392 lines
48 KiB
XML
392 lines
48 KiB
XML
<document id="EF1600B67F8288CDB68D0221673E4104" ID-DOI="10.5852/ejt.2019.558" ID-GBIF-Dataset="5eadba67-d4ee-4eb8-8ec9-bdb75cab1d59" ID-ISSN="2118-9773" ID-Zenodo-Dep="3474974" ID-ZooBank="5B9DC474-1F26-4732-B7B8-32C285FE0099" IM.bibliography_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1570442388906" checkinUser="plazi" docAuthor="Fehér, Zoltán, Deli, Tamás, Erőss, Zoltán P. & Lika, Romilda" docDate="2019" docId="03B17A5AF706FC4BFE0BF6E0689EFE5F" docLanguage="en" docName="ejt-558_feher_deli_eross.pdf.imf" docOrigin="European Journal of Taxonomy 558" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Virpazaria (Aemiliella) ripkeni subsp. arbeni Gittenberger 2015" docType="treatment" docVersion="16" lastPageNumber="22" masterDocId="FF880222F712FC5DFFCFF2516A65FFED" masterDocTitle="Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="21" updateTime="1716577348435" updateUser="carolina">
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<mods:titleInfo id="C0BC1FB9235A7DDA6E3B10647DFCECEA">
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<mods:title id="ED1091D983A72120F3FDB70EB0A1DB8B">Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae)</mods:title>
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<mods:name id="984A0217388DAD273AB1782E44C5A5A1" type="personal">
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<mods:roleTerm id="6749FF41B9EAFE20F9C62CDFAA17CB4D">Author</mods:roleTerm>
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<mods:namePart id="2A1264C15E0EE9CB3731F5FE56503B37">Fehér, Zoltán</mods:namePart>
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<mods:nameIdentifier id="D5270D105F2B406D3E7446E90F54FFAD" type="LSID">E801EC76-8B1E-450B-993E-BBBE57C00EA9</mods:nameIdentifier>
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<mods:affiliation id="91B57A72ACF51053C4600B7DC10301B7">Department of Zoology, Hungarian Natural History Museum, Baross u. 13, Budapest, H- 1088, Hungary.</mods:affiliation>
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<mods:nameIdentifier id="59B99068B2C61D704C662193DB9E042A" type="email">feher.zoltan@nhmus.hu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="4C5B6A314A9C1C2A29DA02392985816A" type="personal">
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<mods:roleTerm id="10386AC0017BE7C74475466EBCD4CCDA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9036E39045E5987DE2D8A7B04709B4F7">Deli, Tamás</mods:namePart>
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<mods:nameIdentifier id="D6F16058F53105B4E7810E4853B46B7A" type="LSID">A33DCDF3-6293-4B34-B1DD-330211905D74</mods:nameIdentifier>
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<mods:affiliation id="B7E5D4989D4EEEC1C0CD29BE15723374">Móricz Zsigmond u. 2, Gyomaendrőd, H- 5500, Hungary.</mods:affiliation>
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<mods:nameIdentifier id="57D8F1CD15F4B5BDBD6AC602D7890843" type="email">virpazaria@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="D78CB01BBD011CD2A9703E296A7B49D5" type="personal">
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<mods:roleTerm id="D83B0BF30AE5A9641C0F2FD10B4FA532">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2274B921C03822CC54C696A0ADE5AD39">Erőss, Zoltán P.</mods:namePart>
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<mods:nameIdentifier id="7C938C8CE612A2C6B42781F8EDE0449E" type="LSID">0B2C74A7-0001-4E25-984C-88CB0C705313</mods:nameIdentifier>
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<mods:affiliation id="85A15F4D864A30957C1DFA75C9E07C00">Bem u. 36., Budapest, H- 1151, Hungary.</mods:affiliation>
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<mods:nameIdentifier id="77F2D2C3172AE5FFC05C4305C306EA8C" type="email">erosspeter@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="5D8EDDE9EE7F1A767D46CDCD09493731" type="personal">
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<mods:roleTerm id="6D4A146A130DBCD70F3ECE9FB0DA0F97">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2DCF2F2B4305ECF0D5A4558C0C4DD566">Lika, Romilda</mods:namePart>
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<mods:nameIdentifier id="F476C0D4369DE2D4F9A43630E89844A4" type="LSID">9662552B-B203-43FF-B427-4330080E1CBA</mods:nameIdentifier>
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<mods:affiliation id="02D70A9BAD06CD8C1745AC36D8AFA174">Departament of Biology, Faculty of Natural Sciences, Tirana University, Bulevardi Zogu i Parë, Tiranë, AL- 1001, Albania.</mods:affiliation>
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<mods:nameIdentifier id="78B1D2A068CBE5848B4602B6D67F82ED" type="email">romildalika@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="A809D423C7AEC29EB0764BA2A4C37570">text</mods:typeOfResource>
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<mods:titleInfo id="DABF4C13485850870C2A7B0F67BCA183">
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<mods:title id="855CC75CBD3F963C8195C04BC172AABA">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="3D39111B2AA8C4E847870352D187DAB3">
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<mods:date id="9FF323B09013184D548FF5F021FD0ED7">2019</mods:date>
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<mods:number id="B6E4C0A56DAF14D8B763DF721BD6BB25">2019-09-26</mods:number>
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</mods:detail>
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<mods:detail id="3A413AA056B7E99A363D6537CDD98925" type="volume">
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<mods:number id="B1AA6388222639E6D79DB65B3AEECCF9">558</mods:number>
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</mods:detail>
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<mods:start id="76A8C47C7FA6EC9A3607524C45A27416">1</mods:start>
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<mods:end id="6D360BF5B9DB1DE1CC3AFE3D8EA91D3B">25</mods:end>
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</mods:extent>
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<mods:classification id="B8C6301170FA8B94B81F711A1A2F89FF">journal article</mods:classification>
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<mods:identifier id="3266E32DD069A3DB8BF00E32AC357F97" type="DOI">10.5852/ejt.2019.558</mods:identifier>
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<mods:identifier id="906F014106B15A61484479851C01DE8B" type="GBIF-Dataset">5eadba67-d4ee-4eb8-8ec9-bdb75cab1d59</mods:identifier>
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<mods:identifier id="6D4B81BEE84C46B6A9E4A2635B3CA178" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="A514D4CF4BD05EE61997A9A0CDD512BF" type="Zenodo-Dep">3474974</mods:identifier>
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<mods:identifier id="0626C5CB30B2BFE592B539389DA0D414" type="ZooBank">5B9DC474-1F26-4732-B7B8-32C285FE0099</mods:identifier>
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</mods:mods>
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<treatment id="03B17A5AF706FC4BFE0BF6E0689EFE5F" ID-DOI="http://doi.org/10.5281/zenodo.3475242" ID-GBIF-Taxon="159654651" ID-Zenodo-Dep="3475242" LSID="urn:lsid:plazi:treatment:03B17A5AF706FC4BFE0BF6E0689EFE5F" httpUri="http://treatment.plazi.org/id/03B17A5AF706FC4BFE0BF6E0689EFE5F" lastPageId="22" lastPageNumber="22" pageId="20" pageNumber="21">
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||
<subSubSection id="C30298C7F706FC49FE0BF6E06E15FB21" box="[452,1136,1201,1228]" pageId="20" pageNumber="21" type="nomenclature">
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||
<paragraph id="8BA7CB4CF706FC49FE0BF6E06E15FB21" blockId="20.[452,1136,1201,1228]" box="[452,1136,1201,1228]" pageId="20" pageNumber="21">
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<heading id="D0EF7C20F706FC49FE0BF6E06E15FB21" box="[452,1136,1201,1228]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
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<taxonomicName id="4C18B0CFF706FC49FE0BF6E06E15FB21" ID-CoL="7J5NK" authority="Gittenberger, 2015" authorityName="Gittenberger" authorityYear="2015" box="[452,1136,1201,1228]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="subSpecies" species="ripkeni" subGenus="Aemiliella" subSpecies="arbeni">
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<emphasis id="B96C175EF706FC49FE0BF6E069E9FB21" bold="true" box="[452,908,1201,1228]" italics="true" pageId="20" pageNumber="21">Virpazaria (Aemiliella) ripkeni arbeni</emphasis>
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<bibRefCitation id="EF89B6BDF706FC49FC5BF6E06E15FB21" author="Gittenberger E." box="[916,1136,1201,1228]" firstAuthor="Gittenberger" journalOrPublisher="Basteria" pageId="20" pageNumber="21" pagination="23 - 29" part="79" refId="ref12813" refString="Gittenberger E. 2015. Caving for snails in northern Albania. Basteria 79 (1 - 3): 23 - 29." title="Caving for snails in northern Albania" type="journal article" year="2015">Gittenberger, 2015</bibRefCitation>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30298C7F706FC49FF72F6A269BBFAE0" box="[189,990,1266,1293]" pageId="20" pageNumber="21" type="reference_group">
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||
<paragraph id="8BA7CB4CF706FC49FF72F6A269BBFAE0" blockId="20.[189,990,1266,1293]" box="[189,990,1266,1293]" pageId="20" pageNumber="21">
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||
<treatmentCitationGroup id="AB08EC62F706FC49FF72F6A269BBFAE0" box="[189,990,1266,1293]" pageId="20" pageNumber="21">
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||
<treatmentCitation id="0AB9ED5DF706FC49FF72F6A269EBFAE0" author="Gittenberger E." box="[189,910,1266,1293]" page="27" pageId="20" pageNumber="21" year="2015">
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<taxonomicName id="4C18B0CFF706FC49FF72F6A269EBFAE0" ID-CoL="7J5NK" authority="Gittenberger, 2015: 27" authorityName="Gittenberger" authorityPageNumber="27" authorityYear="2015" box="[189,910,1266,1293]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="subSpecies" species="ripkeni" subGenus="Aemiliella" subSpecies="arbeni">
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<emphasis id="B96C175EF706FC49FF72F6A26819FAE0" box="[189,636,1267,1293]" italics="true" pageId="20" pageNumber="21">Virpazaria (Aemiliella) ripkeni arbeni</emphasis>
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||
<bibRefCitation id="EF89B6BDF706FC49FD4CF6A269EBFAE0" author="Gittenberger E." box="[643,910,1266,1293]" firstAuthor="Gittenberger" journalOrPublisher="Basteria" pageId="20" pageNumber="21" pagination="23 - 29" part="79" refId="ref12813" refString="Gittenberger E. 2015. Caving for snails in northern Albania. Basteria 79 (1 - 3): 23 - 29." title="Caving for snails in northern Albania" type="journal article" year="2015">Gittenberger, 2015: 27</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, fig. 8.
|
||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30298C7F706FC49FF72F7656F5DFA4D" pageId="20" pageNumber="21" type="diagnosis">
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||
<paragraph id="8BA7CB4CF706FC49FF72F7656B53FAA3" blockId="20.[189,310,1332,1358]" box="[189,310,1332,1358]" pageId="20" pageNumber="21">
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||
<heading id="D0EF7C20F706FC49FF72F7656B53FAA3" bold="true" box="[189,310,1332,1358]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
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||
<emphasis id="B96C175EF706FC49FF72F7656B53FAA3" bold="true" box="[189,310,1332,1358]" pageId="20" pageNumber="21">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="8BA7CB4CF706FC49FF72F7336F5DFA4D" blockId="20.[189,1398,1378,1440]" pageId="20" pageNumber="21">The upper palatal lamella which does not or hardly reaches the peristome and the non-thickened, knobless peristome are the features that distinguish this subspecies from the nominate subspecies.</paragraph>
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</subSubSection>
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||
<subSubSection id="C30298C7F706FC49FF72F7966E8DF8A2" pageId="20" pageNumber="21" type="materials_examined">
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||
<paragraph id="8BA7CB4CF706FC49FF72F7966BCFFA0C" blockId="20.[189,426,1479,1505]" box="[189,426,1479,1505]" pageId="20" pageNumber="21">
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||
<heading id="D0EF7C20F706FC49FF72F7966BCFFA0C" bold="true" box="[189,426,1479,1505]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
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||
<emphasis id="B96C175EF706FC49FF72F7966BCFFA0C" bold="true" box="[189,426,1479,1505]" pageId="20" pageNumber="21">Material examined</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BA7CB4CF706FC49FF25F7A76B39F9FD" blockId="20.[189,1399,1526,1623]" box="[234,348,1526,1552]" pageId="20" pageNumber="21">
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||
<emphasis id="B96C175EF706FC49FF25F7A76B39F9FD" bold="true" box="[234,348,1526,1552]" pageId="20" pageNumber="21">
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||
<typeStatus id="54A375EEF706FC49FF25F7A76B39F9FD" box="[234,348,1526,1552]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
</paragraph>
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||
<paragraph id="8BA7CB4CF706FC49FF72F4486898F9BA" blockId="20.[189,1399,1526,1623]" pageId="20" pageNumber="21">
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||
<materialsCitation id="3B70C111F706FC49FF72F4486898F9BA" ID-GBIF-Occurrence="2424146976" collectingDate="2011-10-18" collectionCode="RMNH" collectorName="E. Gittenberger" country="Albania" latitude="42.0487" location="Rozafa Cave" longLatPrecision="6" longitude="19.4938" pageId="20" pageNumber="21" specimenCode="5003974" specimenCount="1" stateProvince="Shkoder" typeStatus="holotype">
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<collectingCountry id="F30F8BDCF706FC49FF72F4486B5AF9DE" box="[189,319,1561,1587]" name="Albania" pageId="20" pageNumber="21">ALBANIA</collectingCountry>
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||
•
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<collectingRegion id="49DC05AEF706FC49FEAFF4486BA1F9DE" box="[352,452,1561,1587]" country="Albania" name="Shkoder" pageId="20" pageNumber="21">Shkodër</collectingRegion>
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||
,
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||
<location id="8EC79D97F706FC49FE1CF448680BF9DE" LSID="urn:lsid:plazi:treatment:03B17A5AF706FC4BFE0BF6E0689EFE5F:8EC79D97F706FC49FE1CF448680BF9DE" box="[467,622,1561,1587]" country="Albania" latitude="42.0487" longLatPrecision="6" longitude="19.4938" name="Rozafa Cave" pageId="20" pageNumber="21" stateProvince="Shkoder">Rozafa Cave</location>
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||
[= Shpella e Rozafës];
|
||
<geoCoordinate id="EE2CAD8BF706FC49FC59F4486E41F9DE" box="[918,1060,1561,1587]" degrees="42.0487" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="5" value="42.0487">42.0487° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2CAD8BF706FC49FBFBF4486EDAF9DE" box="[1076,1215,1561,1587]" degrees="19.4938" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="5" value="19.4938">19.4938° E</geoCoordinate>
|
||
;
|
||
<date id="FFA6ED8CF706FC49FB00F4486F17F9DE" box="[1231,1394,1561,1587]" pageId="20" pageNumber="21" value="2011-10-18">
|
||
<collectingDate id="EFE21464F706FC49FB00F4486F17F9DE" box="[1231,1394,1561,1587]" pageId="20" pageNumber="21" value="2011-10-18">18 Oct. 2011</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26EDAE9AF706FC49FF72F46C6B17F9BB" box="[189,370,1596,1623]" pageId="20" pageNumber="21">E. Gittenberger</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED095389F706FC49FE7CF46C686EF9BA" box="[435,523,1597,1623]" pageId="20" pageNumber="21">RMNH</collectionCode>
|
||
<specimenCode id="DBBE6337F706FC49FDDCF46D681FF9BA" box="[531,634,1596,1623]" pageId="20" pageNumber="21">5003974</specimenCode>
|
||
[not seen].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA7CB4CF706FC49FF25F42F6B02F975" blockId="20.[189,1398,1662,1758]" box="[234,359,1662,1688]" pageId="20" pageNumber="21">
|
||
<emphasis id="B96C175EF706FC49FF25F42F6B02F975" bold="true" box="[234,359,1662,1688]" pageId="20" pageNumber="21">
|
||
<typeStatus id="54A375EEF706FC49FF25F42F6B02F975" box="[234,359,1662,1688]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA7CB4CF706FC49FF72F4F06B67F933" blockId="20.[189,1398,1662,1758]" pageId="20" pageNumber="21">
|
||
<materialsCitation id="3B70C111F706FC49FF72F4F06A9BF933" ID-GBIF-Occurrence="2424146988" collectingDate="2013-05" collectionCode="RMNH" collectorName="M. Slooff" country="Albania" latitude="42.0487" location="ALBANIA" longLatPrecision="6" longitude="19.4938" pageId="20" pageNumber="21" specimenCode="5003975" specimenCount="2" stateProvince="Shkoder" typeStatus="paratype">
|
||
<collectingCountry id="F30F8BDCF706FC49FF72F4F06B5AF956" box="[189,319,1697,1723]" name="Albania" pageId="20" pageNumber="21">ALBANIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D1E00C5F706FC49FE92F4F06BD1F956" box="[349,436,1697,1723]" count="2" pageId="20" pageNumber="21" type="generic">2 spec.</specimenCount>
|
||
; same locality as for holotype;
|
||
<date id="FFA6ED8CF706FC49FCFFF4F069C8F956" box="[816,941,1697,1723]" pageId="20" pageNumber="21" value="2013-05">
|
||
<collectingDate id="EFE21464F706FC49FCFFF4F069C8F956" box="[816,941,1697,1723]" pageId="20" pageNumber="21" value="2013-05">May 2013</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26EDAE9AF706FC49FC74F4F06E4AF956" box="[955,1071,1697,1723]" pageId="20" pageNumber="21">M. Slooff</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED095389F706FC49FBBAF4F06EA8F956" box="[1141,1229,1697,1723]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="20" pageNumber="21">RMNH</collectionCode>
|
||
<specimenCode id="DBBE6337F706FC49FB18F4F06F5BF951" box="[1239,1342,1697,1724]" pageId="20" pageNumber="21">5003975</specimenCode>
|
||
[not seen]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7CB4CF706FC49FF72F5546B2BF8F2" blockId="20.[189,334,1797,1823]" box="[189,334,1797,1823]" pageId="20" pageNumber="21">
|
||
<heading id="D0EF7C20F706FC49FF72F5546B2BF8F2" bold="true" box="[189,334,1797,1823]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
|
||
<emphasis id="B96C175EF706FC49FF72F5546B2BF8F2" bold="true" box="[189,334,1797,1823]" pageId="20" pageNumber="21">Dimensions</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA7CB4CF706FC49FF72F5646E8DF8A2" blockId="20.[189,1256,1844,1871]" box="[189,1256,1844,1871]" pageId="20" pageNumber="21">
|
||
H
|
||
<quantity id="4CE066A9F706FC49FF16F5656B0FF8A2" box="[217,362,1844,1871]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="20" pageNumber="21" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
|
||
; W
|
||
<quantity id="4CE066A9F706FC49FE57F5656840F8A2" box="[408,549,1844,1871]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="20" pageNumber="21" unit="mm" value="3.15" valueMax="3.2" valueMin="3.1">3.1–3.2 mm</quantity>
|
||
(based on three specimens according to
|
||
<bibRefCitation id="EF89B6BDF706FC49FBCEF5656EB9F8A2" author="Gittenberger E." box="[1025,1244,1844,1871]" firstAuthor="Gittenberger" journalOrPublisher="Basteria" pageId="20" pageNumber="21" pagination="23 - 29" part="79" refId="ref12813" refString="Gittenberger E. 2015. Caving for snails in northern Albania. Basteria 79 (1 - 3): 23 - 29." title="Caving for snails in northern Albania" type="journal article" year="2015">Gittenberger 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30298C7F706FC48FF72F524682CFE7D" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="distribution">
|
||
<paragraph id="8BA7CB4CF706FC49FF72F5246B33F862" blockId="20.[189,342,1909,1935]" box="[189,342,1909,1935]" pageId="20" pageNumber="21">
|
||
<heading id="D0EF7C20F706FC49FF72F5246B33F862" bold="true" box="[189,342,1909,1935]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
|
||
<emphasis id="B96C175EF706FC49FF72F5246B33F862" bold="true" box="[189,342,1909,1935]" pageId="20" pageNumber="21">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA7CB4CF706FC48FF72F5F5682CFE7D" blockId="20.[189,1399,1956,2018]" lastBlockId="21.[189,1399,267,400]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
|
||
Recent records of this species are from the southern rim of the
|
||
<collectingRegion id="49DC05AEF706FC49FC4EF5F56986F853" box="[897,995,1956,1982]" country="Albania" name="Shkoder" pageId="20" pageNumber="21">Shkodër</collectingRegion>
|
||
Lake Basin, from the surroundings of
|
||
<collectingRegion id="49DC05AEF706FC49FF10F5966B26F80C" box="[223,323,1991,2017]" country="Albania" name="Shkoder" pageId="20" pageNumber="21">Shkodër</collectingRegion>
|
||
. The
|
||
<collectingRegion id="49DC05AEF706FC49FE49F5966B8DF80C" box="[390,488,1991,2017]" country="Albania" name="Shkoder" pageId="20" pageNumber="21">Shkodër</collectingRegion>
|
||
site is located on Upper Triassic, the Drisht site on Middle Triassic and the Renc Mts site on Upper Cretaceous limestone. If these were the only known localities, this would already make
|
||
<taxonomicName id="4C18B0CFF707FC48FEA9F37E6BBDFEA4" authorityName="Gittenberger" authorityYear="1969" box="[358,472,303,329]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FEA9F37E6BBDFEA4" box="[358,472,303,329]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
the widest distributed species of its genus. Strangely, the
|
||
<typeStatus id="54A375EEF707FC48FB5CF37E6EA3FEA4" box="[1171,1222,303,329]" pageId="21" pageNumber="22">type</typeStatus>
|
||
locality of the nominate subspecies is located at the northwestern side of the
|
||
<collectingRegion id="49DC05AEF707FC48FC56F303699EFE81" box="[921,1019,338,364]" country="Albania" name="Shkoder" pageId="21" pageNumber="22">Shkodër</collectingRegion>
|
||
Lake Basin, very far from these Albanian sites (but see Remarks).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30298C7F707FC4BFF72F3EF689EFE5F" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="discussion">
|
||
<paragraph id="8BA7CB4CF707FC48FF72F3EF6B4AFE35" blockId="21.[189,303,446,472]" box="[189,303,446,472]" pageId="21" pageNumber="22">
|
||
<heading id="D0EF7C20F707FC48FF72F3EF6B4AFE35" bold="true" box="[189,303,446,472]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
|
||
<emphasis id="B96C175EF707FC48FF72F3EF6B4AFE35" bold="true" box="[189,303,446,472]" pageId="21" pageNumber="22">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA7CB4CF707FC48FF72F3BD692FFC40" blockId="21.[189,1399,492,941]" pageId="21" pageNumber="22">
|
||
This species was described on the basis of eight specimens. Their width and height are 3.5–4.0 mm and
|
||
<quantity id="4CE066A9F707FC48FF3DF05E6BC1FDC7" box="[242,420,527,554]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.25" metricValueMin="2.05" pageId="21" pageNumber="22" unit="mm" value="2.15" valueMax="2.25" valueMin="2.05">2.05–2.25 mm</quantity>
|
||
, respectively, and the number of ribs on the last whorl is 73–86. In the original description (
|
||
<bibRefCitation id="EF89B6BDF707FC48FE9CF0626855FDA3" author="Gittenberger E." box="[339,560,563,590]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="21" pageNumber="22" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger 1969</bibRefCitation>
|
||
) somewhat lower values were given, probably because he applied a different measuring method. The
|
||
<typeStatus id="54A375EEF707FC48FD89F00768C8FD9D" box="[582,685,598,624]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C18B0CFF707FC48FD1AF006691EFD9D" box="[725,891,599,624]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pastorpueri">
|
||
<emphasis id="B96C175EF707FC48FD1AF006691EFD9D" box="[725,891,599,624]" italics="true" pageId="21" pageNumber="22">V. pastorpueri</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<quantity id="4CE066A9F707FC48FC52F007699DFD9C" box="[925,1016,598,625]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="21" pageNumber="22" unit="mm" value="3.6">3.6 mm</quantity>
|
||
wide and
|
||
<quantity id="4CE066A9F707FC48FBBEF0076EBFFD9C" box="[1137,1242,598,625]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="21" pageNumber="22" unit="mm" value="2.15">2.15 mm</quantity>
|
||
high with 60 ribs on its last whorl (although
|
||
<bibRefCitation id="EF89B6BDF707FC48FDE1F028695CFD7E" author="Reischutz A. & Reischutz N. & Reischutz P. L." box="[558,825,633,660]" etAl="et al." firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="21" pageNumber="22" pagination="57 - 60" part="18" refId="ref13283" refString="Reischutz A., Reischutz N. & Reischutz P. L. 2011. Anmerkungen zur Verbreitung von Virpazaria Gittenberger 1969 (Spelaeodiscidae: Gastropoda) und bemerkenswerte Mollusken aus Albanien. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 18: 57 - 60." title="Anmerkungen zur Verbreitung von Virpazaria Gittenberger 1969 (Spelaeodiscidae: Gastropoda) und bemerkenswerte Mollusken aus Albanien" type="journal article" year="2011">
|
||
Reischütz
|
||
<emphasis id="B96C175EF707FC48FD66F02B68B8FD7E" box="[681,733,633,659]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
|
||
. (2011)
|
||
</bibRefCitation>
|
||
provided a smaller width (3.0–
|
||
<quantity id="4CE066A9F707FC48FB64F0286F6CFD79" box="[1195,1289,633,660]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="mm" value="3.5">3.5 mm</quantity>
|
||
), a larger height (
|
||
<quantity id="4CE066A9F707FC48FED6F0CD6B1CFD5A" box="[281,377,668,695]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="22" unit="mm" value="2.5">2.5 mm</quantity>
|
||
) and an average of 65 ribs on the last whorl). Smaller and more tapered shells were defined by
|
||
<bibRefCitation id="EF89B6BDF707FC48FE89F091683CFD37" author="Reischutz A. & Reischutz N. & Reischutz P. L." box="[326,601,704,730]" etAl="et al." firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="21" pageNumber="22" pagination="57 - 60" part="18" refId="ref13283" refString="Reischutz A., Reischutz N. & Reischutz P. L. 2011. Anmerkungen zur Verbreitung von Virpazaria Gittenberger 1969 (Spelaeodiscidae: Gastropoda) und bemerkenswerte Mollusken aus Albanien. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 18: 57 - 60." title="Anmerkungen zur Verbreitung von Virpazaria Gittenberger 1969 (Spelaeodiscidae: Gastropoda) und bemerkenswerte Mollusken aus Albanien" type="journal article" year="2011">
|
||
Reischütz
|
||
<emphasis id="B96C175EF707FC48FE0BF0906867FD37" box="[452,514,704,730]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
as the most important features to distinguish
|
||
<taxonomicName id="4C18B0CFF707FC48FB45F0906F56FD37" box="[1162,1331,705,730]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pastorpueri">
|
||
<emphasis id="B96C175EF707FC48FB45F0906F56FD37" box="[1162,1331,705,730]" italics="true" pageId="21" pageNumber="22">V. pastorpueri</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C18B0CFF707FC48FF72F0B56B55FD10" authorityName="Gittenberger" authorityYear="1969" box="[189,304,739,765]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FF72F0B56B55FD10" box="[189,304,739,765]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
. Later,
|
||
<bibRefCitation id="EF89B6BDF707FC48FE5FF0B26819FD13" author="Gittenberger E." box="[400,636,739,766]" firstAuthor="Gittenberger" journalOrPublisher="Basteria" pageId="21" pageNumber="22" pagination="23 - 29" part="79" refId="ref12813" refString="Gittenberger E. 2015. Caving for snails in northern Albania. Basteria 79 (1 - 3): 23 - 29." title="Caving for snails in northern Albania" type="journal article" year="2015">Gittenberger (2015)</bibRefCitation>
|
||
studied a larger series of topotypical
|
||
<taxonomicName id="4C18B0CFF707FC48FB86F0B56E97FD10" box="[1097,1266,740,765]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pastorpueri">
|
||
<emphasis id="B96C175EF707FC48FB86F0B56E97FD10" box="[1097,1266,740,765]" italics="true" pageId="21" pageNumber="22">V. pastorpueri</emphasis>
|
||
</taxonomicName>
|
||
specimens and concluded that their shells are neither smaller nor relatively higher and thicker discoid than those of typical
|
||
<taxonomicName id="4C18B0CFF707FC48FEDCF17B6BE7FCAE" authorityName="Gittenberger" authorityYear="1969" box="[275,386,809,835]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FEDCF17B6BE7FCAE" box="[275,386,809,835]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
. However, he found that the ribbing of
|
||
<taxonomicName id="4C18B0CFF707FC48FC83F17B6994FCAE" box="[844,1009,810,835]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pastorpueri">
|
||
<emphasis id="B96C175EF707FC48FC83F17B6994FCAE" box="[844,1009,810,835]" italics="true" pageId="21" pageNumber="22">V. pastorpueri</emphasis>
|
||
</taxonomicName>
|
||
is less dense (less than 70 ribs on the last whorl) and therefore he preserved it as a distinct subspecies,
|
||
<taxonomicName id="4C18B0CFF707FC48FC2EF11C6E84FC8B" authorityName="- Gittenberger" authorityYear="2015" box="[993,1249,844,870]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="ripkeni" subSpecies="pastorpueri">
|
||
<emphasis id="B96C175EF707FC48FC2EF11C6E84FC8B" box="[993,1249,844,870]" italics="true" pageId="21" pageNumber="22">V. ripkeni pastorpueri</emphasis>
|
||
</taxonomicName>
|
||
. In the same paper, Gittenberger described another subspecies,
|
||
<taxonomicName id="4C18B0CFF707FC48FCDCF12069BEFC67" authorityName="Gittenberger" authorityYear="2015" box="[787,987,880,906]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="ripkeni" subSpecies="arbeni">
|
||
<emphasis id="B96C175EF707FC48FCDCF12069BEFC67" box="[787,987,880,906]" italics="true" pageId="21" pageNumber="22">V. ripkeni arbeni</emphasis>
|
||
</taxonomicName>
|
||
, which is smaller (H
|
||
<quantity id="4CE066A9F707FC48FB2FF1216F17FC67" box="[1248,1394,880,906]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="21" pageNumber="22" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
|
||
; W 3.0–
|
||
<quantity id="4CE066A9F707FC48FEDAF1C26B0AFC40" box="[277,367,915,941]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="21" pageNumber="22" unit="mm" value="3.1">3.1 mm</quantity>
|
||
) and has a weakly developed peristome.
|
||
</paragraph>
|
||
<paragraph id="8BA7CB4CF707FC48FF72F18D6E21FA97" blockId="21.[189,1399,988,1402]" pageId="21" pageNumber="22">
|
||
We collected several hundred specimens of this species at four different locations in the Renc Mts. Their shell width ranges between 2.7 and
|
||
<quantity id="4CE066A9F707FC48FD94F1AE68D2FBF7" box="[603,695,1023,1050]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="21" pageNumber="22" unit="mm" value="3.6">3.6 mm</quantity>
|
||
, shell height between 1.5 and
|
||
<quantity id="4CE066A9F707FC48FBDDF1AE6E0EFBF7" box="[1042,1131,1023,1050]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="21" pageNumber="22" unit="mm" value="2.1">2.1 mm</quantity>
|
||
and the number of ribs on the last whorl between 60 and 82. Compared to topotypical
|
||
<taxonomicName id="4C18B0CFF707FC48FC67F6726E2BFBD1" box="[936,1102,1059,1084]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pastorpueri">
|
||
<emphasis id="B96C175EF707FC48FC67F6726E2BFBD1" box="[936,1102,1059,1084]" italics="true" pageId="21" pageNumber="22">V. pastorpueri</emphasis>
|
||
</taxonomicName>
|
||
, their size, shape and rib density values largely overlap. Although one might say that they are generally somewhat smaller and more tapered than typical
|
||
<taxonomicName id="4C18B0CFF707FC48FE38F63B680CFB6E" authorityName="Gittenberger" authorityYear="1969" box="[503,617,1129,1155]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FE38F63B680CFB6E" box="[503,617,1129,1155]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
, neither these characters nor the rib density are able to provide a clear-cut distinction. This large variability in rib density refutes Gittenberger’s argument (
|
||
<bibRefCitation id="EF89B6BDF707FC48FB2EF6DD6A99FB27" author="Gittenberger E." firstAuthor="Gittenberger" journalOrPublisher="Basteria" pageId="21" pageNumber="22" pagination="23 - 29" part="79" refId="ref12813" refString="Gittenberger E. 2015. Caving for snails in northern Albania. Basteria 79 (1 - 3): 23 - 29." title="Caving for snails in northern Albania" type="journal article" year="2015">Gittenberger 2015</bibRefCitation>
|
||
), and makes any distinction between
|
||
<taxonomicName id="4C18B0CFF707FC48FD7EF6E16933FB24" box="[689,854,1200,1225]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pastorpueri">
|
||
<emphasis id="B96C175EF707FC48FD7EF6E16933FB24" box="[689,854,1200,1225]" italics="true" pageId="21" pageNumber="22">V. pastorpueri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C18B0CFF707FC48FC40F6E1699BFB24" authorityName="Gittenberger" authorityYear="1969" box="[911,1022,1199,1225]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FC40F6E1699BFB24" box="[911,1022,1199,1225]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
meaningless. However, none of the many adult specimens from the Renc Mts have such a weakly developed peristome as in the three specimens that were described as
|
||
<taxonomicName id="4C18B0CFF707FC48FD9BF6A76979FAE2" authorityName="Gittenberger" authorityYear="2015" box="[596,796,1269,1295]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="ripkeni" subSpecies="arbeni">
|
||
<emphasis id="B96C175EF707FC48FD9BF6A76979FAE2" box="[596,796,1269,1295]" italics="true" pageId="21" pageNumber="22">V. ripkeni arbeni</emphasis>
|
||
</taxonomicName>
|
||
. Having no more material available from the
|
||
<typeStatus id="54A375EEF707FC48FA8BF6A76F12FAFD" box="[1348,1399,1270,1296]" pageId="21" pageNumber="22">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C18B0CFF707FC48FE8DF74B682FFAD9" authorityName="Gittenberger" authorityYear="2015" box="[322,586,1305,1332]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="ripkeni" subSpecies="arbeni">
|
||
<emphasis id="B96C175EF707FC48FE8DF74B6BADFADE" box="[322,456,1305,1331]" italics="true" pageId="21" pageNumber="22">V. r. arbeni</emphasis>
|
||
(
|
||
<collectingRegion id="49DC05AEF707FC48FE15F7486827FAD9" box="[474,578,1305,1332]" country="Albania" name="Shkoder" pageId="21" pageNumber="22">Shkodër</collectingRegion>
|
||
)
|
||
</taxonomicName>
|
||
, it is not possible to test whether the entire population consistently has an underdeveloped peristome there or the subspecies was described on the basis of three freak individuals. Until this question is settled, we must preserve
|
||
<taxonomicName id="4C18B0CFF707FC48FCB6F7316E58FA94" authorityName="Gittenberger" authorityYear="2015" box="[889,1085,1375,1401]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="ripkeni" subSpecies="arbeni">
|
||
<emphasis id="B96C175EF707FC48FCB6F7316E58FA94" box="[889,1085,1375,1401]" italics="true" pageId="21" pageNumber="22">V. ripkeni arbeni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7CB4CF707FC4BFF72F7F9689EFE5F" blockId="21.[189,1399,1448,2003]" lastBlockId="22.[189,1399,267,435]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
The type material of
|
||
<taxonomicName id="4C18B0CFF707FC48FE05F7F86825FA2F" authorityName="Gittenberger" authorityYear="1969" box="[458,576,1448,1474]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FE05F7F86825FA2F" box="[458,576,1448,1474]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
was presumably collected by Petar Dabović in the 1930s, passed somehow to the Viennese private collector Aemilian Edlauer, and finally landed in the NHMW Mollusc Collection. It appears that Edlauer, or someone else who studied his collection, recognized, but never described, three different species of
|
||
<taxonomicName id="4C18B0CFF707FC48FD49F4426966F9C1" authorityName="Gittenberger" authorityYear="1969" box="[646,771,1555,1580]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B96C175EF707FC48FD49F4426966F9C1" box="[646,771,1555,1580]" italics="true" pageId="21" pageNumber="22">Virpazaria</emphasis>
|
||
</taxonomicName>
|
||
. The proposed names were ʻ
|
||
<taxonomicName id="4C18B0CFF707FC48FBBAF4436F03F9C1" authorityName="Fehér & Deli & Erőss & Lika" authorityYear="2019" box="[1141,1382,1554,1580]" class="Gastropoda" family="Trissexodontidae" genus="Trissexodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kusceri">
|
||
<emphasis id="B96C175EF707FC48FBBAF4436F03F9C1" box="[1141,1382,1554,1580]" italics="true" pageId="21" pageNumber="22">Trissexodon kuščeri</emphasis>
|
||
</taxonomicName>
|
||
ʼ, ʻ
|
||
<taxonomicName id="4C18B0CFF707FC48FF08F4646865F9A2" authorityName="Fehér & Deli & Erőss & Lika" authorityYear="2019" box="[199,512,1589,1615]" class="Gastropoda" family="Trissexodontidae" genus="Trissexodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="montenegrina">
|
||
<emphasis id="B96C175EF707FC48FF08F4646865F9A2" box="[199,512,1589,1615]" italics="true" pageId="21" pageNumber="22">Trissexodon montenegrina</emphasis>
|
||
</taxonomicName>
|
||
ʼand ʻ
|
||
<taxonomicName id="4C18B0CFF707FC48FD88F4646956F9A2" authority="Kaufeli" authorityName="Kaufeli" box="[583,819,1589,1615]" class="Gastropoda" family="Trissexodontidae" genus="Trissexodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B96C175EF707FC48FD88F4646956F9A2" box="[583,819,1589,1615]" italics="true" pageId="21" pageNumber="22">Trissexodon Käufeli</emphasis>
|
||
</taxonomicName>
|
||
ʼ. The latter name was obviously proposed for
|
||
<taxonomicName id="4C18B0CFF707FC48FAAEF4676B63F99F" authorityName="Gittenberger" authorityYear="1969" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pageti">
|
||
<emphasis id="B96C175EF707FC48FAAEF4676B63F99F" italics="true" pageId="21" pageNumber="22">V. pageti</emphasis>
|
||
</taxonomicName>
|
||
, but the situation is more dubious with the other two. Among the lots of
|
||
<taxonomicName id="4C18B0CFF707FC48FBB7F4086F64F99F" authorityName="Gittenberger" authorityYear="1969" box="[1144,1281,1624,1650]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="adrianae">
|
||
<emphasis id="B96C175EF707FC48FBB7F4086F64F99F" box="[1144,1281,1624,1650]" italics="true" pageId="21" pageNumber="22">V. adrianae</emphasis>
|
||
</taxonomicName>
|
||
we could find both ʻ
|
||
<taxonomicName id="4C18B0CFF707FC48FEF8F42A6844F978" authorityName="Fehér & Deli & Erőss & Lika" authorityYear="2019" box="[311,545,1659,1685]" class="Gastropoda" family="Trissexodontidae" genus="Trissexodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kusceri">
|
||
<emphasis id="B96C175EF707FC48FEF8F42A6844F978" box="[311,545,1659,1685]" italics="true" pageId="21" pageNumber="22">Trissexodon kuščeri</emphasis>
|
||
</taxonomicName>
|
||
ʼ and ʻ
|
||
<taxonomicName id="4C18B0CFF707FC48FDA2F42A69C1F978" authorityName="Fehér & Deli & Erőss & Lika" authorityYear="2019" box="[621,932,1659,1685]" class="Gastropoda" family="Trissexodontidae" genus="Trissexodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="montenegrina">
|
||
<emphasis id="B96C175EF707FC48FDA2F42A69C1F978" box="[621,932,1659,1685]" italics="true" pageId="21" pageNumber="22">Trissexodon montenegrina</emphasis>
|
||
</taxonomicName>
|
||
’ labels, whereas the name ʻ
|
||
<taxonomicName id="4C18B0CFF707FC48FB26F42A6B77F955" authorityName="Fehér & Deli & Erőss & Lika" authorityYear="2019" class="Gastropoda" family="Trissexodontidae" genus="Trissexodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kusceri">
|
||
<emphasis id="B96C175EF707FC48FB26F42A6B77F955" italics="true" pageId="21" pageNumber="22">Trissexodon kuščeri</emphasis>
|
||
</taxonomicName>
|
||
’ also appears in the material of
|
||
<taxonomicName id="4C18B0CFF707FC48FD44F4CE6899F955" authorityName="Gittenberger" authorityYear="1969" box="[651,764,1694,1720]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FD44F4CE6899F955" box="[651,764,1694,1720]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF89B6BDF707FC48FCC4F4CF6991F954" author="Gittenberger E." box="[779,1012,1694,1721]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="21" pageNumber="22" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger (1969)</bibRefCitation>
|
||
mentioned eight type specimens of
|
||
<taxonomicName id="4C18B0CFF707FC48FF11F4926B2BF931" authorityName="Gittenberger" authorityYear="1969" box="[222,334,1730,1756]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FF11F4926B2BF931" box="[222,334,1730,1756]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
from the same original lot (of which one was designated as a
|
||
<typeStatus id="54A375EEF707FC48FBFEF4936EFEF931" box="[1073,1179,1730,1756]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
, one made its way to the RMNH and six remained in the original vial and kept the original labels (NHMW 77153a)). Interestingly, another site, “Pečina Grbočica bei Virpazar”, is indicated on the original label in that vial, whereas “Soko Höhle bei Dupilo”, which is given in the description, can be found on a newer label written in someone else’s handwriting. Furthermore, there is another lot (NHMW 77154), which is from Soko Cave according to the original label and contains one specimen of
|
||
<taxonomicName id="4C18B0CFF707FC48FBCDF5226E14F861" authorityName="Gittenberger" authorityYear="1969" box="[1026,1137,1906,1932]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF707FC48FBCDF5226E14F861" box="[1026,1137,1906,1932]" italics="true" pageId="21" pageNumber="22">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
, but not mentioned by
|
||
<bibRefCitation id="EF89B6BDF707FC48FF72F5C46BCCF85D" author="Gittenberger E." box="[189,425,1941,1968]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="21" pageNumber="22" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger (1969)</bibRefCitation>
|
||
. Even stranger, this lot contains an original Edlauer label with the same locality, the same voucher number (E-57441) and the same proposed name, ʻ
|
||
<taxonomicName id="4C18B0CFF707FC48FC7EF5E96EFEF83F" authorityName="Fehér & Deli & Erőss & Lika" authorityYear="2019" box="[945,1179,1976,2002]" class="Gastropoda" family="Trissexodontidae" genus="Trissexodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kusceri">
|
||
<emphasis id="B96C175EF707FC48FC7EF5E96EFEF83F" box="[945,1179,1976,2002]" italics="true" pageId="21" pageNumber="22">Trissexodon kusceri</emphasis>
|
||
</taxonomicName>
|
||
’, as the type lot of
|
||
<taxonomicName id="4C18B0CFF704FC4BFF72F35D6B20FEC8" authorityName="Gittenberger" authorityYear="1969" box="[189,325,267,293]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="adrianae">
|
||
<emphasis id="B96C175EF704FC4BFF72F35D6B20FEC8" box="[189,325,267,293]" italics="true" pageId="22" pageNumber="23">V. adrianae</emphasis>
|
||
</taxonomicName>
|
||
. It seems very difficult, if not impossible, to trace back whether there was any mixup with the labels or the specimens in the past, and, if so, where exactly the type specimens of
|
||
<taxonomicName id="4C18B0CFF704FC4BFB0FF37E6F55FEA4" authorityName="Gittenberger" authorityYear="1969" box="[1216,1328,303,329]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF704FC4BFB0FF37E6F55FEA4" box="[1216,1328,303,329]" italics="true" pageId="22" pageNumber="23">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
came from (Soko?, Grbočica? or from somewhere else?). The fact that neither
|
||
<bibRefCitation id="EF89B6BDF704FC4BFBC9F3036E8AFE80" author="Gittenberger E." box="[1030,1263,338,365]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="22" pageNumber="23" pagination="263 - 277" part="48" refId="ref12736" refString="Gittenberger E. 1975. Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag. Zoologische Mededelingen 48 (23): 263 - 277." title="Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag" type="journal article" year="1975">Gittenberger (1975)</bibRefCitation>
|
||
nor anyone else has been able to refind
|
||
<taxonomicName id="4C18B0CFF704FC4BFDC7F327681CFE62" authorityName="Gittenberger" authorityYear="1969" box="[520,633,373,399]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="ripkeni">
|
||
<emphasis id="B96C175EF704FC4BFDC7F327681CFE62" box="[520,633,373,399]" italics="true" pageId="22" pageNumber="23">V. ripkeni</emphasis>
|
||
</taxonomicName>
|
||
at its alleged type locality or anywhere else in the Virpazar area further discredits the validity of the type locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |