480,586,1665,1691]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="391" type="Museum">NHMUK</collectionCode>
specimens from “
<collectingCountry id="F3647636FF808C3FBC66D186FCC9F975" box="[788,857,1665,1691]" name="Spain" pageId="4" pageNumber="391">Spain</collectingCountry>
” as the
<typeStatus id="54C88804FF808C3FBCC3D186FB8BF975" box="[945,1051,1665,1691]" pageId="4" pageNumber="391" type="lectotype">lectotype</typeStatus>
, under Art. 74.6 (
<bibRefCitation id="EFE24B57FF808C3FBBACD186FAF4F975" author="ICZN" box="[1246,1380,1665,1691]" pageId="4" pageNumber="391" refId="ref6898" refString="ICZN (1999) (and later updates) International Code of Zoological Nomenclature. Available from: http: // www. nhm. ac. uk / hosted-sites / iczn / code / (accessed 5 March 2020)" type="url" year="1999">ICZN 1999</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>