143 lines
17 KiB
XML
143 lines
17 KiB
XML
<document id="8917B5F0CD9DD080CD6A6F32CB9065F3" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA242ABD72FF37DF831EA3A257" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma congesta Wendland ex Spruce 1871" docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="53" updateTime="1698765326825" updateUser="ExternalLinkService">
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<mods:mods id="43FFDA1E699DC8FE4AA64C8500127B65" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B091B0E654940528E60DD0D5ACC38836">
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<mods:title id="D4E6CAC0954E944E85DF12FC37247C65">A revision of Geonoma (Arecaceae)</mods:title>
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</mods:titleInfo>
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<mods:name id="66A767CC4E0472A85516F4361EA223A0" type="personal">
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<mods:role id="62457DF0C14506BFB709AC0CC8542092">
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<mods:roleTerm id="E4D7FE737B3923A1CFAFFDC3451D3E88">Author</mods:roleTerm>
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<mods:namePart id="E7DC104F7F2D228E9B718FE4C6CEF438">Henderson, Andrew</mods:namePart>
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<mods:typeOfResource id="4F8B8CE6AC9198F236C8BA6CBFFCC8A5">text</mods:typeOfResource>
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<mods:titleInfo id="FB05FB1C46C2F462CBFAF81780E9279E">
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<mods:title id="36E7949A93502A42EB64A1260E57002F">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="052172F3DB0B6D38B489148B6FAB47BF">
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<mods:date id="1685812D9080AA8D1546121A4BFA1447">2011</mods:date>
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<mods:detail id="20EB816A775D4FAD77726BECD19381CE" type="pubDate">
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<mods:number id="791ADAF2471CDC2F6677AC173C2C7F5A">2011-02-18</mods:number>
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<mods:detail id="B1074022937D084C63948FAEA91CE70C" type="volume">
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<mods:number id="CE4CFC182324F0DAA02B11F93EAA02B4">17</mods:number>
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<mods:classification id="A73BA9A897EBD4256C04573570C08C5A">journal article</mods:classification>
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<mods:identifier id="5D4EEEA4D9A2F89C418708A3D1FAA0FB" type="CLB-Dataset">24911</mods:identifier>
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<mods:identifier id="85C0E53CD5AE5C03B9ED799178D0C0F5" type="DOI">10.11646/phytotaxa.17.1</mods:identifier>
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<mods:identifier id="AF9D1AD8CB4D48D304670C0622F21822" type="GBIF-Dataset">65962ba7-eec1-40e7-aed6-cef94e99ca39</mods:identifier>
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<mods:identifier id="532475E471FF737C840B9E258640FBCC" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="9D9CF227528E3E5DC3B0593192FC0F0B" type="Zenodo-Dep">3538362</mods:identifier>
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<treatment id="FA4887FA242ABD72FF37DF831EA3A257" ID-DOI="http://doi.org/10.5281/zenodo.5608938" ID-GBIF-Taxon="160342489" ID-Zenodo-Dep="5608938" LSID="urn:lsid:plazi:treatment:FA4887FA242ABD72FF37DF831EA3A257" httpUri="http://treatment.plazi.org/id/FA4887FA242ABD72FF37DF831EA3A257" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
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<subSubSection id="3AFB6567242ABD73FF37DF8318C9A100" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="725E36EC242ABD73FF37DF8318C9A100" blockId="52.[151,1437,448,514]" pageId="52" pageNumber="53">
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<emphasis id="4095EAFE242ABD73FF37DF831B82A2DA" bold="true" box="[151,189,449,475]" pageId="52" pageNumber="53">13.</emphasis>
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<taxonomicName id="B5E14D6F242ABD73FF66DF821865A2DA" ID-CoL="3FQHT" authority="Wendland ex Spruce (1871: 112)" authorityName="Wendland ex Spruce" authorityPageNumber="112" authorityYear="1871" box="[198,858,448,475]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="congesta">
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<emphasis id="4095EAFE242ABD73FF66DF821A89A2DB" bold="true" box="[198,438,448,474]" italics="true" pageId="52" pageNumber="53">Geonoma congesta</emphasis>
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Wendland ex
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<bibRefCitation id="16704B1D242ABD73FDC8DF831865A2DA" author="Spruce R." box="[616,858,449,475]" journalOrPublisher="Journal Linnaean Society, Botany" pageId="52" pageNumber="53" pagination="65 - 183" part="11" refId="ref124389" refString="Spruce R. (1871) Palmae Amazonicae. Journal Linnaean Society, Botany 11: 65 - 183." title="Palmae Amazonicae" type="journal article" year="1871">Spruce (1871: 112)</bibRefCitation>
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</taxonomicName>
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. Type:
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<collectingCountry id="0AF6767C242ABD73FC15DF831F54A2DA" box="[949,1131,449,475]" name="Costa Rica" pageId="52" pageNumber="53">COSTA RICA</collectingCountry>
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.
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<collectingRegion id="B025F80E242ABD73FBD8DF831FE1A2DA" box="[1144,1246,449,475]" country="Costa Rica" name="Heredia" pageId="52" pageNumber="53">Heredia</collectingRegion>
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: Río Sarapiquí, between Pedegral and San Miguel, 1857,
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<emphasis id="4095EAFE242ABD73FDDFDFAB1878A103" box="[639,839,487,514]" italics="true" pageId="52" pageNumber="53">H. Wendland s.n.</emphasis>
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(
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<typeStatus id="AD5A884E242ABD73FCF6DFA51881A100" box="[854,958,487,513]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
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K!).
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</paragraph>
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</subSubSection>
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<subSubSection id="3AFB6567242ABD73FF37DC76195EA470" pageId="52" pageNumber="53" type="description">
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<paragraph id="725E36EC242ABD73FF37DC7619FEA6EF" blockId="52.[151,1438,564,1906]" pageId="52" pageNumber="53">
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<emphasis id="4095EAFE242ABD73FF37DC761BDDA14F" box="[151,226,564,590]" italics="true" pageId="52" pageNumber="53">Plants</emphasis>
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3.4(1.5–6.0) m tall; stems 4.1(1.0–8.0) m tall, 1.6(1.0–2.7) cm in diameter, solitary or clustered, canelike; internodes 2.0(0.9–5.5) cm long, yellowish and smooth.
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<emphasis id="4095EAFE242ABD73FC04DC1F18C4A177" box="[932,1019,605,630]" italics="true" pageId="52" pageNumber="53">Leaves</emphasis>
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10(6–13) per stem, undivided or irregularly pinnate, not plicate, bases of blades running diagonally into the rachis; sheaths 23.2(15.5–27.0) cm long; petioles 18.7(5.5–51.0) cm long, drying green or yellowish; rachis 66.1(38.8–132.0) cm long, 5.1(2.4– 15.2) mm in diameter; veins raised and rectangular in cross-section adaxially; pinnae 4(1–24) per side of rachis; basal pinna 59.2(33.0–82.0) cm long, 18.0(4.5–35.5) cm wide, forming an angle of 25(15–38)° with the rachis; apical pinna 34.9(21.7–48.0) cm long, 20.7(7.1–42.0) cm wide, forming an angle of 26(20–32)° with the rachis.
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<emphasis id="4095EAFE242ABD73FEF0DD011AC7A05C" box="[336,504,835,861]" italics="true" pageId="52" pageNumber="53">Inflorescences</emphasis>
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branched 1–3 orders; prophylls and peduncular bracts not ribbed with elongate, unbranched fibers, flattened, deciduous or persistent; prophylls 15.9(6.0–29.7) cm long, not short and asymmetrically apiculate, the surfaces ridged with close, equal, parallel, non-dividing ridges, scarcely tomentose between the ridges, without unequally wide ridges; peduncular bracts 16.5(13.0–26.5) cm long, well-developed, inserted 0.6(0.3–1.0) cm above the prophyll; peduncles 7.8(4.2–13.0) cm long, 6.2(3.4–9.8) mm in diameter; rachillae 9(3–16), 12.8(6.0–23.0) cm long, 5.8(4.1–7.2) mm in diameter, the surfaces without spiky, fibrous projections or ridges, drying brown or yellow-brown, without short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits spirally arranged, glabrous internally; proximal lips without a central notch before anthesis, not recurved after anthesis, not hood-shaped; proximal and distal lips drying the same color as the rachillae, not joined to form a raised cupule, the proximal lip margins overlapping the distal lip margins; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted almost directly onto the filament apices, the connectives bifid but scarcely developed; anthers short and curled over at anthesis; non-fertilized pistillate flowers persistent after anthesis; staminodial tubes crenulate at the apex, those of non-fertilized flowers projecting and persistent after anthesis;
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<emphasis id="4095EAFE242ABD73FB44DB231E1DA67A" box="[1252,1314,1377,1403]" italics="true" pageId="52" pageNumber="53">fruits</emphasis>
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12.4(9.7– 15.2) mm long, 9.8(8.1–11.7) mm in diameter, the bases with a prominent, asymmetric stipe, the apices not conical, the surfaces not splitting at maturity, with fibers emerging, not bumpy, not apiculate; locular epidermis with operculum, smooth, with pores.
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</paragraph>
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<paragraph id="725E36EC242ABD73FF66DBB91966A53D" blockId="52.[151,1438,564,1906]" pageId="52" pageNumber="53">
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<emphasis id="4095EAFE242ABD73FF66DBB91AFDA514" bold="true" box="[198,450,1531,1557]" pageId="52" pageNumber="53">Taxonomic notes:—</emphasis>
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<bibRefCitation id="16704B1D242ABD73FE62DBB919DCA514" author="Henderson, A. & Galeano, G. & Bernal, R." box="[450,739,1531,1558]" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="52" pageNumber="53" pagination="1 - 352" refId="ref122861" refString="Henderson, A., Galeano, G. & Bernal, R. (1995) A field guide to the palms of the Americas. Princeton University Press, Princeton, New Jersey, pp. 1 - 352." title="A field guide to the palms of the Americas" type="book chapter" year="1995">
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Henderson
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<emphasis id="4095EAFE242ABD73FDECDBBF19B5A514" box="[588,650,1531,1558]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
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(1995)
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</bibRefCitation>
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included
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<taxonomicName id="B5E14D6F242ABD73FCFCDBB91F9AA514" authorityName="Burret" authorityYear="1930" box="[860,1189,1531,1557]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="calyptrogynoidea">
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<emphasis id="4095EAFE242ABD73FCFCDBB91F9AA514" box="[860,1189,1531,1557]" italics="true" pageId="52" pageNumber="53">Geonoma calyptrogynoidea</emphasis>
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</taxonomicName>
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as a synonym of
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<taxonomicName id="B5E14D6F242ABD73FA23DBB91BC0A53D" authorityName="Wendland ex Spruce" authorityYear="1871" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="congesta">
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<emphasis id="4095EAFE242ABD73FA23DBB91BC0A53D" italics="true" pageId="52" pageNumber="53">G. congesta</emphasis>
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</taxonomicName>
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–see notes under that species.
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</paragraph>
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<paragraph id="725E36EC242ABD73FF66D80B18F0A5B1" blockId="52.[151,1438,564,1906]" pageId="52" pageNumber="53">
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<emphasis id="4095EAFE242ABD73FF66D80B1ACAA562" bold="true" box="[198,501,1609,1635]" pageId="52" pageNumber="53">Subspecific variation:—</emphasis>
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No trait apart from stem branching and leaf division varies within this species. There is geographic discontinuity and there is an isolated population on the Pacific slope of
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<collectingCountry id="0AF6767C242ABD73FB7CD82D1E62A588" box="[1244,1373,1647,1673]" name="Costa Rica" pageId="52" pageNumber="53">Costa Rica</collectingCountry>
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(Osa Peninsula and adjacent areas). There are thus two potential subgroups.
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</paragraph>
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<paragraph id="725E36EC242ABD73FF66D8FF195EA470" blockId="52.[151,1438,564,1906]" pageId="52" pageNumber="53">
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Specimens from Osa differ significantly from other specimens in 10 variables (plant height, rachis length, rachis width, number of pinnae, apical pinna length, peduncle width, rachillae length, rachillae width, number of rachillae, fruit diameter)(
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<emphasis id="4095EAFE242ABD73FE56D9491AC1A425" box="[502,510,1803,1828]" italics="true" pageId="52" pageNumber="53">t</emphasis>
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-test,
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<emphasis id="4095EAFE242ABD73FDE6D9491967A425" box="[582,600,1803,1828]" italics="true" pageId="52" pageNumber="53">P</emphasis>
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<0.05). Specimens from Osa have larger mean values for all these variables. Based on these results, and geographic discontinuity, the two subgroups are recognized as subspecies (subspp.
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<taxonomicName id="B5E14D6F242ABD73FE25D91B1AD2A473" authorityName="Wendland ex Spruce" authorityYear="1871" box="[389,493,1881,1906]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="congesta">
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<emphasis id="4095EAFE242ABD73FE25D91B1AD2A473" box="[389,493,1881,1906]" italics="true" pageId="52" pageNumber="53">congesta</emphasis>
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</taxonomicName>
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,
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<emphasis id="4095EAFE242ABD73FE5CD91B196FA473" box="[508,592,1881,1906]" italics="true" pageId="52" pageNumber="53">osensis</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="3AFB6567242BBD72FF37DEDC1EA3A257" pageId="53" pageNumber="54" type="key">
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<paragraph id="725E36EC242BBD72FF37DEDC196DA3B9" blockId="53.[151,594,158,184]" box="[151,594,158,184]" pageId="53" pageNumber="54">
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<heading id="29168180242BBD72FF37DEDC196DA3B9" bold="true" box="[151,594,158,184]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="2">
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<emphasis id="4095EAFE242BBD72FF37DEDC196DA3B9" bold="true" box="[151,594,158,184]" pageId="53" pageNumber="54">
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Key to the subspecies of
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<taxonomicName id="B5E14D6F242BBD72FE68DEDC196DA3B9" ID-CoL="3FQHT" authorityName="Wendland ex Spruce" authorityYear="1871" box="[456,594,158,184]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="congesta">
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<emphasis id="4095EAFE242BBD72FE68DEDC196DA3B9" bold="true" box="[456,594,158,184]" italics="true" pageId="53" pageNumber="54">G. congesta</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="725E36EC242BBD72FF37DE9C1B9BA3F4" blockId="53.[151,164,222,245]" box="[151,164,222,245]" pageId="53" pageNumber="54">1</paragraph>
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<paragraph id="725E36EC242BBD72FF37DF5C1EA3A257" blockId="53.[151,160,286,309]" lastBlockId="53.[199,1436,222,342]" pageId="53" pageNumber="54">
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- Rachillae 11.9(6.0–21.5) cm long;
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<collectingCountry id="0AF6767C242BBD72FD9FDE9C1996A3F4" box="[575,681,222,245]" name="Honduras" pageId="53" pageNumber="54">Honduras</collectingCountry>
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,
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<collectingCountry id="0AF6767C242BBD72FD15DE9C181AA3F4" box="[693,805,222,245]" name="Nicaragua" pageId="53" pageNumber="54">Nicaragua</collectingCountry>
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,
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<collectingCountry id="0AF6767C242BBD72FC91DE9C1899A3F4" box="[817,934,222,245]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
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(excluding Osa Peninsula and adjacent areas), and western and central
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<collectingCountry id="0AF6767C242BBD72FE6ADEBC1921A214" box="[458,542,254,277]" name="Panama" pageId="53" pageNumber="54">Panama</collectingCountry>
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............................................................................................................
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<taxonomicName id="B5E14D6F242BBD72FB59DEBC1A12A234" ID-CoL="5HFVK" authority="Rachillae" authorityName="Rachillae" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="subSpecies" species="congesta" subSpecies="congesta">
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subsp.
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<emphasis id="4095EAFE242BBD72FA9DDEBD1EA3A217" box="[1341,1436,255,278]" italics="true" pageId="53" pageNumber="54">congesta</emphasis>
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Rachillae
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</taxonomicName>
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19.1(15.0–23.0) cm long; Osa Peninsula and adjacent areas on the Pacific slope in
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<collectingCountry id="0AF6767C242BBD72FB06DF5C1E25A234" box="[1190,1306,286,309]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
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.................. .................................................................................................................................................................
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<taxonomicName id="B5E14D6F242BBD72FAA5DF7C1EA3A257" ID-CoL="5HFVL" authorityName="Henderson" authorityYear="2011" box="[1285,1436,318,342]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="subSpecies" species="congesta" subSpecies="osensis">
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subsp.
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<emphasis id="4095EAFE242BBD72FAEFDF7D1EA3A257" box="[1359,1436,319,342]" italics="true" pageId="53" pageNumber="54">osensis</emphasis>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |