treatments-xml/data/44/45/4C/44454C2AFFD4CA792F6F2357250C0995.xml
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<document id="CCAB41CFF2F69B5C3425ABE5FE4C47AB" ID-DOI="10.11646/zootaxa.1011.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5048990" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105215244" checkinUser="felipe" docAuthor="Jaschhof, Mathias &amp; Hippa, Heikki" docDate="2005" docId="44454C2AFFD4CA792F6F2357250C0995" docLanguage="en" docName="zt01011p054.pdf" docOrigin="Zootaxa 1011 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1011.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Manota squamulata Jaschhof &amp; Hippa 2005, sp. n." docType="treatment" docVersion="3" lastPageNumber="18" masterDocId="B87C3452FFDBCA682E67255B20100B0B" masterDocTitle="The genus Manota in Costa Rica (Diptera: Mycetophilidae)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="16" updateTime="1698980072965" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="5D8857F055BADF2A72D18FE44F4E4D44" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="82309CD287A7FB8A784377D0828064F5">
<mods:title id="CCD5C219D2A23EBEDD2BC1DBBC2542DA">The genus Manota in Costa Rica (Diptera: Mycetophilidae)</mods:title>
</mods:titleInfo>
<mods:name id="FDC50AC5B68B556DFDD4BE26CCEA3540" type="personal">
<mods:role id="8752FC842AD6805D8AECEC5491D52CEF">
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<mods:namePart id="1B74B383A9E5628438A46B5AAF4E5E2E">Jaschhof, Mathias</mods:namePart>
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<mods:namePart id="C2021F5E57E3540175DE074F0FBA8C31">Hippa, Heikki</mods:namePart>
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<mods:title id="1806C031A0748DCE75DB3F1FD5021A9C">Zootaxa</mods:title>
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<mods:part id="337BE3312C303CC599DF9052C91F15C9">
<mods:date id="7661E472C98F6E8EFCE203D090CE55BD">2005</mods:date>
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<mods:number id="AF368F5D89D44F96E1E3C4BB3F8FCCD3">2005-06-24</mods:number>
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<mods:number id="8B52E4FE8A8DAD66160E878119E03070">1011</mods:number>
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<mods:start id="48FD555FA3CA360362AB74443D3388B1">1</mods:start>
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<mods:url id="8E0118B7B3D8C76C582D944287B3A349">https://biotaxa.org/Zootaxa/article/view/zootaxa.1011.1.1</mods:url>
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<mods:classification id="4F24157B1FA1E8351ABA509CBF22C43B">journal article</mods:classification>
<mods:identifier id="D6807DFCDA29472CA00DB98E4763B9B5" type="DOI">10.11646/zootaxa.1011.1.1</mods:identifier>
<mods:identifier id="3FDE952022A2543E80272230D05739DA" type="ISSN">1175­5334</mods:identifier>
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<treatment id="44454C2AFFD4CA792F6F2357250C0995" ID-DOI="http://doi.org/10.5281/zenodo.5049036" ID-Zenodo-Dep="5049036" LSID="urn:lsid:plazi:treatment:44454C2AFFD4CA792F6F2357250C0995" httpUri="http://treatment.plazi.org/id/44454C2AFFD4CA792F6F2357250C0995" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="84F6AEB7FFD4CA672F6F235722B50D2D" box="[264,677,1548,1574]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="CC53FD3CFFD4CA672F6F235722B50D2D" blockId="15.[264,677,1548,1574]" box="[264,677,1548,1574]" pageId="15" pageNumber="16">
<heading id="971B4A50FFD4CA672F6F235722B50D2D" bold="true" box="[264,677,1548,1574]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="FE98212EFFD4CA672F6F235722B50D2D" bold="true" box="[264,677,1548,1574]" pageId="15" pageNumber="16">
<taxonomicName id="0BEC86BFFFD4CA672F6F235721EA0D2D" ID-CoL="3XVYQ" authority="Jaschhof &amp; Hippa, 2005" authorityName="Jaschhof &amp; Hippa" authorityYear="2005" box="[264,506,1548,1574]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="squamulata" status="sp. nov.">
<emphasis id="FE98212EFFD4CA672F6F235721EA0D2D" bold="true" box="[264,506,1548,1574]" italics="true" pageId="15" pageNumber="16">Manota squamulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AB9C55FFD4CA672C65235722540D2D" box="[514,580,1548,1574]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="54D7E1B9FFD4CA672C332357228C0D2D" box="[596,668,1548,1574]" captionStart="FIGURE 7" captionStartId="16.[264,368,1652,1676]" captionTargetBox="[268,1316,773,1616]" captionTargetId="figure-233@16.[264,1323,749,1659]" captionTargetPageId="16" captionText="FIGURE 7. Manota squamulata sp. n. A: Male terminalia, ventral view (holotype). B: Male terminalia, dorsal view (paratype from Cerro Puma). Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049012" httpUri="https://zenodo.org/record/5049012/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="84F6AEB7FFD4CA792F6F230C250C0995" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="CC53FD3CFFD4CA672F6F230C225A0D7A" blockId="15.[264,1324,1623,1889]" box="[264,586,1623,1649]" pageId="15" pageNumber="16">
<emphasis id="FE98212EFFD4CA672F6F230C21820D7B" box="[264,402,1623,1648]" italics="true" pageId="15" pageNumber="16">Description</emphasis>
. Small species.
</paragraph>
<paragraph id="CC53FD3CFFD4CA672F5F232422010DE2" blockId="15.[264,1324,1623,1889]" pageId="15" pageNumber="16">
Male.
<emphasis id="FE98212EFFD4CA672FE2232521DF0D93" bold="true" box="[389,463,1662,1688]" pageId="15" pageNumber="16">Head.</emphasis>
Antenna with fourth flagellomere 1.8 times as long as wide. Maxillary palpus with third palpomere bearing (3)4(5) curved sensilla; fourth palpomere with a small process apically.
</paragraph>
<paragraph id="CC53FD3CFFD4CA672F5F23AD22C90C6A" blockId="15.[264,1324,1623,1889]" pageId="15" pageNumber="16">
<emphasis id="FE98212EFFD4CA672F5F23AD218C0C1B" bold="true" box="[312,412,1782,1808]" pageId="15" pageNumber="16">Thorax.</emphasis>
Anterior basalare and laterotergite each setose. Episternum 3 with ventral portion setose.
<emphasis id="FE98212EFFD4CA672FDD224522150C33" bold="true" box="[442,517,1822,1848]" pageId="15" pageNumber="16">Wing.</emphasis>
Length
<quantity id="0B1450D9FFD4CA672C01224422D30C32" box="[614,707,1823,1849]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="mm" value="1.5">1.5 mm</quantity>
. Membrane with a few setae along posterior margin. Sc bare. CuA2 fading away proximally.
</paragraph>
<paragraph id="CC53FD3CFFCBCA782F5F2440251C09CD" blockId="16.[264,1323,283,710]" pageId="16" pageNumber="17">
<emphasis id="FE98212EFFCBCA782F5F244021D10A3E" bold="true" box="[312,449,283,309]" pageId="16" pageNumber="17">Terminalia</emphasis>
(
<figureCitation id="54D7E1B9FFCBCA782FB42447220C0A3D" box="[467,540,284,310]" captionStart="FIGURE 7" captionStartId="16.[264,368,1652,1676]" captionTargetBox="[268,1316,773,1616]" captionTargetId="figure-233@16.[264,1323,749,1659]" captionTargetPageId="16" captionText="FIGURE 7. Manota squamulata sp. n. A: Male terminalia, ventral view (holotype). B: Male terminalia, dorsal view (paratype from Cerro Puma). Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049012" httpUri="https://zenodo.org/record/5049012/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
). Sternite 9 largely merged with gonocoxites; only posterior margin distinct, situated at almost same level as posterior gonocoxal margin, straight or very slightly emarginate; on interior surface with extensive scale­like pattern; on exterior surface densely covered with setae, sometimes with a small non­setose area medially. Distolateral lobes of gonocoxites absent. Gonocoxites in position I+II with 1214 thick, blunt and pointed setae on lobe; in position III+IV with 1 pointed megaseta. Gonostyli short, slightly bilobed; ventral lobe on inner margin with some 5 large setae increasing in length posteriorly; dorsal lobe on posterior margin with some 4 large setae curved dorsally. Parastylar lobes large, rounded; each with some 5 setae posteriorly. Tegmen strongly narrowed in distal half, very wide basally with dorsal portions of tegmen extending laterally beyond width of parameral apodemes. Hypoproct tapering to tip; with long setae disto­laterally.
</paragraph>
<caption id="9893ADB4FFCBCA782F6F232F23A60DA5" ID-DOI="http://doi.org/10.5281/zenodo.5049012" ID-Zenodo-Dep="5049012" httpUri="https://zenodo.org/record/5049012/files/figure.png" pageId="16" pageNumber="17" startId="16.[264,368,1652,1676]" targetBox="[268,1316,773,1616]" targetPageId="16">
<paragraph id="CC53FD3CFFCBCA782F6F232F23A60DA5" blockId="16.[264,1322,1652,1710]" pageId="16" pageNumber="17">
<emphasis id="FE98212EFFCBCA782F6F232F219A0D87" bold="true" box="[264,394,1652,1676]" pageId="16" pageNumber="17">FIGURE 7.</emphasis>
<taxonomicName id="0BEC86BFFFCBCA782FF6232F22760D80" authority="Jaschhof &amp; Hippa, 2005" authorityName="Jaschhof &amp; Hippa" authorityYear="2005" box="[401,614,1652,1675]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="squamulata" status="sp. nov.">
<emphasis id="FE98212EFFCBCA782FF6232F22760D80" box="[401,614,1652,1675]" italics="true" pageId="16" pageNumber="17">Manota squamulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AB9C55FFCBCA782C0B232F22B50D80" box="[620,677,1652,1675]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="FE98212EFFCBCA782CCB232F22D80D87" bold="true" box="[684,712,1652,1676]" pageId="16" pageNumber="17">A:</emphasis>
Male terminalia, ventral view (holotype).
<emphasis id="FE98212EFFCBCA782AEC232F24B60D87" bold="true" box="[1163,1190,1652,1676]" pageId="16" pageNumber="17">B:</emphasis>
Male terminalia, dorsal view (paratype from Cerro Puma). Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="CC53FD3CFFCBCA792F5F2259236F0A55" blockId="16.[264,1323,1794,1900]" lastBlockId="17.[264,1323,284,670]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="FE98212EFFCBCA782F5F2259222D0C10" box="[312,573,1794,1819]" italics="true" pageId="16" pageNumber="17">Diagnostic characters</emphasis>
. Together with
<taxonomicName id="0BEC86BFFFCBCA782C9E225923760C10" authorityName="Jaschhof &amp; Hippa" authorityYear="2005" box="[761,870,1794,1819]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="major">
<emphasis id="FE98212EFFCBCA782C9E225923760C10" box="[761,870,1794,1819]" italics="true" pageId="16" pageNumber="17">M. major</emphasis>
</taxonomicName>
(next species), this is the only species with both the anterior basalare and laterotergites setose. The terminalia with sternite 9 are largely merged with the gonocoxites and with a scale­like pattern on the interior surface; disto­lateral lobes of the gonocoxites are absent; gonostyli are stout and with large setae apically; and the parastylar lobes are large and setose.
</paragraph>
<paragraph id="CC53FD3CFFCACA792F5F243722560AA5" blockId="17.[264,1323,284,670]" pageId="17" pageNumber="18">
<emphasis id="FE98212EFFCACA792F5F243721A30A8E" box="[312,435,364,389]" italics="true" pageId="17" pageNumber="18">Etymology</emphasis>
. From Latin,
<emphasis id="FE98212EFFCACA792C3B243722E00A8E" box="[604,752,364,389]" italics="true" pageId="17" pageNumber="18">squamulatus</emphasis>
, with small scales, referring to the interior surface structure on sternite 9.
</paragraph>
<paragraph id="CC53FD3CFFCACA792F5F24E7250C0995" blockId="17.[264,1323,284,670]" pageId="17" pageNumber="18">
<emphasis id="FE98212EFFCACA792F5F24E7216A0ADE" box="[312,378,444,469]" italics="true" pageId="17" pageNumber="18">Types</emphasis>
.
<materialsCitation id="7C84F761FFCACA792FEF24E722D6092D" collectingDate="2003-09-18" collectingDateMax="2003-10-07" collectingDateMin="2003-09-18" collectorName="M. Jaschhof" country="Costa Rica" county="Male" elevation="325" location="Sendero Espavel" municipality="Hitoy Cerere Biological Reserve" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
<emphasis id="FE98212EFFCACA792FEF24E721E30ADE" box="[392,499,444,469]" italics="true" pageId="17" pageNumber="18">
<typeStatus id="1357439EFFCACA792FEF24E721E30ADE" box="[392,499,444,469]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCounty id="253285B0FFCACA792C6524E722510ADD" box="[514,577,444,470]" pageId="17" pageNumber="18">Male</collectingCounty>
,
<collectingCountry id="B4FBBDACFFCACA792C2A24E722DF0ADD" box="[589,719,444,470]" name="Costa Rica" pageId="17" pageNumber="18">Costa Rica</collectingCountry>
,
<collectingRegion id="0E2833DEFFCACA792CBC24E723890ADD" box="[731,921,444,470]" country="Costa Rica" name="Limon" pageId="17" pageNumber="18">Limón province</collectingRegion>
,
<collectingMunicipality id="2C376746FFCACA792DC224E725360ADD" box="[933,1318,444,470]" pageId="17" pageNumber="18">Hitoy Cerere Biological Reserve</collectingMunicipality>
,
<location id="C933ABE7FFCACA792F6F24BF21C60AF5" LSID="urn:lsid:plazi:treatment:44454C2AFFD4CA792F6F2357250C0995:C933ABE7FFCACA792F6F24BF21C60AF5" box="[264,470,484,510]" country="Costa Rica" county="Male" municipality="Hitoy Cerere Biological Reserve" name="Sendero Espavel" pageId="17" pageNumber="18" stateProvince="Limon">Sendero Espavel</location>
,
<quantity id="0B1450D9FFCACA792F8F24BF22650AF5" box="[488,629,484,510]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="5.5" metricValueMin="1.0" pageId="17" pageNumber="18" unit="m" value="325.0" valueMax="550.0" valueMin="100.0">
<elevation id="47C11A0FFFCACA792F8F24BF22650AF5" box="[488,629,484,510]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="5.5" metricValueMin="1.0" pageId="17" pageNumber="18" unit="m" value="325.0" valueMax="550.0" valueMin="100.0">100550 m</elevation>
</quantity>
, in tall secondary rain­forest,
<date id="B852DBFCFFCACA792D9324BF24EA0AF5" box="[1012,1274,484,510]" pageId="17" pageNumber="18" value="2003-09-18" valueMax="2003-10-07" valueMin="2003-09-18">
<collectingDate id="A8162214FFCACA792D9324BF24EA0AF5" box="[1012,1274,484,510]" pageId="17" pageNumber="18" value="2003-09-18" valueMax="2003-10-07" valueMin="2003-09-18">18 Sep.7 Oct. 2003</collectingDate>
</date>
, by sweepnet and aspirator,
<collectorName id="611998EAFFCACA792C48275722D6092D" box="[559,710,524,550]" pageId="17" pageNumber="18">M. Jaschhof</collectorName>
</materialsCitation>
.
<materialsCitation id="7C84F761FFCACA792CB02757251B092D" box="[727,1291,524,550]" collectingDate="2003-09-18" collectingDateMax="2003-10-07" collectingDateMin="2003-09-18" collectorName="M. Jaschhof" country="Costa Rica" county="Male" elevation="325" location="Sendero Espavel" municipality="Hitoy Cerere Biological Reserve" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<emphasis id="FE98212EFFCACA792CB02757235E092E" box="[727,846,524,549]" italics="true" pageId="17" pageNumber="18">
<typeStatus id="1357439EFFCACA792CB02757235E092E" box="[727,846,524,549]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCount id="DAEA36B5FFCACA792D06275723A8092D" box="[865,952,524,550]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
, same data as the holotype
</materialsCitation>
;
<materialsCitation id="7C84F761FFCACA792B7C275721C60945" collectingDate="2003-09-18" collectingDateMax="2003-10-07" collectingDateMin="2003-09-18" collectorName="M. Jaschhof" country="Costa Rica" county="Male" elevation="325" location="Sendero Espavel" municipality="Hitoy Cerere Biological Reserve" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" stateProvince="Limon" typeStatus="paratype">
<specimenCount id="DAEA36B5FFCACA792B7C2757215F0945" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
, same data
</materialsCitation>
, but
<quantity id="0B1450D9FFCACA792C76276F22700945" box="[529,608,564,590]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="18" unit="m" value="550.0">550 m</quantity>
, by Malaise trap, E. Rojas, B. Gamboa, W. Arana, M. &amp; C. Jaschhof;
<materialsCitation id="7C84F761FFCACA792F19270725080995" collectingDate="2003-06-19" collectingDateMax="2003-07-08" collectingDateMin="2003-06-19" collectionCode="INBio" collectorName="M. Moraga &amp; A. Azofeifa &amp; Caballero" country="Costa Rica" elevation="200" location="Cerro Puma" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="DAEA36B5FFCACA792F19270721C0097D" box="[382,464,604,630]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
,
<collectingRegion id="0E2833DEFFCACA792FBA270722DE097D" box="[477,718,604,630]" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="18">Puntarenas province</collectingRegion>
,
<location id="C933ABE7FFCACA792CBD2707237A097D" LSID="urn:lsid:plazi:treatment:44454C2AFFD4CA792F6F2357250C0995:C933ABE7FFCACA792CBD2707237A097D" box="[730,874,604,630]" country="Costa Rica" name="Cerro Puma" pageId="17" pageNumber="18" stateProvince="Puntarenas">Cerro Puma</location>
,
<quantity id="0B1450D9FFCACA792D10270723EE097D" box="[887,1022,604,630]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="17" pageNumber="18" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">
<elevation id="47C11A0FFFCACA792D10270723EE097D" box="[887,1022,604,630]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="17" pageNumber="18" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">100300 m</elevation>
</quantity>
,
<date id="B852DBFCFFCACA792A6C27072510097D" box="[1035,1280,604,630]" pageId="17" pageNumber="18" value="2003-06-19" valueMax="2003-07-08" valueMin="2003-06-19">
<collectingDate id="A8162214FFCACA792A6C27072510097D" box="[1035,1280,604,630]" pageId="17" pageNumber="18" value="2003-06-19" valueMax="2003-07-08" valueMin="2003-06-19">19 June8 July 2003</collectingDate>
</date>
,
<collectingMethod id="15AD852BFFCACA792B6A270722020995" pageId="17" pageNumber="18">by flight interception trap</collectingMethod>
,
<collectorName id="611998EAFFCACA792C7927DF22B50995" box="[542,677,644,670]" pageId="17" pageNumber="18">M. Moraga</collectorName>
,
<collectorName id="611998EAFFCACA792CD627DF232B0995" box="[689,827,644,670]" pageId="17" pageNumber="18">A. Azofeifa</collectorName>
&amp; K.
<collectorName id="611998EAFFCACA792DE327DF23E60995" box="[900,1014,644,670]" pageId="17" pageNumber="18">Caballero</collectorName>
(
<collectionCode id="AAFD65F9FFCACA792A6127DF24410995" box="[1030,1105,644,670]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
sample #74480)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>