150 lines
22 KiB
XML
150 lines
22 KiB
XML
<document id="494A6D6BF2DF01CEF3DA28C08363529F" ID-CLB-Dataset="4737" ID-DOI="10.11646/zootaxa.4500.3.4" ID-GBIF-Dataset="d44cea23-cf29-4e99-b971-7b36ea9ce911" ID-ISSN="1175-5326" ID-Zenodo-Dep="5297691" ID-ZooBank="CAC076FD-030D-4851-AAE9-44068F7B7A75" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630085217845" checkinUser="felipe" docAuthor="Grebennikov, Vasily V." docDate="2018" docId="AC2887EFFFAEFF86FF56FA73FBA69B61" docLanguage="en" docName="zootaxa.4500.3.4.pdf" docOrigin="Zootaxa 4500 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Devernodes chthonia Grebennikov 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="369" masterDocId="5011FF97FFA8FF80FFC1FF9DFFAF9C0F" masterDocTitle="Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia" masterLastPageNumber="380" masterPageNumber="363" pageNumber="369" updateTime="1699146272368" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5C30431F7BDB235239447F61A45CD4F0">Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia</mods:title>
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<mods:namePart id="5C9CE8BC4387E1D0986DB9F9A728B0B1">Grebennikov, Vasily V.</mods:namePart>
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<mods:title id="D46D52F30884147D8C377ADF98B29F4C">Zootaxa</mods:title>
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<mods:part id="D13E92A996192747C5A153960FE64363">
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<mods:date id="C6EFFF0D38B7A0236925A8B7506CAFAF">2018</mods:date>
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<mods:number id="DDAE844E4080918C2402BE75D488A4BA">2018-10-17</mods:number>
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<treatment id="AC2887EFFFAEFF86FF56FA73FBA69B61" ID-DOI="http://doi.org/10.5281/zenodo.5777921" ID-GBIF-Taxon="185119318" ID-Zenodo-Dep="5777921" LSID="urn:lsid:plazi:treatment:AC2887EFFFAEFF86FF56FA73FBA69B61" httpUri="http://treatment.plazi.org/id/AC2887EFFFAEFF86FF56FA73FBA69B61" lastPageNumber="369" pageId="6" pageNumber="369">
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<subSubSection id="6C9B6572FFAEFF86FF56FA73FE4E9A07" box="[151,481,1518,1544]" pageId="6" pageNumber="369" type="nomenclature">
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<paragraph id="243E36F9FFAEFF86FF56FA73FE4E9A07" blockId="6.[151,625,1518,1578]" box="[151,481,1518,1544]" pageId="6" pageNumber="369">
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<heading id="7F768195FFAEFF86FF56FA73FE4E9A07" bold="true" box="[151,481,1518,1544]" fontSize="11" level="1" pageId="6" pageNumber="369" reason="1">
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<taxonomicName id="E3814D7AFFAEFF86FF56FA73FE379A07" authority="Grebennikov, 2018" authorityName="Grebennikov" authorityYear="2018" box="[151,408,1518,1544]" class="Insecta" family="Curculionidae" genus="Devernodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="chthonia" status="sp. nov.">
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<emphasis id="16F5EAEBFFAEFF86FF56FA73FE379A07" bold="true" box="[151,408,1518,1544]" italics="true" pageId="6" pageNumber="369">Devernodes chthonia</emphasis>
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<taxonomicNameLabel id="0DC65790FFAEFF86FE5FFA73FE4E9A07" box="[414,481,1518,1544]" pageId="6" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="6C9B6572FFAEFF86FF56F98CFB5F9A9A" pageId="6" pageNumber="369" type="description">
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<paragraph id="243E36F9FFAEFF86FF56F98CFDDE9A25" blockId="6.[151,625,1518,1578]" box="[151,625,1553,1578]" pageId="6" pageNumber="369">
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<figureCitation id="BCBA2A7CFFAEFF86FF56F98CFEBD9A25" box="[151,274,1553,1578]" captionStart="FIGURE 4" captionStartId="8.[151,250,1365,1387]" captionTargetBox="[151,1426,195,1340]" captionTargetId="figure-144@8.[151,1436,195,1343]" captionTargetPageId="8" captionText="FIGURE 4. Devernodes chthonia sp. n., holotype. A–D: habitus, dorsal (A), lateral (B), ventral (C), fronto-lateral (D); E–G: aedeagus, lateral (E), dorsal (F), ventral (G)." figureDoi="http://doi.org/10.5281/zenodo.5297701" httpUri="https://zenodo.org/record/5297701/files/figure.png" pageId="6" pageNumber="369">Figs 4A–G</figureCitation>
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,
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<figureCitation id="BCBA2A7CFFAEFF86FEDCF98FFEFC9A26" box="[285,339,1554,1577]" captionStart="FIGURE 7" captionStartId="11.[151,250,1590,1612]" captionTargetBox="[151,1436,195,1566]" captionTargetId="figure-194@11.[151,1436,195,1566]" captionTargetPageId="11" captionText="FIGURE 7. Devernodes, clades A–F, heads dorsal (A–F) and lateral (G–L), holotypes (A, C–F) and paratype 2345 (B). A,B,G,H: D. alkippe sp. n., C, I: D. asteria sp. n., D, J: D. chthonia sp. n., E, K: D. drimo sp. n., F, L: D. methone sp. n." figureDoi="http://doi.org/10.5281/zenodo.5297709" httpUri="https://zenodo.org/record/5297709/files/figure.png" pageId="6" pageNumber="369">7D,J</figureCitation>
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,
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<figureCitation id="BCBA2A7CFFAEFF86FE9EF98CFE399A25" box="[351,406,1553,1578]" captionStart="FIGURE 8" captionStartId="12.[151,250,1271,1293]" captionTargetBox="[151,1436,189,1209]" captionTargetId="figure-356@12.[151,1436,189,1209]" captionTargetPageId="12" captionText="FIGURE 8. Devernodes, clades A–F, right antennae (A–F) and declivity of left elytron (G–L), holotypes (A, C–F) and paratype 2345 (B). A,B,G,H: D. alkippe sp. n., C, I: D. asteria sp. n., D, J: D. chthonia sp. n., E, K: D. drimo sp. n., F, L: D. methone sp. n. Numbering of elytral striae does not necessarily imply homology." figureDoi="http://doi.org/10.5281/zenodo.5297711" httpUri="https://zenodo.org/record/5297711/files/figure.png" pageId="6" pageNumber="369">8D,J</figureCitation>
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,
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<figureCitation id="BCBA2A7CFFAEFF86FE60F98CFDB69A25" box="[417,537,1553,1578]" captionStart="FIGURE 9" captionStartId="13.[151,250,1128,1150]" captionTargetBox="[151,1436,187,1106]" captionTargetId="figure-136@13.[151,1436,187,1106]" captionTargetPageId="13" captionText="FIGURE 9. Devernodes, details. A–B: D. chthonia sp. n., holotype, male, ventral (A: mouthparts, B: genitalia and apical sclerites); C: D. asteria sp. n., holotype, male sternites VIII and IX; D–G: D. chthonia sp. n., paratype 4368, female genitalia and apical sclerites (D, F: ventral, E: right dorso-lateral)." figureDoi="http://doi.org/10.5281/zenodo.5297713" httpUri="https://zenodo.org/record/5297713/files/figure.png" pageId="6" pageNumber="369">9A,B,D–G</figureCitation>
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,
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<figureCitation id="BCBA2A7CFFAEFF86FDE4F98CFDEC9A25" box="[549,579,1553,1578]" captionStart="FIGURE 10" captionStartId="13.[151,250,1898,1920]" captionTargetBox="[155,1433,1263,1873]" captionTargetId="figure-145@13.[151,1436,1261,1876]" captionTargetPageId="13" captionText="FIGURE 10. Neighbour-joining clustering of 26 DNA barcoded Devernodes type specimens performed using the online BOLD tree-building engine and Kimura 2-parameter. Branch length shown is proportional to relative divergence with scale bars indicating 0.02 units. Red dots denote illustrated specimens from each of six clusters; “HT” denotes holotypes; images are to scale. Unless otherwise indicated, all type specimens are stored in CNC." figureDoi="http://doi.org/10.5281/zenodo.5297717" httpUri="https://zenodo.org/record/5297717/files/figure.png" pageId="6" pageNumber="369">10</figureCitation>
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,
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<figureCitation id="BCBA2A7CFFAEFF86FD91F98CFDC19A25" box="[592,622,1553,1578]" captionStart="FIGURE 11" captionStartId="14.[151,250,1548,1570]" captionTargetBox="[151,1436,196,1523]" captionTargetId="figure-185@14.[151,1436,193,1526]" captionTargetPageId="14" captionText="FIGURE 11. Maximum Likelihood inference phylogram of Devernodes weevils from a three fragment dataset with 3223 aligned positions. Digits at internodes are bootstrap values of 60% and more. Branch length shown is proportional to relative divergence with scale bars indicating 0.2 units." figureDoi="http://doi.org/10.5281/zenodo.5297719" httpUri="https://zenodo.org/record/5297719/files/figure.png" pageId="6" pageNumber="369">11</figureCitation>
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.
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</paragraph>
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<paragraph id="243E36F9FFAEFF86FF56F9C4FB5F9A9A" blockId="6.[151,1437,1625,1902]" pageId="6" pageNumber="369">
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<emphasis id="16F5EAEBFFAEFF86FF56F9C4FE079A7D" bold="true" box="[151,424,1625,1650]" inLineHeading="true" pageId="6" pageNumber="369" reason="1">Diagnostic description.</emphasis>
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<typeStatus id="FB3A885BFFAEFF86FE6EF9C4FDB89A7D" box="[431,535,1625,1650]" pageId="6" pageNumber="369" type="holotype">Holotype</typeStatus>
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male (
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<figureCitation id="BCBA2A7CFFAEFF86FDA3F9C4FD709A7D" box="[610,735,1625,1650]" captionStart="FIGURE 4" captionStartId="8.[151,250,1365,1387]" captionTargetBox="[151,1426,195,1340]" captionTargetId="figure-144@8.[151,1436,195,1343]" captionTargetPageId="8" captionText="FIGURE 4. Devernodes chthonia sp. n., holotype. A–D: habitus, dorsal (A), lateral (B), ventral (C), fronto-lateral (D); E–G: aedeagus, lateral (E), dorsal (F), ventral (G)." figureDoi="http://doi.org/10.5281/zenodo.5297701" httpUri="https://zenodo.org/record/5297701/files/figure.png" pageId="6" pageNumber="369">Figs 4A–G</figureCitation>
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,
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||
<figureCitation id="BCBA2A7CFFAEFF86FD2BF9C7FC8F9A7E" box="[746,800,1626,1649]" captionStart="FIGURE 7" captionStartId="11.[151,250,1590,1612]" captionTargetBox="[151,1436,195,1566]" captionTargetId="figure-194@11.[151,1436,195,1566]" captionTargetPageId="11" captionText="FIGURE 7. Devernodes, clades A–F, heads dorsal (A–F) and lateral (G–L), holotypes (A, C–F) and paratype 2345 (B). A,B,G,H: D. alkippe sp. n., C, I: D. asteria sp. n., D, J: D. chthonia sp. n., E, K: D. drimo sp. n., F, L: D. methone sp. n." figureDoi="http://doi.org/10.5281/zenodo.5297709" httpUri="https://zenodo.org/record/5297709/files/figure.png" pageId="6" pageNumber="369">7D,J</figureCitation>
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,
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||
<figureCitation id="BCBA2A7CFFAEFF86FCEAF9C4FC049A7D" box="[811,939,1625,1650]" captionStart="FIGURE 8" captionStartId="12.[151,250,1271,1293]" captionTargetBox="[151,1436,189,1209]" captionTargetId="figure-356@12.[151,1436,189,1209]" captionTargetPageId="12" captionText="FIGURE 8. Devernodes, clades A–F, right antennae (A–F) and declivity of left elytron (G–L), holotypes (A, C–F) and paratype 2345 (B). A,B,G,H: D. alkippe sp. n., C, I: D. asteria sp. n., D, J: D. chthonia sp. n., E, K: D. drimo sp. n., F, L: D. methone sp. n. Numbering of elytral striae does not necessarily imply homology." figureDoi="http://doi.org/10.5281/zenodo.5297711" httpUri="https://zenodo.org/record/5297711/files/figure.png" pageId="6" pageNumber="369">8D,J, 8A,B</figureCitation>
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,
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<figureCitation id="BCBA2A7CFFAEFF86FC76F9C4FC7A9A7D" box="[951,981,1625,1650]" captionStart="FIGURE 10" captionStartId="13.[151,250,1898,1920]" captionTargetBox="[155,1433,1263,1873]" captionTargetId="figure-145@13.[151,1436,1261,1876]" captionTargetPageId="13" captionText="FIGURE 10. Neighbour-joining clustering of 26 DNA barcoded Devernodes type specimens performed using the online BOLD tree-building engine and Kimura 2-parameter. Branch length shown is proportional to relative divergence with scale bars indicating 0.02 units. Red dots denote illustrated specimens from each of six clusters; “HT” denotes holotypes; images are to scale. Unless otherwise indicated, all type specimens are stored in CNC." figureDoi="http://doi.org/10.5281/zenodo.5297717" httpUri="https://zenodo.org/record/5297717/files/figure.png" pageId="6" pageNumber="369">10</figureCitation>
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,
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<figureCitation id="BCBA2A7CFFAEFF86FC21F9C4FC529A7D" box="[992,1021,1625,1650]" captionStart="FIGURE 11" captionStartId="14.[151,250,1548,1570]" captionTargetBox="[151,1436,196,1523]" captionTargetId="figure-185@14.[151,1436,193,1526]" captionTargetPageId="14" captionText="FIGURE 11. Maximum Likelihood inference phylogram of Devernodes weevils from a three fragment dataset with 3223 aligned positions. Digits at internodes are bootstrap values of 60% and more. Branch length shown is proportional to relative divergence with scale bars indicating 0.2 units." figureDoi="http://doi.org/10.5281/zenodo.5297719" httpUri="https://zenodo.org/record/5297719/files/figure.png" pageId="6" pageNumber="369">11</figureCitation>
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), body length
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<quantity id="E3799B1CFFAEFF86FB5CF9C4FB5A9A7E" box="[1181,1269,1625,1650]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="369" unit="mm" value="3.1">3.1 mm</quantity>
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, DNA barcode GenBank accession number in
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<figureCitation id="BCBA2A7CFFAEFF86FE32F9E1FDE99A9A" box="[499,582,1660,1685]" captionStart="FIGURE 10" captionStartId="13.[151,250,1898,1920]" captionTargetBox="[155,1433,1263,1873]" captionTargetId="figure-145@13.[151,1436,1261,1876]" captionTargetPageId="13" captionText="FIGURE 10. Neighbour-joining clustering of 26 DNA barcoded Devernodes type specimens performed using the online BOLD tree-building engine and Kimura 2-parameter. Branch length shown is proportional to relative divergence with scale bars indicating 0.02 units. Red dots denote illustrated specimens from each of six clusters; “HT” denotes holotypes; images are to scale. Unless otherwise indicated, all type specimens are stored in CNC." figureDoi="http://doi.org/10.5281/zenodo.5297717" httpUri="https://zenodo.org/record/5297717/files/figure.png" pageId="6" pageNumber="369">Fig. 10</figureCitation>
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; unique combination of morphological characters in
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<tableCitation id="69030342FFAEFF86FB57F9E1FB459A9A" box="[1174,1258,1660,1685]" captionStart="TABLE 3" captionStartId="15.[151,239,1210,1233]" captionText="TABLE 3. Discrete characters for morphological diagnostics of Devernodes weevils" pageId="6" pageNumber="369">Table 3</tableCitation>
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.
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</paragraph>
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<emphasis id="16F5EAEBFFAEFF86FF06F93CFE1B9AB5" bold="true" box="[199,436,1697,1722]" inLineHeading="true" pageId="6" pageNumber="369" reason="1">Material examined.</emphasis>
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<materialsCitation id="94E93CA4FFAEFF86FE7EF93CFD7F9AD1" ID-GBIF-Occurrence="3339719307" accessionNumber="E10538" collectingDate="2012-05-12" collectionCode="CNC, V" collectorName="Tam Dao, N" country="Vietnam" location="VIETNAM" pageId="6" pageNumber="369" specimenCount="1" typeStatus="holotype">
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<typeStatus id="FB3A885BFFAEFF86FE7EF93CFD889AB5" box="[447,551,1697,1722]" pageId="6" pageNumber="369" type="holotype">Holotype</typeStatus>
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(
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): “
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<collectingCountry id="5C967669FFAEFF86FD5AF93CFC8E9AB5" box="[667,801,1697,1722]" name="Vietnam" pageId="6" pageNumber="369">VIETNAM</collectingCountry>
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,
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<collectorName id="8974532FFFAEFF86FCF1F93FFC6C9AB5" box="[816,963,1697,1722]" pageId="6" pageNumber="369">Tam Dao, N</collectorName>
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2127'54" E10538'44",
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<date id="503F1039FFAEFF86FB06F93CFA979AB5" box="[1223,1336,1697,1722]" pageId="6" pageNumber="369" value="2012-05-12">
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<collectingDate id="407BE9D1FFAEFF86FB06F93CFA979AB5" box="[1223,1336,1697,1722]" pageId="6" pageNumber="369" value="2012-05-12">12.v.2012</collectingDate>
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</date>
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, 1218m, sift02,
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<collectionCode id="4290AE3CFFAEFF86FF27F958FF559AD1" box="[230,250,1733,1758]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="369" type="Museum">V</collectionCode>
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.Grebennikov”, “CNCCOLVG00004339”
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</materialsCitation>
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.
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<materialsCitation id="94E93CA4FFAEFF86FD1DF958FF529B0D" ID-GBIF-Occurrence="3339719302" accessionNumber="E10538" collectingDate="2012-05-12" collectionCode="CNC" collectorName="Tam Dao, N" country="Vietnam" location="VIETNAM" pageId="6" pageNumber="369" specimenCount="6" typeStatus="paratype">
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<typeStatus id="FB3A885BFFAEFF86FD1DF958FCE69AD2" box="[732,841,1733,1757]" pageId="6" pageNumber="369" type="paratype">Paratypes</typeStatus>
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(
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<collectionCode id="4290AE3CFFAEFF86FC98F958FC369AD1" box="[857,921,1733,1758]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="6" pageNumber="369">CNC</collectionCode>
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):
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, as in
|
||
<figureCitation id="BCBA2A7CFFAEFF86FB47F958FB769AD1" box="[1158,1241,1733,1758]" captionStart="FIGURE 10" captionStartId="13.[151,250,1898,1920]" captionTargetBox="[155,1433,1263,1873]" captionTargetId="figure-145@13.[151,1436,1261,1876]" captionTargetPageId="13" captionText="FIGURE 10. Neighbour-joining clustering of 26 DNA barcoded Devernodes type specimens performed using the online BOLD tree-building engine and Kimura 2-parameter. Branch length shown is proportional to relative divergence with scale bars indicating 0.02 units. Red dots denote illustrated specimens from each of six clusters; “HT” denotes holotypes; images are to scale. Unless otherwise indicated, all type specimens are stored in CNC." figureDoi="http://doi.org/10.5281/zenodo.5297717" httpUri="https://zenodo.org/record/5297717/files/figure.png" pageId="6" pageNumber="369">Fig. 10</figureCitation>
|
||
, same locality as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6C9B6572FFAEFF86FF06F890FE6E9B46" pageId="6" pageNumber="369" type="distribution">
|
||
<paragraph id="243E36F9FFAEFF86FF06F890FE6E9B46" blockId="6.[151,1437,1625,1902]" pageId="6" pageNumber="369">
|
||
<emphasis id="16F5EAEBFFAEFF86FF06F890FECF9B29" bold="true" box="[199,352,1805,1830]" inLineHeading="true" pageId="6" pageNumber="369" reason="1">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="FB3A885BFFAEFF86FDB5F890FD0B9B2A" box="[628,676,1805,1829]" pageId="6" pageNumber="369">type</typeStatus>
|
||
locality, the Tam Dao Mountains in northern
|
||
<collectingCountry id="5C967669FFAEFF86FB04F890FA8B9B29" box="[1221,1316,1805,1830]" name="Vietnam" pageId="6" pageNumber="369">Vietnam</collectingCountry>
|
||
(
|
||
<figureCitation id="BCBA2A7CFFAEFF86FAF6F890FA209B29" box="[1335,1423,1805,1830]" captionStart="FIGURE 12" captionStartId="15.[151,250,1052,1074]" captionTargetBox="[242,1361,205,1024]" captionTargetId="figure-399@15.[229,1372,195,1032]" captionTargetPageId="15" captionText="FIGURE 12. Geographical distribution of recent Lymantini. Northernmost limits in North America follow Anderson (2016); southernmost limits are based on country records (Alonso-Zarazaga & Lyal 1999) and are fuzzy to reflect uncertainty. New country records (Peru, Guyana, Surinam) are based on unpublished records of Ithaura (Robert Anderson, personal communication). Map design: Jessica Hsiung." figureDoi="http://doi.org/10.5281/zenodo.5297721" httpUri="https://zenodo.org/record/5297721/files/figure.png" pageId="6" pageNumber="369">Fig. 12</figureCitation>
|
||
). Elevation:
|
||
<quantity id="E3799B1CFFAEFF86FED0F8ACFE129B46" box="[273,445,1841,1866]" metricMagnitude="3" metricUnit="m" metricValue="1.1785" metricValueMax="1.229" metricValueMin="1.128" pageId="6" pageNumber="369" unit="m" value="1178.5" valueMax="1229.0" valueMin="1128.0">
|
||
<geoCoordinate id="41B5503EFFAEFF86FED0F8ACFEE09B45" box="[273,335,1841,1866]" degrees="1,128" direction="north" orientation="latitude" pageId="6" pageNumber="369" precision="55" value="1.128">1,128</geoCoordinate>
|
||
<geoCoordinate id="41B5503EFFAEFF86FE8EF8ACFE329B45" box="[335,413,1841,1866]" degrees="1,229" direction="west" orientation="longitude" pageId="6" pageNumber="369" precision="55" value="-1.229">–1,229</geoCoordinate>
|
||
m
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6C9B6572FFAEFF86FF06F8C8FBA69B61" box="[199,1033,1877,1902]" pageId="6" pageNumber="369" type="etymology">
|
||
<paragraph id="243E36F9FFAEFF86FF06F8C8FBA69B61" blockId="6.[151,1437,1625,1902]" box="[199,1033,1877,1902]" pageId="6" pageNumber="369">
|
||
<emphasis id="16F5EAEBFFAEFF86FF06F8C8FEE19B61" bold="true" box="[199,334,1877,1902]" inLineHeading="true" pageId="6" pageNumber="369" reason="1">Etymology.</emphasis>
|
||
One of the seven daughters of Alcyoneus; noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |