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<document id="201EA886C3516C2CBE819AED8DD45662" ID-DOI="10.1649/072.066.0209" ID-ISSN="1938-4394" ID-Zenodo-Dep="4893146" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1622638001525" checkinUser="carolina" docAuthor="Mora-Aguilar, Eder F. &amp; Delgado, Leonardo" docDate="2012" docId="039B9A4AFFB3FFC8FF1ECE402D2157B6" docLanguage="en" docName="TheColeopBull.66.2.139-142.pdf.imf" docOrigin="The Coleopterists Bulletin 66 (2)" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Cyclocephala berti Delgado 1992" docType="treatment" docVersion="3" lastPageNumber="142" masterDocId="FFA2E232FFB0FFCBFFFFCC4B2C035177" masterDocTitle="A New Species of Cyclocephala Dejean (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) from the Cloud Forests of Southeastern Mexico and Description of the Female of Cyclocephala berti Delgado" masterLastPageNumber="142" masterPageNumber="139" pageNumber="142" updateTime="1698952918332" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="77B5D0B5EBB2FD6A18C1A8AD2A325DAA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3119A14B24CD58B32C61AE0CE7054316">
<mods:title id="057506B49791C49EFD831D49FE0471D7">A New Species of Cyclocephala Dejean (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) from the Cloud Forests of Southeastern Mexico and Description of the Female of Cyclocephala berti Delgado</mods:title>
</mods:titleInfo>
<mods:name id="8129069050FF0C86B738977579D7F573" type="personal">
<mods:role id="D1B90BB35DFBD69475EA1D3A75FD68BB">
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</mods:role>
<mods:namePart id="8902EAF31509C6B25DFA92F2DF2CF2E7">Mora-Aguilar, Eder F.</mods:namePart>
<mods:affiliation id="EB4B8DC12EF906C2FCCD159802775121">Instituto de Ecología A. C., Carretera antigua a Coatepec No. 351 91070 Xalapa, Veracruz, MEXICO</mods:affiliation>
<mods:nameIdentifier id="80E7951F64D9D42D556B432FEC58040F" type="email">edynastes@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7FCF77E28B8AB33C19B5FB1887FA1063" type="personal">
<mods:role id="0C8448D9E18F442486EB78353B6571B8">
<mods:roleTerm id="14A0B3F3487505A94ED34BB491EE04EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="121E7670EBACA5720FDD36713244D3E7">Delgado, Leonardo</mods:namePart>
<mods:affiliation id="ECD619D6AE756097208C22DD39539C95">Instituto de Ecología A. C., Carretera antigua a Coatepec No. 351 91070 Xalapa, Veracruz, MEXICO</mods:affiliation>
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<mods:titleInfo id="369974C6FE4DEB9494AD4885D2483C9E">
<mods:title id="1B16FC6014038BC6D60E3D141364DC55">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="B994EF90054DBE1318C2F91F8243360F">
<mods:date id="76DBDAEED95E28DF1777CFCFCA63A488">2012</mods:date>
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<mods:number id="27AADFDD19F5303A29141915E544F7D3">2012-06-20</mods:number>
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<mods:number id="6CA96172A083564A5724410922B191B4">66</mods:number>
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<mods:classification id="6FD81C351A2D3D0588C4F3566828077E">journal article</mods:classification>
<mods:identifier id="67471E5BE5552A4A975752E59269C4B9" type="DOI">10.1649/072.066.0209</mods:identifier>
<mods:identifier id="B1C19D7F9B290E12F3BF29C58BE765C8" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="750FE4D5BA3184AB7182B5BFD425852B" type="Zenodo-Dep">4893146</mods:identifier>
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<treatment id="039B9A4AFFB3FFC8FF1ECE402D2157B6" ID-DOI="http://doi.org/10.5281/zenodo.4893136" ID-Zenodo-Dep="4893136" LSID="urn:lsid:plazi:treatment:039B9A4AFFB3FFC8FF1ECE402D2157B6" httpUri="http://treatment.plazi.org/id/039B9A4AFFB3FFC8FF1ECE402D2157B6" lastPageNumber="142" pageId="3" pageNumber="142">
<subSubSection id="C32878D7FFB3FFC8FF1ECE402E375357" box="[225,564,523,544]" pageId="3" pageNumber="142" type="nomenclature">
<paragraph id="8B8D2B5CFFB3FFC8FF1ECE402E375357" blockId="3.[225,564,523,544]" box="[225,564,523,544]" pageId="3" pageNumber="142">
<heading id="D0C59C30FFB3FFC8FF1ECE402E375357" box="[225,564,523,544]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="142" reason="2">
<taxonomicName id="4C3250DFFFB3FFC8FF1ECE402E375357" authority="Delgado, 1992" authorityName="Delgado" authorityYear="1992" box="[225,564,523,544]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="berti">
<emphasis id="B946F74EFFB3FFC8FF1ECE402E375357" bold="true" box="[225,564,523,544]" pageId="3" pageNumber="142">
<emphasis id="B946F74EFFB3FFC8FF1ECE402D995357" bold="true" box="[225,410,523,544]" italics="true" pageId="3" pageNumber="142">Cyclocephala berti</emphasis>
<bibRefCitation id="EFA356ADFFB3FFC8FE5ECE402E375357" author="Delgado" box="[417,564,523,544]" pageId="3" pageNumber="142" pagination="75 - 78" refId="ref2813" refString="Delgado, L. 1992. Una especie nueva mexicana de Cyclocephala (Coleoptera: Melolonthidae; Dynastinae). Anales del Instituto de Biologia de la Universidad Nacional Autonoma de Mexico, Serie Zoologia 63 (1): 75 - 78." type="journal article" year="1992">Delgado, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32878D7FFB3FFC8FF50CE7D2D2157B6" pageId="3" pageNumber="142" type="materials_examined">
<paragraph id="8B8D2B5CFFB3FFC8FF50CE7D2D3C52FD" blockId="3.[151,639,566,1729]" pageId="3" pageNumber="142">
<emphasis id="B946F74EFFB3FFC8FF50CE7D2D80533C" bold="true" box="[175,387,566,587]" pageId="3" pageNumber="142">Material Examined.</emphasis>
<materialsCitation id="3B5A2101FFB3FFC8FE6FCE7D2CE35311" location="Two" pageId="3" pageNumber="142" specimenCount="1" specimenCount-female="1">
<location id="8EED7D87FFB3FFC8FE6FCE7D2DBD533C" LSID="urn:lsid:plazi:treatment:039B9A4AFFB3FFC8FF1ECE402D2157B6:8EED7D87FFB3FFC8FE6FCE7D2DBD533C" box="[400,446,566,587]" name="Two" pageId="3" pageNumber="142">Two</location>
males and
<specimenCount id="9D34E0D5FFB3FFC8FDACCE7D2CE35311" pageId="3" pageNumber="142" type="female">five females</specimenCount>
</materialsCitation>
<materialsCitation id="3B5A2101FFB3FFC8FF15CE1A2D19524C" collectingDate="2011-05-29" collectingDateMax="2011-07-31" collectingDateMin="2011-05-29" collectorName="E. Mora &amp; L. Delgado &amp; L. Lara" country="Mexico" county="Zongolica" elevation="1250" location="Amatitla." municipality="La Quinta" pageId="3" pageNumber="142" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave">
labeled: “
<collectingCountry id="F3256BCCFFB3FFC8FEB7CE1A2DAA5311" box="[328,425,593,614]" name="Mexico" pageId="3" pageNumber="142">MÉXICO</collectingCountry>
:
<collectingRegion id="49F6E5BEFFB3FFC8FE4ACE1A2E0F5311" box="[437,524,593,614]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="142">Veracruz</collectingRegion>
,
<collectingCounty id="62EC53D0FFB3FFC8FDE8CE1A2E795311" box="[535,634,593,614]" pageId="3" pageNumber="142">Zongolica</collectingCounty>
,
<collectingMunicipality id="6BE9B126FFB3FFC8FF68CE202CFC53F7" box="[151,255,619,640]" pageId="3" pageNumber="142">La Quinta</collectingMunicipality>
,
<location id="8EED7D87FFB3FFC8FEF2CE202D6A53F7" LSID="urn:lsid:plazi:treatment:039B9A4AFFB3FFC8FF1ECE402D2157B6:8EED7D87FFB3FFC8FEF2CE202D6A53F7" box="[269,361,619,640]" country="Mexico" county="Zongolica" municipality="La Quinta" name="Amatitla." pageId="3" pageNumber="142" stateProvince="Veracruz de Ignacio de la Llave">Amatitla.</location>
<date id="FF8C0D9CFFB3FFC8FE8DCE202DFE53F7" box="[370,509,619,640]" pageId="3" pageNumber="142" value="2011-05-29" valueMax="2011-05-30" valueMin="2011-05-29">
<collectingDate id="EFC8F474FFB3FFC8FE8DCE202DFE53F7" box="[370,509,619,640]" pageId="3" pageNumber="142" value="2011-05-29" valueMax="2011-05-30" valueMin="2011-05-29">29-30-V-2011</collectingDate>
</date>
.
<location id="8EED7D87FFB3FFC8FDF5CE202E2C53F7" LSID="urn:lsid:plazi:treatment:039B9A4AFFB3FFC8FF1ECE402D2157B6:8EED7D87FFB3FFC8FDF5CE202E2C53F7" box="[522,559,619,640]" country="Mexico" county="Zongolica" municipality="La Quinta" name="Luz" pageId="3" pageNumber="142" stateProvince="Veracruz de Ignacio de la Llave">Luz</location>
mercurial.
<location id="8EED7D87FFB3FFC8FF38CECD2D0C53EC" LSID="urn:lsid:plazi:treatment:039B9A4AFFB3FFC8FF1ECE402D2157B6:8EED7D87FFB3FFC8FF38CECD2D0C53EC" box="[199,271,646,667]" country="Mexico" county="Zongolica" municipality="La Quinta" name="Bosque" pageId="3" pageNumber="142" stateProvince="Veracruz de Ignacio de la Llave">Bosque</location>
mesófilo de montaña.
<location id="8EED7D87FFB3FFC8FE03CECD2E2253EC" LSID="urn:lsid:plazi:treatment:039B9A4AFFB3FFC8FF1ECE402D2157B6:8EED7D87FFB3FFC8FE03CECD2E2253EC" box="[508,545,646,667]" country="Mexico" county="Zongolica" municipality="La Quinta" name="Alt." pageId="3" pageNumber="142" stateProvince="Veracruz de Ignacio de la Llave">Alt.</location>
<quantity id="4CCA86B9FFB3FFC8FDD4CECD2E7953EC" box="[555,634,646,667]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="142" unit="m" value="1250.0">
<elevation id="001FCC6FFFB3FFC8FDD4CECD2E7953EC" box="[555,634,646,667]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="142" unit="m" value="1250.0">1250 m</elevation>
</quantity>
,
<collectorName id="26C74E8AFFB3FFC8FF68CEEB2CE953C2" box="[151,234,672,693]" pageId="3" pageNumber="142">E. Mora</collectorName>
,
<collectorName id="26C74E8AFFB3FFC8FF09CEEB2D6153C2" box="[246,354,672,693]" pageId="3" pageNumber="142">L. Delgado</collectorName>
y
<collectorName id="26C74E8AFFB3FFC8FE80CEEB2DC453C2" box="[383,455,672,693]" pageId="3" pageNumber="142">L. Lara</collectorName>
cols”. One female with same data, except
<date id="FF8C0D9CFFB3FFC8FE6CCEF02E0F53A7" box="[403,524,699,720]" pageId="3" pageNumber="142" value="2011-06-31">
<collectingDate id="EFC8F474FFB3FFC8FE6CCEF02E0F53A7" box="[403,524,699,720]" pageId="3" pageNumber="142" value="2011-06-31">31-VI-2011</collectingDate>
</date>
. One male with same data, except
<date id="FF8C0D9CFFB3FFC8FE75CE9E2E05539D" box="[394,518,725,746]" pageId="3" pageNumber="142" value="2011-07-31">
<collectingDate id="EFC8F474FFB3FFC8FE75CE9E2E05539D" box="[394,518,725,746]" pageId="3" pageNumber="142" value="2011-07-31">31-VII-2011</collectingDate>
</date>
. The specimens are deposited in the following collections: University of Nebraska State Museum (Nebraska, USA)
<specimenCount id="9D34E0D5FFB3FFC8FF2BCF6E2CF4524C" box="[212,247,805,827]" pageId="3" pageNumber="142" type="male">1 ♂</specimenCount>
<specimenCount id="9D34E0D5FFB3FFC8FF03CF6E2D19524C" box="[252,282,805,827]" pageId="3" pageNumber="142" type="female">1♀</specimenCount>
</materialsCitation>
; Instituto de Ecología, A. C. (Xalapa, México)
<specimenCount id="9D34E0D5FFB3FFC8FF10CF0B2D0F5222" box="[239,268,831,853]" pageId="3" pageNumber="142" type="female">2♀</specimenCount>
; L. Delgado Collection (Mexico City)
<specimenCount id="9D34E0D5FFB3FFC8FF68CF112CBE5207" box="[151,189,858,880]" pageId="3" pageNumber="142" type="male">1 ♂</specimenCount>
<specimenCount id="9D34E0D5FFB3FFC8FF3ACF112CE05207" box="[197,227,858,880]" pageId="3" pageNumber="142" type="female">1♀</specimenCount>
; and E. Mora-Aguilar Collection (Xalapa, México)
<specimenCount id="9D34E0D5FFB3FFC8FF10CF3E2D1752FD" box="[239,276,884,906]" pageId="3" pageNumber="142" type="male">1 ♂</specimenCount>
<specimenCount id="9D34E0D5FFB3FFC8FEE4CF3E2D3A52FD" box="[283,313,884,906]" pageId="3" pageNumber="142" type="female">2♀</specimenCount>
.
</paragraph>
<paragraph id="8B8D2B5CFFB3FFC8FF50CFC42D685444" blockId="3.[151,639,566,1729]" pageId="3" pageNumber="142">
<materialsCitation id="3B5A2101FFB3FFC8FF50CFC42D675445" collectorName="H. &amp; A. Howden &amp; de Zongolica &amp; Male &amp; Female" country="Canada" county="Canadian Museum of Nature" location="Volcan de San Martin" municipality="Los Tuxtlas region" pageId="3" pageNumber="142" specimenCount="2" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave">
This species was described based on only
<specimenCount id="9D34E0D5FFB3FFC8FDA4CFC42CC652C8" pageId="3" pageNumber="142" type="male">one male</specimenCount>
specimen from the
<collectorName id="26C74E8AFFB3FFC8FE67CFE12E4E52C8" box="[408,589,938,959]" pageId="3" pageNumber="142">H. &amp; A. Howden</collectorName>
collection (now in
<collectingCounty id="62EC53D0FFB3FFC8FEB8CF8F2E7952AE" box="[327,634,964,985]" pageId="3" pageNumber="142">Canadian Museum of Nature</collectingCounty>
, Ottawa,
<collectingCountry id="F3256BCCFFB3FFC8FF13CF942D3A5283" box="[236,313,991,1012]" name="Canada" pageId="3" pageNumber="142">Canada</collectingCountry>
). The specimen was collected in the
<collectingMunicipality id="6BE9B126FFB3FFC8FF41CFB12D7B5578" box="[190,376,1018,1039]" pageId="3" pageNumber="142">Los Tuxtlas region</collectingMunicipality>
on the southeastern slope of the
<location id="8EED7D87FFB3FFC8FF22C85F2DB8555E" LSID="urn:lsid:plazi:treatment:039B9A4AFFB3FFC8FF1ECE402D2157B6:8EED7D87FFB3FFC8FF22C85F2DB8555E" box="[221,443,1044,1065]" country="Canada" county="Canadian Museum of Nature" municipality="Los Tuxtlas region" name="Volcan de San Martin" pageId="3" pageNumber="142" stateProvince="Veracruz de Ignacio de la Llave">Volcán de San Martín</location>
,
<collectingRegion id="49F6E5BEFFB3FFC8FE36C85F2E23555E" box="[457,544,1044,1065]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="142">Veracruz</collectingRegion>
,
<collectingCountry id="F3256BCCFFB3FFC8FDD1C85F2E79555E" box="[558,634,1044,1065]" name="Mexico" pageId="3" pageNumber="142">Mexico</collectingCountry>
. Recently,
<specimenCount id="9D34E0D5FFB3FFC8FF00C8642D995533" box="[255,410,1071,1092]" pageId="3" pageNumber="142" type="generic">nine specimens</specimenCount>
were collected at the
<location id="8EED7D87FFB3FFC8FF68C8022D675529" LSID="urn:lsid:plazi:treatment:039B9A4AFFB3FFC8FF1ECE402D2157B6:8EED7D87FFB3FFC8FF68C8022D675529" box="[151,356,1097,1118]" country="Canada" county="Canadian Museum of Nature" municipality="Los Tuxtlas region" name="Sierra de Zongolica" pageId="3" pageNumber="142" stateProvince="Veracruz de Ignacio de la Llave">
Sierra
<collectorName id="26C74E8AFFB3FFC8FF22C8022D675529" box="[221,356,1097,1118]" pageId="3" pageNumber="142">de Zongolica</collectorName>
</location>
,
<collectingRegion id="49F6E5BEFFB3FFC8FE8CC8022DCE5529" box="[371,461,1097,1118]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="142">Veracruz</collectingRegion>
.
<collectorName id="26C74E8AFFB3FFC8FE24C8022E0E5529" box="[475,525,1097,1118]" pageId="3" pageNumber="142">Male</collectorName>
specimens agree with the description of
<taxonomicName id="4C3250DFFFB3FFC8FE40C8282E08550F" box="[447,523,1123,1144]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="berti">
<emphasis id="B946F74EFFB3FFC8FE40C8282E08550F" box="[447,523,1123,1144]" italics="true" pageId="3" pageNumber="142">C. berti</emphasis>
</taxonomicName>
, except for their larger size (length
<quantity id="4CCA86B9FFB3FFC8FE63C8342E3555E3" box="[412,566,1150,1172]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.68" metricValueMin="1.3699999999999999" pageId="3" pageNumber="142" unit="mm" value="15.25" valueMax="16.8" valueMin="13.7">13.716.8 mm</quantity>
, width across the humeri
<quantity id="4CCA86B9FFB3FFC8FEAFC8D22DCA55D9" box="[336,457,1176,1198]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.3" metricValueMin="6.5" pageId="3" pageNumber="142" unit="mm" value="6.9" valueMax="7.3" valueMin="6.5">6.57.3 mm</quantity>
), head completely covered with long and erect setae, and each elytron with 3 or 4 diffuse spots.
<collectorName id="26C74E8AFFB3FFC8FE33C8852E125594" box="[460,529,1230,1251]" pageId="3" pageNumber="142">Female</collectorName>
specimens collected are similar to males except in the dimorphic characters, and are different from other species of
<taxonomicName id="4C3250DFFFB3FFC8FF1DC9562D675445" box="[226,356,1309,1330]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B946F74EFFB3FFC8FF1DC9562D675445" box="[226,356,1309,1330]" italics="true" pageId="3" pageNumber="142">Cyclocephala</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8D2B5CFFB3FFC8FF50C9732D2157B6" blockId="3.[151,639,566,1729]" pageId="3" pageNumber="142">
Description of the previously unknown female of
<taxonomicName id="4C3250DFFFB3FFC8FF4EC9192CFB5410" box="[177,248,1362,1383]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="berti">
<emphasis id="B946F74EFFB3FFC8FF4EC9192CFB5410" box="[177,248,1362,1383]" italics="true" pageId="3" pageNumber="142">C. berti</emphasis>
</taxonomicName>
is as follows: Total length
<quantity id="4CCA86B9FFB3FFC8FE0EC9182E79541F" box="[497,634,1362,1384]" metricMagnitude="-2" metricUnit="m" metricValue="1.395" metricValueMax="1.48" metricValueMin="1.31" pageId="3" pageNumber="142" unit="mm" value="13.95" valueMax="14.8" valueMin="13.1">13.114.8 mm</quantity>
, width across the humeri
<quantity id="4CCA86B9FFB3FFC8FE6CC9252E0D54F4" box="[403,526,1389,1411]" metricMagnitude="-3" metricUnit="m" metricValue="6.45" metricValueMax="6.7" metricValueMin="6.2" pageId="3" pageNumber="142" unit="mm" value="6.45" valueMax="6.7" valueMin="6.2">6.26.7 mm</quantity>
. Similar to male except in the following respects: Head and pronotum glabrous. Each elytron with 34 small, fuscous spots. Elytra and scutellum glabrous, except for the apex and posterior lateral third of elytra with small, sparse setae. Epipleuron expanded at level of metacoxa to abdominal sternites 23, with small, rounded, obtuse tooth between abdominal sternites 23. Pygidium almost flat, glabrous, except with small, erect setae on the base. Protibiae with basal tooth normal, teeth equidistant. Protarsus not enlarged, claws simple. Metatarsi slightly longer than metatibia.
</paragraph>
</subSubSection>
</treatment>
</document>