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<document id="BF78745E298113FA2736393DA2A64502" ID-DOI="10.3897/asp.82.e112587" ID-ZooBank="CB81B265-46A0-4247-A2F2-C9AC2394BC4E" ID-publisher-id="112587" URI-arpha="E54D30EB-4688-54F3-9F8B-C2351EFB7165" URI-zoobank="http://zoobank.org/CB81B265-46A0-4247-A2F2-C9AC2394BC4E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1716297392854" checkinUser="pensoft" docAuthor="Vilarino, Albane &amp; Calor, Adolfo R." docDate="2024" docId="B1E6960C7A1359EE8013F4386196B7B8" docLanguage="en" docName="Arthropod Systematics &amp; Phylogeny 82: 407-431" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 82" docSource="https://arthropod-systematics.arphahub.com/article/112587/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Xiphocentron (Sphagocentron) guanacaste Vilarino &amp; Calor 2024, sp. nov." docType="treatment" docUuid="229026A6-84A4-4D23-BDA9-52074A7DF801" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="431" masterDocId="E54D30EB468854F39F8BC2351EFB7165" masterDocTitle="Comparative geometric morphometrics of male genitalia in Xiphocentron subgenera (Trichoptera: Xiphocentronidae): new species, revision and phylogenetic systematics of the subgenus Sphagocentron" masterLastPageNumber="431" masterPageNumber="407" pageNumber="407" updateTime="1716324065396" updateUser="admin">
<mods:mods id="63A9F57EA564655C95F8D20BE8F890A2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9AC8F65714B06AF0567DC880760705AF">
<mods:title id="B59B9C9D7939CD4D8EAF6FE964E4AF0C">Comparative geometric morphometrics of male genitalia in Xiphocentron subgenera (Trichoptera: Xiphocentronidae): new species, revision and phylogenetic systematics of the subgenus Sphagocentron</mods:title>
</mods:titleInfo>
<mods:name id="4B53CB696F4C1B8961CC3E1A8DF6E6A1" type="personal">
<mods:role id="638A3736E1F515EE0C7146E8A854BDBF">
<mods:roleTerm id="25CE616B44037567FE1603CE86B78F22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6B2854D5D89C2068B0351EA9613FDAE">Vilarino, Albane</mods:namePart>
<mods:nameIdentifier id="6A18C5BF868C9BD95F0B49BF083E8417" type="ORCID">0000-0003-3912-8928</mods:nameIdentifier>
<mods:affiliation id="BD9AB562C4E64D7ED88BC29846383226">Universidade Federal da Bahia, Instituto de Biologia, Laboratório de Entomologia Aquática, Salvador, Bahia, Brazil</mods:affiliation>
</mods:name>
<mods:name id="C13E251348853C231A89097B5EE7745C" type="personal">
<mods:role id="CFD7FEC62E95CCEAF3EF91F52870CAFE">
<mods:roleTerm id="FBD2D544493AA3E5B1342D45B37F25F7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="803ABB51E6E6B50D8CF3A29B81133302">Calor, Adolfo R.</mods:namePart>
<mods:nameIdentifier id="593CE72B39797C02BB4C597FA4B14C08" type="ORCID">0000-0003-3975-3176</mods:nameIdentifier>
<mods:affiliation id="B27BA1830AABD365FB0125959E13DBAE">Universidade Federal da Bahia, Instituto de Biologia, Laboratório de Entomologia Aquática, Salvador, Bahia, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7CF6014BE07951142EB2293D3EB2FE7E">text</mods:typeOfResource>
<mods:relatedItem id="49530004918DC0BABFAC32E460BCDF21" type="host">
<mods:titleInfo id="76C884658EDD52557324EEF38E55CC94">
<mods:title id="E20A1B87D4F856B4E9E3C6A6F947515D">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part id="82E93AEE8CC2F01A39AA4A544C63A311">
<mods:date id="D27E1017E090F171EEDB93DEF14899C7">2024</mods:date>
<mods:detail id="6725146C6FE642CC4C36244B5E52FF44" type="pubDate">
<mods:number id="EE97D6C019488620CDA85DE0E1E46DEB">2024-05-21</mods:number>
</mods:detail>
<mods:detail id="91E7ADB286E36366CA7FE21A294A27D2" type="volume">
<mods:number id="DA3450A107FFD7A19290C96A72E84684">82</mods:number>
</mods:detail>
<mods:extent id="6046A705C6791E728909E0A0F480643E" unit="page">
<mods:start id="EC88786354BC52298F0FD6527FAB67C9">407</mods:start>
<mods:end id="2FF40925E5F5AA95876288BED7E56C34">431</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E9F01B113F7B56EBDB9072E67EC4D600">journal article</mods:classification>
<mods:identifier id="22363A3397A44B94B2968DEAF42987A0" type="DOI">10.3897/asp.82.e112587</mods:identifier>
<mods:identifier id="193BB956238700BDF7D7D9E7899CCF4D" type="ZooBank">CB81B265-46A0-4247-A2F2-C9AC2394BC4E</mods:identifier>
</mods:mods>
<treatment id="B1E6960C7A1359EE8013F4386196B7B8" ID-DOI="http://doi.org/10.5281/zenodo.11235534" ID-Zenodo-Dep="11235534" ID-arpha="B1E6960C-7A13-59EE-8013-F4386196B7B8" ID-zoobank="https://zoobank.org/229026A6-84A4-4D23-BDA9-52074A7DF801" LSID="urn:lsid:zoobank.org:act:229026A6-84A4-4D23-BDA9-52074A7DF801" LSID-ZBK="urn:lsid:zoobank.org:act:229026A6-84A4-4D23-BDA9-52074A7DF801" httpUri="http://treatment.plazi.org/id/B1E6960C7A1359EE8013F4386196B7B8">
<subSubSection id="DFA792D32E0DB9A57B7520A1C4C7CA5B" type="nomenclature">
<paragraph id="4D31C4702200DA09B66F615DF8C67973">
<taxonomicName id="0664C9B6C926D5AAC4744D51BCAE58EC" authority="Vilarino &amp; Calor, 2024" authorityName="Vilarino &amp; Calor" authorityYear="2024" class="Insecta" family="Xiphocentronidae" genus="Xiphocentron" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="guanacaste" status="sp. nov." subGenus="Sphagocentron">
<emphasis id="FF7199A0C8862C993AED906082CAD842" italics="true">Xiphocentron</emphasis>
(
<emphasis id="5B40D85137D7CABFF236C7551D183F59" italics="true">Sphagocentron</emphasis>
)
<emphasis id="2A764FAD53922B6163564BB6C7ECF459" italics="true">guanacaste</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2E06DDBCEE524CE775A8EBB2CE800DF4" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="200959B64DA5EAD66C4F1161C891CCEC" type="description">
<paragraph id="DA6457E9DE11719C12D20D6339F9750C">
<figureCitation id="42C54F4A330B8A474B5320B47AAE3537" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">Figures 9 A – D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAVAI" type="type material">
<paragraph id="4E2D79ADAF9B9F4C7F21DD969F8F6E8A">
<heading id="B0579D9F06D856A7491CB4612894A1A2" reason="title">Type material.</heading>
</paragraph>
<paragraph id="3E87E3A7F8370CC4484AF338A7CAE681">
<materialsCitation id="BD015278B2B56EC8CD1A91A27672F2BD" collectingDate="2013-08-26" collectingMethod="Malaise Trap" collectionCode="BIOUG" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Area de Conservacion Guanacaste" elevation="575" latitude="10.88" location="Estacion San Gerardo" longLatPrecision="1" longitude="-85.389" municipality="Sector San Cristobal" specimenCode="BIOUG 28044" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="3E305D86D7EB63D499465E03EDD53512" bold="true">
<typeStatus id="C379E1097BA1185731CF30A9E801A0A4" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="DE8723DB2FB6DDE6DB341353DB412012" name="Costa Rica">COSTA RICA</collectingCountry>
•
<specimenCount id="483145E56BE64313F3C655B49CF86195" count="1" type="male">♂</specimenCount>
;
<collectingRegion id="9E5D1C6E1978FCE2C9C61E7D65B5D5EC" country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
,
<collectingCounty id="9B5D2D595BE8D9EFCB9C7FF8F0285EE9">Area de Conservacion Guanacaste</collectingCounty>
,
<collectingMunicipality id="6F9A3AA16A5534CC7E857339F55661E5">Sector San Cristobal</collectingMunicipality>
,
<location id="F8048DA53D192F032BAB14199B5910F4" LSID="urn:lsid:plazi:treatment:B1E6960C7A1359EE8013F4386196B7B8:F8048DA53D192F032BAB14199B5910F4" country="Costa Rica" county="Area de Conservacion Guanacaste" latitude="10.88" longLatPrecision="1" longitude="-85.389" municipality="Sector San Cristobal" name="Estacion San Gerardo" stateProvince="Guanacaste">Estacion San Gerardo</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELVAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-85.389000,10.880000]}">
<geoCoordinate id="236214B2B8AAF985057E0CAB8AAF24EE" degrees="10" direction="north" minutes="52" orientation="latitude" precision="1" seconds="48.00" value="10.88">10 ° 52 ′ 48.00 ” N</geoCoordinate>
,
<geoCoordinate id="615DEADB08521B7E3AA799FA23FC8666" degrees="85" direction="west" minutes="23" orientation="longitude" precision="1" seconds="20.40" value="-85.389">85 ° 23 ′ 20.40 ” W</geoCoordinate>
</named-content>
, el.
<quantity id="AF1A4CB593C035E7EC3D5FAA818D0F9D" metricMagnitude="2" metricUnit="m" metricValue="5.75" unit="m" value="575.0">
<elevation id="3B58A2265BD1E7D9DA23C2EF484DF30B" metricMagnitude="2" metricUnit="m" metricValue="5.75" unit="m" value="575.0">575 m</elevation>
</quantity>
,
<date id="166F9F1CC7F04A339796F1C9CF0B8BA9" value="2013-08-26">
<collectingDate id="5E23A26F84268A299532FF488C3A1E8D" value="2013-08-26">26. viii. 2013</collectingDate>
</date>
,
<collectingMethod id="A351940032DB258DAC8F2788D030A75D">Malaise Trap</collectingMethod>
,
<collectorName id="2DB31036997C0A262AA72191787957BF">D. H. Janzen</collectorName>
,
<collectorName id="AA8EBE15473185897920601990D6EAEE">W. Hallwachs</collectorName>
leg., [BOLD:
<ext-link id="B075E2781F2085D2F4EF447BEF498361" ext-link-type="uri" xlink_href="http://www.boldsystems.org/index.php/Public_RecordView?processid=GMAAG1127-16" xlink_type="simple">GMAAG 1127-16</ext-link>
]
<specimenCode id="A6A99C27F6C28E8E2C4017807804FF87" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" type="University or college">BIOUG 28044</specimenCode>
- C 01
</materialsCitation>
. —
<materialsCitation id="5BD8B9014978F8DE296B65F11C9DF6F2" collectingDate="2013-09-09" collectingMethod="Malaise Trap" collectionCode="BIOUG" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Area de Conservacion" elevation="575" latitude="10.88" location="Estacion San Gerardo" longLatPrecision="1" longitude="-85.389" municipality="Sector San Cristobal" specimenCode="BIOUG 19725" specimenCount="2" specimenCount-male="2" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="DE6D11E53BBB09AE4DFC5E56D8FFA457" bold="true">
<typeStatus id="F44D0D45C957E931080A589DD985F7A7" type="paratype">Paratypes</typeStatus>
</emphasis>
: Same data as holotype, except 09. ix. 2013. •
<specimenCount id="25A5E80B02BFBFA32AE8C89B6885AE53" count="2" type="male">2 ♂</specimenCount>
;
<date id="657DFB2BF070030509D51F2AA775CEBA" value="2013-09-09">
<collectingDate id="6579140CB20EECFFACFDBF1CC9C1889C" value="2013-09-09">09. ix. 2013</collectingDate>
</date>
, [BOLD:
<ext-link id="93D641DFAAE029B78F50975BCA98DFB3" ext-link-type="uri" xlink_href="http://www.boldsystems.org/index.php/Public_RecordView?processid=GMACB1559-15" xlink_type="simple">GMACB 1559-15</ext-link>
]
<specimenCode id="4F3DC9DD762C00B73C0B85FE8D29B679" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" type="University or college">BIOUG 19725</specimenCode>
- A 07, [BOLD:
<ext-link id="06F9A6A84A6C4FD4A9445D1BDA6CDEC9" ext-link-type="uri" xlink_href="http://www.boldsystems.org/index.php/Public_RecordView?processid=GMACB1565-15" xlink_type="simple">GMACB 1565-15</ext-link>
]
<specimenCode id="B047A2B85E68C8DF5624F21965285E37" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" type="University or college">BIOUG 19725</specimenCode>
- B 01
</materialsCitation>
. •
<materialsCitation id="6A4D77581FD41439AA83B3C29775A911" collectingDate="2013-08-26" collectingMethod="Malaise Trap" collectionCode="BIOUG" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Area de Conservacion" elevation="575" latitude="10.88" location="Estacion San Gerardo" longLatPrecision="1" longitude="-85.389" municipality="Sector San Cristobal" specimenCode="GMAAG 1127, BIOUG 28044" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="E9115473BC8EFCF7AD870006C1CEEF50" count="1" type="male">♂</specimenCount>
; same data, except
<date id="20BAFE39D0516B2F26FDC5A792981B52" value="2013-08-26">
<collectingDate id="AA1D5F83FF0EFB6AD4A7DAC4FC58B831" value="2013-08-26">26. viii. 2013</collectingDate>
</date>
, [BOLD:
<ext-link id="21680FF88E71815E69FB00FFEE1229E6" ext-link-type="uri" xlink_href="http://www.boldsystems.org/index.php/Public_RecordView?processid=GMAAG1127-16" xlink_type="simple">
<specimenCode id="33362CC72F4FAF2154B39E27AADA32B4">GMAAG 1127</specimenCode>
-16
</ext-link>
]
<specimenCode id="46F71D25C11EDA92249FF1434E26BDE5" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" type="University or college">BIOUG 28044</specimenCode>
- C 01
</materialsCitation>
. •
<materialsCitation id="31C85A914C0988D0237204B9114D32C7" collectingDate="2014-03-31" collectingMethod="Malaise Trap" collectionCode="BIOUG" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Area de Conservacion" elevation="575" latitude="10.88" location="Estacion San Gerardo" longLatPrecision="1" longitude="-85.389" municipality="Sector San Cristobal" specimenCode="GMAAQ 746, BIOUG 28344" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="425E62CF6C14BBFD05CA573F20F0CBA9" count="1" type="male">♂</specimenCount>
; same data, except
<date id="222C3474355247D5C24D54183DD2940F" value="2014-03-31">
<collectingDate id="9CFAEEC8C5664F7A55027897802D21D0" value="2014-03-31">31. iii. 2014</collectingDate>
</date>
, [BOLD:
<ext-link id="0A14756206CF95F06A80B25FA29D158C" ext-link-type="uri" xlink_href="http://www.boldsystems.org/index.php/Public_RecordView?processid=GMAAQ746-16" xlink_type="simple">
<specimenCode id="1A3043B1D0B7CD1C68C5CFC94415D4B2">GMAAQ 746</specimenCode>
-16
</ext-link>
]
<specimenCode id="EDDBE8ABEB6BEA26E19945D7ECB125B0" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" type="University or college">BIOUG 28344</specimenCode>
- B 02
</materialsCitation>
. •
<materialsCitation id="457CA841C526B151D84311F71E91FD56" collectingDate="2013-09-11" collectingMethod="Malaise Trap" collectionCode="MZUSP" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Area de Conservacion" elevation="575" latitude="10.88" location="Estacion San Gerardo" longLatPrecision="1" longitude="-85.389" municipality="Sector San Cristobal" specimenCode="GMACA 1002" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="DFBBEDE82C4E8B09CBE4EB521995CC28" count="1" type="male">♂</specimenCount>
; same data, except
<date id="CFE065FDB78F43EC0F14B69DC47F5271" value="2013-09-11">
<collectingDate id="B368D2667DACA4E702EB75EB51D5D9E7" value="2013-09-11">11. ix. 2013</collectingDate>
</date>
, [BOLD:
<ext-link id="508E9F55ADB17D4C87E55BAEF2A165F6" ext-link-type="uri" xlink_href="http://www.boldsystems.org/index.php/Public_RecordView?processid=GMACA1002-15" xlink_type="simple">
<specimenCode id="C9EE410525DD8776738EA0BF0BF9C7E1">GMACA 1002</specimenCode>
-15
</ext-link>
]
<named-content id="4DCB263C3E48C90AE2CA5F19DE9C838A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoloogia da Universidade de São Paulo">
<collectionCode id="26C376393B15AB008FB6AC967E228A2E" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" type="">MZUSP</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUWAI" type="diagnosis">
<paragraph id="34B3275F9F08F912962D4B468803620C">
<heading id="8B47DAD4DF7A26FADEC31E2D3CF04EFF" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="3FC53CAD46BDD0CFDB39281932A3F9D2">
The new species is most similar to
<taxonomicName id="D1A103D14A50B1DD29B0020871F3F0A9" authority="Vilarino &amp; Calor, 2024" authorityName="Vilarino &amp; Calor" authorityYear="2024" class="Insecta" family="Xiphocentronidae" genus="Xiphocentron" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tuxtla" status="sp. nov.">
<emphasis id="543DB5B4D9831E8D6986371E17796499" italics="true">X. tuxtla</emphasis>
</taxonomicName>
<emphasis id="B5443E29AE2BFB2E4B9D3B1E269CE0D0" bold="true">
<taxonomicNameLabel id="924CE2BCC997A571AE558A8DCE7132FE" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by sharing the prominent acute lobes at the posterior margin of sternum IX and the slightly divergent lobes of tergum IX posterior margin. The new species can be diagnosed in lateral view by (1) the preanal appendage with about the same width throughout length; (2) the spine-like setae of inferior appendage numerous and very dense (sparser basely in
<taxonomicName id="5CDBAF3285F4D98F46DEA2795B108B91" authority="Vilarino &amp; Calor, 2024" authorityName="Vilarino &amp; Calor" authorityYear="2024" class="Insecta" family="Xiphocentronidae" genus="Xiphocentron" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tuxtla" status="sp. nov.">
<emphasis id="97414F51DC8D271E3930827442B9ADCA" italics="true">X. tuxtla</emphasis>
</taxonomicName>
<emphasis id="D3F37C60EBF10B31B8F79E15434C51B9" bold="true">
<taxonomicNameLabel id="238CBFF2B29CA3FA3EF7F0BFC58AC2BE" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); and (3) the shape of paraproct apicodorsaly round (truncate in
<taxonomicName id="47F0F9A9E76B283741493BCA765BD23A" authority="Vilarino &amp; Calor, 2024" authorityName="Vilarino &amp; Calor" authorityYear="2024" class="Insecta" family="Xiphocentronidae" genus="Xiphocentron" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tuxtla" status="sp. nov.">
<emphasis id="57C1A822ED90DA4F4FD2B7305D999E65" italics="true">X. tuxtla</emphasis>
</taxonomicName>
<emphasis id="93E0C6524892F25606C03843ECF98AC9" bold="true">
<taxonomicNameLabel id="7C5F58AE8421FBC2EFCA7A59A21DAA45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). In dorsal view (4) the tergum IX is longer than wide (about as long as wide in
<taxonomicName id="DFC6C4754D4D2FCE41D783B6D04B8491" authority="Vilarino &amp; Calor, 2024" authorityName="Vilarino &amp; Calor" authorityYear="2024" class="Insecta" family="Xiphocentronidae" genus="Xiphocentron" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tuxtla" status="sp. nov.">
<emphasis id="00CF98966E79873E38F3FA15A14B7EC4" italics="true">X. tuxtla</emphasis>
</taxonomicName>
<emphasis id="3A06107AC1E975F6F60C0497170EC4F8" bold="true">
<taxonomicNameLabel id="66EDB75904C6979BB8B5D1632737D4E5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUYAI" type="description">
<paragraph id="74C00B2E3F3D07654763FAF4A6EEB4C0">
<heading id="82FB48766A81A9AAC89D19D4CD1A2814" reason="title">Description.</heading>
</paragraph>
<paragraph id="9D1E6E48142A794EE3CBA179C998FBD8">
<emphasis id="DCC32634AE823861EAAC13204B056947" bold="true" italics="true">Male</emphasis>
: Forewing length
<quantity id="85BDA24541E8E2D5F1AB7C3B60033466" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" unit="mm" value="4.1" valueMax="4.2" valueMin="4.0">4–4.2 mm</quantity>
(n = 3). Color overall pale brown, forewing uniformly dark brown. Maxillary palp segment length formula (I = II = III) &lt;IV &lt;V. Tibial spur formula 2: 4: 3; spurs unmodified. Forewing forks II and IV present; fork II sessile at discoidal cell; discoidal cell half as long as thyridial cell. Hind wing forks II and V present (Fig.
<figureCitation id="7043F6F6515BCBF82146C07A38BDB4B4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 A</figureCitation>
). Sternum V with anterolateral reticulated region and wide sclerotized lateral projection.
</paragraph>
<caption id="90D036961C78C6CC37371B47A1AEBED3" ID-DOI="10.3897/asp.82.e112587.figure9" ID-arpha="225ACA67-4AA2-543A-B057-38DD859FC8AA" httpUri="https://binary.pensoft.net/fig/1053147" startId="F9">
<paragraph id="E300742720536BA521DB3AFE1960A11A">
<label id="A10F13A31B3F2263DA674DA4CC869745">Figure 9.</label>
</paragraph>
<paragraph id="AF745A31D2002485A93A29F59D7209EB">
<taxonomicName id="883D0331324A25E741EB959D2C3F9905" authority="Vilarino &amp; Calor, 2024" authorityName="Vilarino &amp; Calor" authorityYear="2024" class="Insecta" family="Xiphocentronidae" genus="Xiphocentron" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="guanacaste" status="sp. nov." subGenus="Sphagocentron">
<emphasis id="FEB70648EABB49C5F9DC649C937FFFCA" italics="true">Xiphocentron</emphasis>
(
<emphasis id="013B4D79593D52C541D5B3189C884183" italics="true">Sphagocentron</emphasis>
)
<emphasis id="75B0AA75A82386DD5591466E3713F371" italics="true">guanacaste</emphasis>
</taxonomicName>
<emphasis id="7E9E66F0F1FB2AC42399500EC9AD2A78" bold="true">
<taxonomicNameLabel id="FA9F060505735D101D7FD87EB23963F7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (
<named-content id="C54535D0FD123160955C8995FC603F35" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/university-guelph" xlink_title="Centre for Biodiversity Genomics, University of Guelph">BIOUG</named-content>
).
<emphasis id="0CAB88EEE72DDE5CB85C49341E20B15B" bold="true">A</emphasis>
wing venation left. Genitalia:
<emphasis id="FB5F9CB61BE0BA68DBD65EB64E855789" bold="true">B</emphasis>
left lateral,
<emphasis id="A1CB6D753412F98226477FFE7284D31F" bold="true">C</emphasis>
dorsal,
<emphasis id="DB73CAA294F431F519527DAF6C06FA72" bold="true">D</emphasis>
ventral. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="3EA883CDB2F1AA295C0F0EA0F5339E6F">
Genitalia (Fig.
<figureCitation id="113F9298F22E88D9EE5F311014E8A9F5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 B – D</figureCitation>
). Tergum IX, in lateral view, wider basally, narrower apically (Fig.
<figureCitation id="75B78CA611405B4986D6B81C43FA93D1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 B</figureCitation>
); in dorsal view, anterior margin with deep wide concave incision; posterior margin with round lobes and deep v-shape mesal incision (Fig.
<figureCitation id="69297FDF9B1D243CF0F8A3188266620A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 C</figureCitation>
). Sternum IX, in lateral view, about 3 × as long as high, apex subtruncate, with narrow, straight anterior apodeme, tapering to narrow flange (Fig.
<figureCitation id="F19566D8F5115F1501CBC77A2C23EB5A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 B</figureCitation>
); in ventral view, longer than wide, posterior margin produced mesally, forming prominent acute lobes divided by v-shaped mesal incision (Fig.
<figureCitation id="19C08CB3484AD2B2FC4CA5422564F03C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 D</figureCitation>
). Tergum X membranous fused basodorsally to each paraproct. Paraproct, in lateral view, oblong, width subequal throughout length, dorsoapically round, with narrow, short ventral lobe (Fig.
<figureCitation id="4430D6D1B66D7AD22D76BB32DD257BC8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 B</figureCitation>
); in dorsal view, wide basally, divided apicomesally, each side partially fused at midlenght, without clear mesal band, apex with several sensillae (Fig.
<figureCitation id="DD280850800D16ABC58CCF212B9A5D48" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 C</figureCitation>
). Preanal appendage setose, in lateral view more than 2 × as long as tergum IX, mostly straight to slightly wavy, with same width throughout length, apex tapered, acute (Fig.
<figureCitation id="7E21D4388B0295BCC5AE234AF60AD440" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 B</figureCitation>
); in dorsal view, about same width throughout length (Fig.
<figureCitation id="19463779B07D01F5079703972B6963E4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 C</figureCitation>
). Inferior appendage in lateral view about 3 × as long as tergum IX; coxopodite and harpago completly fused, without suture line between each article, inner face with area of dense, long spine-like setae, and row of regular setae from midlenght to apex (Fig.
<figureCitation id="F3BBE919576BDEBF6C2A66979F9E2BD3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 B</figureCitation>
); basal region (coxopodite) wide, without basomesal spine-like setae, mesal sclerite region with dense, long spine-like setae, longer than other spinelike-setae; apical region (harpago) narrow, digitate, 1.5 × as long as basal region, apex slightly enlarged (Fig.
<figureCitation id="EC0DA9630AA384D0C198B7B39B29297E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 B</figureCitation>
); basal plate in lateral view wide, anteriorly with long, narrow flange (Fig.
<figureCitation id="68B39807B4A87BA4D12BB4C9BD40A7EF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xiphocentron (Sphagocentron) guanacaste sp. nov., holotype, male (BIOUG). A wing venation left. Genitalia: B left lateral, C dorsal, D ventral. Scale bar 0.1 mm." figureDoi="10.3897/asp.82.e112587.figure9" httpUri="https://binary.pensoft.net/fig/1053147">9 B</figureCitation>
). Phallus tubular, very long and narrow, reaching segment V, basally conical, subapically annulate, weakly sclerotized, apex slightly enlarged.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E52AI" type="etymology">
<paragraph id="285C20FED7D0C4BC4DECA85C02EBC2C1">
<heading id="D3E9093B13D570A38593BEBC0C43FB79" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="12DDE6BC2A47D497E64583EB501B1D3E">
Name in apposition; from the indigenous Nahuatl language: guaitil, ‘ tree’, and nacaztli, ‘ ear’. It is the popular name of the tree
<tp_taxon-name id="4135A8AC36413A705775FA64359029AD">
<emphasis id="20C053F32946C94381F65572EA1A6DA9" italics="true">
<tp_taxon-name-part id="D8C8510978380BFCA72447C90A4046FB" reg="Entelorobium" taxon-name-part-type="genus">Entelorobium</tp_taxon-name-part>
<tp_taxon-name-part id="0FD76CA181AFEFB0E2FE58FD4642418E" reg="cyclocarpum" taxon-name-part-type="species">cyclocarpum</tp_taxon-name-part>
</emphasis>
</tp_taxon-name>
and also the name of the conservation area where the species was collected.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EP3AI" type="distribution">
<paragraph id="C7680229405F6EFD5839E85A78809BAC">
<heading id="E60E1BA3330622BF87F4D4C542A701B3" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="F686DECF3A63F29B3735481BE57B3263">
<collectingCountry id="28151D4F2C8C7C8471C51695555E0136" name="Costa Rica">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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