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<document ID-DOI="http://doi.org/10.5281/zenodo.5072601" ID-GBIF-Dataset="2ab421bf-d013-4c38-97a7-60ce19394e0d" ID-GBIF-Taxon="183325065" ID-ISSN="1982-0224" ID-Zenodo-Dep="5072601" checkinTime="1614363209436" checkinUser="carolina" docAuthor="González-Bergonzoni, Iván, Loureiro, Marcelo &amp; Oviedo, Sebastián" docDate="2009" docId="A91CAC3FFFE6FFD9D5D95C1F54F602B0" docLanguage="en" docName="NeotropIchthyol.7.1.19-24.pdf" docOrigin="Neotropical Ichthyology 7 (1)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252009000100003&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Gymnogeophagus tiraparae González-Bergonzoni &amp; Loureiro &amp; Oviedo 2009, new species" docType="treatment" docVersion="5" lastPageId="2" lastPageNumber="21" masterDocId="5525D447FFE7FFDBD5065F6550130538" masterDocTitle="A new species of Gymnogeophagus from the río Negro and río Tacuarí basins, Uruguay (Teleostei: Perciformes)" masterLastPageNumber="24" masterPageNumber="19" pageId="1" pageNumber="20" updateTime="1636472636074" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Gymnogeophagus from the río Negro and río Tacuarí basins, Uruguay (Teleostei: Perciformes)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>González-Bergonzoni, Iván</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Loureiro, Marcelo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oviedo, Sebastián</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-03-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>24</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252009000100003&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252009000100003</mods:identifier>
<mods:identifier type="GBIF-Dataset">2ab421bf-d013-4c38-97a7-60ce19394e0d</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4567199</mods:identifier>
<mods:identifier type="ZooBank">685B88DD-2FCE-4A42-B352-4D2E9DD4AEC8</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5072601" ID-GBIF-Taxon="183325065" ID-Zenodo-Dep="5072601" LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0" httpUri="http://treatment.plazi.org/id/A91CAC3FFFE6FFD9D5D95C1F54F602B0" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<subSubSection pageId="1" pageNumber="20" type="nomenclature">
<paragraph blockId="1.[223,676,890,945]" pageId="1" pageNumber="20">
<emphasis bold="true" box="[223,676,890,913]" pageId="1" pageNumber="20">
<taxonomicName authority="González-Bergonzoni &amp; Loureiro &amp; Oviedo, 2009" authorityName="González-Bergonzoni &amp; Loureiro &amp; Oviedo" authorityYear="2009" box="[223,532,890,913]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="tiraparae" status="new species">
<emphasis bold="true" box="[223,532,890,913]" italics="true" pageId="1" pageNumber="20">Gymnogeophagus tiraparae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[545,676,890,913]" pageId="1" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</emphasis>
<figureCitation box="[423,515,922,945]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[140,179,1879,1902]" captionStartId-1="2.[816,855,510,533]" captionTargetBox-0="[127,1451,1372,1862]" captionTargetBox-1="[816,1459,178,496]" captionTargetId-0="figure-685@1.[127,1451,1372,1862]" captionTargetId-1="figure-856@2.[816,1459,178,496]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Gymnogeophagus tiraparae, holotype, ZVC-P 7870, 99.7 mm SL, male, río Tacuarembó on Road 26, Pueblo Ansina, tributary to the río Negro (lower río Uruguay drainage)." captionText-1="Fig. 2. Gymnogeophagus tiraparae, male not preserved from the río Yí, tributary to the río Negro (lower río Uruguay drainage)." figureDoi-0="http://doi.org/10.5281/zenodo.4567201" figureDoi-1="http://doi.org/10.5281/zenodo.4567203" httpUri-0="https://zenodo.org/record/4567201/files/figure.png" httpUri-1="https://zenodo.org/record/4567203/files/figure.png" pageId="1" pageNumber="20">Figs. 1-2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="20" type="materials_examined">
<paragraph blockId="1.[129,772,985,1121]" pageId="1" pageNumber="20">
<materialsCitation ID-GBIF-Occurrence="3313990312" collectingDate="2005-11" collectorName="M. Loureiro &amp; F. Teixeira de Mello &amp; S. Oviedo &amp; A. D'Anatro &amp; I. Gonzalez." country="Uruguay" county="Pueblo Ansina" latitude="-31.883612" location="Teixeira de Mello" longLatPrecision="20" longitude="-55.477222" municipality="Road" pageId="1" pageNumber="20" specimenCount="1" stateProvince="Tacuarembo" typeStatus="holotype">
<emphasis bold="true" box="[129,229,985,1006]" pageId="1" pageNumber="20">
<typeStatus box="[129,225,985,1006]" pageId="1" pageNumber="20">Holotype</typeStatus>
.
</emphasis>
ZVC-P 7870,
<quantity box="[366,450,985,1006]" metricMagnitude="-2" metricUnit="m" metricValue="9.97" pageId="1" pageNumber="20" unit="mm" value="99.7">99.7 mm</quantity>
SL, male,
<collectingCountry box="[552,638,985,1006]" name="Uruguay" pageId="1" pageNumber="20">Uruguay</collectingCountry>
, Tacuarembó,
<collectingCounty box="[129,273,1014,1035]" pageId="1" pageNumber="20">Pueblo Ansina</collectingCounty>
, río
<collectingRegion box="[316,436,1014,1035]" country="Uruguay" name="Tacuarembo" pageId="1" pageNumber="20">Tacuarembó</collectingRegion>
at
<collectingMunicipality box="[466,517,1014,1035]" pageId="1" pageNumber="20">Road</collectingMunicipality>
26, tributary of río Negro (lower río Uruguay drainage),
<geoCoordinate box="[415,532,1043,1064]" degrees="31" direction="south" minutes="53" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="01" value="-31.883612">31º5301S</geoCoordinate>
,
<geoCoordinate box="[540,664,1043,1064]" degrees="55" direction="west" minutes="28" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="38" value="-55.477222">55º2838W</geoCoordinate>
,
<date box="[672,767,1043,1064]" pageId="1" pageNumber="20" value="2005-11">
<collectingDate box="[672,767,1043,1064]" pageId="1" pageNumber="20" value="2005-11">Nov 2005</collectingDate>
</date>
,
<collectorName box="[129,253,1071,1092]" pageId="1" pageNumber="20">M. Loureiro</collectorName>
,
<collectorName box="[265,471,1071,1092]" pageId="1" pageNumber="20">
F.
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD4245B4A51C4017C" box="[290,471,1071,1092]" country="Uruguay" county="Pueblo Ansina" latitude="-31.883612" longLatPrecision="20" longitude="-55.477222" municipality="Road" name="Teixeira de Mello" pageId="1" pageNumber="20" stateProvince="Tacuarembo">Teixeira de Mello</location>
</collectorName>
,
<collectorName box="[483,585,1071,1092]" pageId="1" pageNumber="20">S. Oviedo</collectorName>
,
<collectorName box="[596,721,1071,1092]" pageId="1" pageNumber="20">A. DAnatro</collectorName>
&amp;
<collectorName pageId="1" pageNumber="20">I. González.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="1.[129,772,1158,1352]" lastBlockId="1.[809,1452,177,1033]" pageId="1" pageNumber="20">
<materialsCitation ID-GBIF-Occurrence="3313990311" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<emphasis bold="true" box="[129,237,1158,1179]" pageId="1" pageNumber="20">
<typeStatus box="[129,233,1158,1179]" pageId="1" pageNumber="20">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty box="[242,273,1158,1179]" pageId="1" pageNumber="20">All</collectingCounty>
from
<collectingCountry box="[328,414,1158,1179]" name="Uruguay" pageId="1" pageNumber="20">Uruguay</collectingCountry>
.
<collectingMunicipality box="[421,484,1158,1179]" pageId="1" pageNumber="20">Lower</collectingMunicipality>
río
<collectingCountry box="[519,603,1158,1179]" name="Uruguay" pageId="1" pageNumber="20">Uruguay</collectingCountry>
drainage: ZVC-P 2690, 2,
<quantity box="[217,357,1187,1208]" metricMagnitude="-2" metricUnit="m" metricValue="4.87" metricValueMax="5.22" metricValueMin="4.52" pageId="1" pageNumber="20" unit="mm" value="48.7" valueMax="52.2" valueMin="45.2">45.2-52.2 mm</quantity>
SL,
<collectingRegion box="[406,492,1187,1208]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD4F15BC652B60180" box="[503,677,1187,1208]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Arroyo Cordobes" pageId="1" pageNumber="20" stateProvince="Durazno">Arroyo Cordobés</location>
,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD7B65BC6510201EC" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Estancia Las Pitangas" pageId="1" pageNumber="20" stateProvince="Durazno">Estancia Las Pitangas</location>
,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD4275BDA527801EC" box="[289,619,1215,1236]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Cercanias de Paso del Gordo" pageId="1" pageNumber="20" stateProvince="Durazno">Cercanías de Paso del Gordo</location>
,
<geoCoordinate box="[634,766,1215,1236]" degrees="32" direction="south" minutes="33" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="31" value="-32.55861">32º3331S</geoCoordinate>
,
<geoCoordinate box="[129,255,1244,1265]" degrees="55" direction="west" minutes="17" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="38" value="-55.293888">55º1738W</geoCoordinate>
,
<date box="[263,352,1244,1265]" pageId="1" pageNumber="20" value="1963-01">
<collectingDate box="[263,352,1244,1265]" pageId="1" pageNumber="20" value="1963-01">Jan 1963</collectingDate>
</date>
,
<collectorName box="[360,431,1244,1265]" pageId="1" pageNumber="20">Dolber</collectorName>
; ZVC-P 3703, 2,
<quantity box="[596,732,1244,1265]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.07" metricValueMin="6.93" pageId="1" pageNumber="20" unit="mm" value="75.0" valueMax="80.7" valueMin="69.3">69.3-80.7 mm</quantity>
SL,
<collectingRegion box="[129,215,1273,1294]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<collectingRegion box="[222,317,1273,1294]" country="Uruguay" name="Rio Negro" pageId="1" pageNumber="20">río Negro</collectingRegion>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990313" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<collectorName box="[325,385,1273,1294]" pageId="1" pageNumber="20">Paraje</collectorName>
329,
<collectorName box="[437,488,1273,1294]" pageId="1" pageNumber="20">Road</collectorName>
6,
<geoCoordinate box="[515,633,1273,1294]" degrees="32" direction="south" minutes="26" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="44" value="-32.445557">32º2644S</geoCoordinate>
,
<geoCoordinate box="[642,767,1273,1294]" degrees="55" direction="west" minutes="26" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="03" value="-55.434166">55º2603W</geoCoordinate>
,
<date box="[129,220,1302,1323]" pageId="1" pageNumber="20" value="1998-10">
<collectingDate box="[129,220,1302,1323]" pageId="1" pageNumber="20" value="1998-10">Oct 1998</collectingDate>
</date>
,
<collectorName box="[229,334,1302,1323]" pageId="1" pageNumber="20">F. Achaval</collectorName>
; ZVC-P 6684, 9,
<quantity box="[500,637,1302,1323]" metricMagnitude="-2" metricUnit="m" metricValue="5.159999999999999" metricValueMax="6.029999999999999" metricValueMin="4.29" pageId="1" pageNumber="20" unit="mm" value="51.599999999999994" valueMax="60.3" valueMin="42.9">42.9-60.3 mm</quantity>
SL,
<collectingRegion box="[681,767,1302,1323]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD5875A5650AF0070" box="[129,188,1331,1352]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="rio Yi" pageId="1" pageNumber="20" stateProvince="Durazno">río Yí</location>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990305" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<collectorName box="[200,354,1331,1352]" pageId="1" pageNumber="20">Paso San Borja</collectorName>
,
<geoCoordinate box="[364,484,1331,1352]" degrees="33" direction="south" minutes="23" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="50" value="-33.397224">33º2350S</geoCoordinate>
,
<geoCoordinate box="[495,621,1331,1352]" degrees="56" direction="west" minutes="24" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="10" value="-56.40278">56º2410W</geoCoordinate>
,
<date box="[632,733,1331,1352]" pageId="1" pageNumber="20" value="2005-11">
<collectingDate box="[632,733,1331,1352]" pageId="1" pageNumber="20" value="2005-11">Nov 2005</collectingDate>
</date>
,
<collectorName box="[744,901,177,1352]" pageId="1" pageNumber="20">M. Loureiro</collectorName>
,
<collectorName box="[915,1133,177,198]" pageId="1" pageNumber="20">
F.
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD6A95FD4547E05FE" box="[943,1133,177,198]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Teixeira de Mello" pageId="1" pageNumber="20" stateProvince="Durazno">Teixeira de Mello</location>
</collectorName>
,
<collectorName box="[1147,1254,177,198]" pageId="1" pageNumber="20">S. Oviedo</collectorName>
,
<collectorName box="[1267,1398,177,198]" pageId="1" pageNumber="20">A. DAnatro</collectorName>
&amp;
<collectorName pageId="1" pageNumber="20">I. González.</collectorName>
; ZVC-P 6687, 1,
<quantity box="[1090,1179,206,227]" metricMagnitude="-2" metricUnit="m" metricValue="5.72" pageId="1" pageNumber="20" unit="mm" value="57.2">57.2 mm</quantity>
SL,
<collectingRegion box="[1229,1328,206,227]" country="Uruguay" name="Rio Negro" pageId="1" pageNumber="20">río Negro</collectingRegion>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990302" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<collectorName box="[1340,1401,206,227]" pageId="1" pageNumber="20">Paraje</collectorName>
329,
<collectorName box="[809,859,235,256]" pageId="1" pageNumber="20">Road</collectorName>
6,
<collectingRegion box="[885,969,235,256]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<geoCoordinate box="[976,1092,235,256]" degrees="32" direction="south" minutes="26" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="44" value="-32.445557">32º2644S</geoCoordinate>
,
<geoCoordinate box="[1099,1221,235,256]" degrees="55" direction="west" minutes="26" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="03" value="-55.434166">55º2603W</geoCoordinate>
,
<date box="[1229,1324,235,256]" pageId="1" pageNumber="20" value="2005-11">
<collectingDate box="[1229,1324,235,256]" pageId="1" pageNumber="20" value="2005-11">Nov 2005</collectingDate>
</date>
,
<collectorName box="[1332,1448,235,256]" pageId="1" pageNumber="20">M. Loureiro</collectorName>
,
<collectorName box="[809,1005,264,285]" pageId="1" pageNumber="20">
F.
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD6395E6D53FE0425" box="[831,1005,264,285]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Teixeira de Mello" pageId="1" pageNumber="20" stateProvince="Durazno">Teixeira de Mello</location>
</collectorName>
,
<collectorName box="[1014,1113,264,285]" pageId="1" pageNumber="20">S. Oviedo</collectorName>
,
<collectorName box="[1120,1242,264,285]" pageId="1" pageNumber="20">A. DAnatro</collectorName>
&amp;
<collectorName box="[1271,1386,264,285]" pageId="1" pageNumber="20">I. González</collectorName>
; ZVC- P 6688, 1,
<quantity box="[904,987,292,313]" metricMagnitude="-2" metricUnit="m" metricValue="7.29" pageId="1" pageNumber="20" unit="mm" value="72.9">72.9 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD1025E41542F0401" box="[1028,1084,292,313]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="rio Yi" pageId="1" pageNumber="20" stateProvince="Durazno">río Yí</location>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990301" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<collectorName box="[1091,1235,292,313]" pageId="1" pageNumber="20">Paso San Borja</collectorName>
,
<collectingRegion box="[1242,1326,292,313]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<geoCoordinate box="[1333,1447,292,313]" degrees="33" direction="south" minutes="23" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="50" value="-33.397224">33º2350S</geoCoordinate>
,
<geoCoordinate box="[809,930,321,342]" degrees="56" direction="west" minutes="24" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="10" value="-56.40278">56º2410W</geoCoordinate>
,
<date box="[938,1031,321,342]" pageId="1" pageNumber="20" value="2005-11">
<collectingDate box="[938,1031,321,342]" pageId="1" pageNumber="20" value="2005-11">Nov 2005</collectingDate>
</date>
,
<collectorName box="[1039,1154,321,342]" pageId="1" pageNumber="20">M. Loureiro</collectorName>
,
<collectorName box="[1161,1346,321,342]" pageId="1" pageNumber="20">
F.
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD1985E245551046E" box="[1182,1346,321,342]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Teixeira de Mello" pageId="1" pageNumber="20" stateProvince="Durazno">Teixeira de Mello</location>
</collectorName>
,
<collectorName box="[1354,1448,321,342]" pageId="1" pageNumber="20">S. Oviedo</collectorName>
,
<collectorName box="[809,934,350,371]" pageId="1" pageNumber="20">A. DAnatro</collectorName>
&amp;
<collectorName box="[969,1086,350,371]" pageId="1" pageNumber="20">I. González</collectorName>
; ZVC-P 6689, 3,
<quantity box="[1266,1409,350,371]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" metricValueMax="6.59" metricValueMin="4.71" pageId="1" pageNumber="20" unit="mm" value="56.5" valueMax="65.9" valueMin="47.1">47.165.9 mm</quantity>
SL, same data as holotype; ZVC-P 6692, 3,
<quantity box="[1202,1340,379,400]" metricMagnitude="-2" metricUnit="m" metricValue="6.92" metricValueMax="7.95" metricValueMin="5.89" pageId="1" pageNumber="20" unit="mm" value="69.2" valueMax="79.5" valueMin="58.9">58.9-79.5 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD06D5E1E55B404A8" box="[1387,1447,379,400]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="rio Yi" pageId="1" pageNumber="20" stateProvince="Durazno">río Yí</location>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990314" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<collectorName box="[809,961,408,429]" pageId="1" pageNumber="20">Paso San Borja</collectorName>
,
<collectingRegion box="[971,1057,408,429]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<geoCoordinate box="[1068,1187,408,429]" degrees="33" direction="south" minutes="23" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="50" value="-33.397224">33º2350S</geoCoordinate>
,
<geoCoordinate box="[1198,1324,408,429]" degrees="56" direction="west" minutes="24" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="10" value="-56.40278">56º2410W</geoCoordinate>
, 2005,
<collectorName pageId="1" pageNumber="20">Pablo Laurino</collectorName>
; ZVC-P 6694, 1,
<quantity box="[1073,1162,436,457]" metricMagnitude="-2" metricUnit="m" metricValue="8.59" pageId="1" pageNumber="20" unit="mm" value="85.9">85.9 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD1B95ED154EE04F1" box="[1215,1277,436,457]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="rio Yi" pageId="1" pageNumber="20" stateProvince="Durazno">río Yí</location>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990308" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<collectorName box="[1291,1448,436,457]" pageId="1" pageNumber="20">Paso San Borja</collectorName>
,
<collectingRegion box="[809,895,465,486]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<geoCoordinate box="[905,1024,465,486]" degrees="33" direction="south" minutes="23" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="50" value="-33.397224">33º2350S</geoCoordinate>
,
<geoCoordinate box="[1034,1160,465,486]" degrees="56" direction="west" minutes="24" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="10" value="-56.40278">56º2410W</geoCoordinate>
, 2005,
<collectorName box="[1230,1372,465,486]" pageId="1" pageNumber="20">Pablo Laurino</collectorName>
; ZVC-P 6695, 1,
<quantity box="[900,989,494,515]" metricMagnitude="-2" metricUnit="m" metricValue="9.19" pageId="1" pageNumber="20" unit="mm" value="91.9">91.9 mm</quantity>
SL,
<collectingRegion box="[1041,1140,494,515]" country="Uruguay" name="Rio Negro" pageId="1" pageNumber="20">río Negro</collectingRegion>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990306" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<collectorName box="[1153,1214,494,515]" pageId="1" pageNumber="20">Paraje</collectorName>
329,
<collectorName box="[1274,1325,494,515]" pageId="1" pageNumber="20">Road</collectorName>
6,
<collectingRegion box="[1362,1448,494,515]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<geoCoordinate box="[809,928,523,544]" degrees="32" direction="south" minutes="26" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="44" value="-32.445557">32º2644S</geoCoordinate>
,
<geoCoordinate box="[937,1063,523,544]" degrees="55" direction="west" minutes="26" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="03" value="-55.434166">55º2603W</geoCoordinate>
,
<date box="[1071,1168,523,544]" pageId="1" pageNumber="20" value="2005-08">
<collectingDate box="[1071,1168,523,544]" pageId="1" pageNumber="20" value="2005-08">Aug 2005</collectingDate>
</date>
,
<collectorName box="[1177,1291,523,544]" pageId="1" pageNumber="20">I. González</collectorName>
; ZVC-P 7656, 2,
<quantity box="[809,950,552,573]" metricMagnitude="-2" metricUnit="m" metricValue="6.045" metricValueMax="7.63" metricValueMin="4.46" pageId="1" pageNumber="20" unit="mm" value="60.45" valueMax="76.3" valueMin="44.6">44.6-76.3 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD6EC5D4D543B0705" box="[1002,1064,552,573]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="rio Yi" pageId="1" pageNumber="20" stateProvince="Durazno">río Yí</location>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990309" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD1335D4D54DE0705" box="[1077,1229,552,573]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Polanco del Yi" pageId="1" pageNumber="20" stateProvince="Durazno">Polanco del Yí</location>
,
<collectingRegion box="[1243,1316,552,573]" country="Uruguay" name="Florida" pageId="1" pageNumber="20">Florida</collectingRegion>
,
<geoCoordinate box="[1329,1448,552,573]" degrees="33" direction="south" minutes="18" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="45" value="-33.3125">33º1845S</geoCoordinate>
,
<geoCoordinate box="[809,935,580,601]" degrees="56" direction="west" minutes="09" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="08" value="-56.152225">56º0908W</geoCoordinate>
,
<date box="[949,1050,580,601]" pageId="1" pageNumber="20" value="2008-03">
<collectingDate box="[949,1050,580,601]" pageId="1" pageNumber="20" value="2008-03">Mar 2008</collectingDate>
</date>
,
<collectorName box="[1063,1188,580,601]" pageId="1" pageNumber="20">M. Loureiro</collectorName>
,
<collectorName box="[1201,1411,580,601]" pageId="1" pageNumber="20">
F.
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD1CA5D2155900761" box="[1228,1411,580,601]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Teixeira de Mello" pageId="1" pageNumber="20" stateProvince="Durazno">Teixeira de Mello</location>
</collectorName>
,
<collectorName pageId="1" pageNumber="20">M. Zarucki</collectorName>
,
<collectorName box="[895,990,609,630]" pageId="1" pageNumber="20">L. Ziegler</collectorName>
&amp;
<collectorName box="[1019,1132,609,630]" pageId="1" pageNumber="20">I. González</collectorName>
; ZVC-P 7658, 1, 62.0 mm SL, río
<collectingRegion box="[809,927,638,659]" country="Uruguay" name="Tacuarembo" pageId="1" pageNumber="20">Tacuarembó</collectingRegion>
<collectorName box="[932,992,638,659]" pageId="1" pageNumber="20">Chico</collectorName>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990310" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<collectorName box="[1000,1045,638,659]" pageId="1" pageNumber="20">Paso</collectorName>
de los
<collectorName box="[1110,1195,638,659]" pageId="1" pageNumber="20">Novillos</collectorName>
,
<collectingRegion box="[1203,1324,638,659]" country="Uruguay" name="Tacuarembo" pageId="1" pageNumber="20">Tacuarembó</collectingRegion>
, 32º1109S, 55º2748,
<date box="[936,1043,667,688]" pageId="1" pageNumber="20" value="2008-03">
<collectingDate box="[936,1043,667,688]" pageId="1" pageNumber="20" value="2008-03">Mar 2008</collectingDate>
</date>
,
<collectorName box="[1057,1279,667,688]" pageId="1" pageNumber="20">
F.
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD1385DFE54EC0788" box="[1086,1279,667,688]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Teixeira de Mello" pageId="1" pageNumber="20" stateProvince="Durazno">Teixeira de Mello</location>
</collectorName>
,
<collectorName box="[1293,1415,667,688]" pageId="1" pageNumber="20">M. Masdeu</collectorName>
,
<collectorName pageId="1" pageNumber="20">G. Goyenola</collectorName>
,
<collectorName box="[915,1045,696,717]" pageId="1" pageNumber="20">M. Goyenola</collectorName>
,
<collectorName box="[1054,1168,696,717]" pageId="1" pageNumber="20">J. Clemente</collectorName>
&amp;
<collectorName box="[1198,1312,696,717]" pageId="1" pageNumber="20">I. González</collectorName>
; ZVC-P 7938, 5,
<quantity box="[834,972,724,745]" metricMagnitude="-2" metricUnit="m" metricValue="7.969999999999999" metricValueMax="9.709999999999999" metricValueMin="6.2299999999999995" pageId="1" pageNumber="20" unit="mm" value="79.69999999999999" valueMax="97.1" valueMin="62.3">62.3-97.1 mm</quantity>
SL,
<collectingRegion box="[1019,1115,724,745]" country="Uruguay" name="Rio Negro" pageId="1" pageNumber="20">río Negro</collectingRegion>
, embalse de represa
<collectorName box="[1318,1448,724,745]" pageId="1" pageNumber="20">de Baygorria</collectorName>
,
<collectingRegion box="[809,895,753,774]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<geoCoordinate box="[903,1020,753,774]" degrees="32" direction="south" minutes="52" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="21" value="-32.872498">32º5221S</geoCoordinate>
,
<geoCoordinate box="[1029,1154,753,774]" degrees="56" direction="west" minutes="48" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="13" value="-56.80361">56º4813W</geoCoordinate>
,
<date box="[1163,1253,753,774]" pageId="1" pageNumber="20" value="2008-10">
<collectingDate box="[1163,1253,753,774]" pageId="1" pageNumber="20" value="2008-10">Oct 2008</collectingDate>
</date>
,
<collectorName box="[1261,1452,753,774]" pageId="1" pageNumber="20">
F.
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD0055D9455BF063E" box="[1283,1452,753,774]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Teixeira de Mello" pageId="1" pageNumber="20" stateProvince="Durazno">Teixeira de Mello</location>
</collectorName>
&amp;
<collectorName box="[832,981,782,803]" pageId="1" pageNumber="20">Federico Viana</collectorName>
;
<specimenCode box="[989,1119,782,803]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="20">MNHN 3277</specimenCode>
,
<specimenCode box="[1127,1141,782,803]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="20">6</specimenCode>
,
<quantity box="[1149,1288,782,803]" metricMagnitude="-2" metricUnit="m" metricValue="5.709999999999999" metricValueMax="6.659999999999999" metricValueMin="4.76" pageId="1" pageNumber="20" unit="mm" value="57.099999999999994" valueMax="66.6" valueMin="47.6">47.666.6 mm</quantity>
SL, same data as
<typeStatus box="[809,898,811,832]" pageId="1" pageNumber="20">holotype</typeStatus>
. laguna Merin drainage: ZVC-P 7869, 1,
<quantity box="[1295,1380,811,832]" metricMagnitude="-2" metricUnit="m" metricValue="4.57" pageId="1" pageNumber="20" unit="mm" value="45.7">45.7 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD0975C4E53670665" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="rio Tacuari" pageId="1" pageNumber="20" stateProvince="Durazno">río Tacuarí</location>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990303" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
Road 18,
<collectingRegion box="[993,1116,840,861]" country="Uruguay" name="Cerro Largo" pageId="1" pageNumber="20">Cerro Largo</collectingRegion>
,
<geoCoordinate box="[1128,1246,840,861]" degrees="32" direction="south" minutes="45" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="46" value="-32.76278">32º4546S</geoCoordinate>
,
<geoCoordinate box="[1259,1385,840,861]" degrees="53" direction="west" minutes="42" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="58" value="-53.71611">53º4258W</geoCoordinate>
, 2005, Pablo Laurino.
<specimenCode box="[954,1097,868,889]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="20">UFRGS 10203</specimenCode>
,
<specimenCode box="[1105,1118,868,889]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="20">3</specimenCode>
,
<quantity box="[1128,1260,868,889]" metricMagnitude="-2" metricUnit="m" metricValue="5.834999999999999" metricValueMax="6.43" metricValueMin="5.24" pageId="1" pageNumber="20" unit="mm" value="58.349999999999994" valueMax="64.3" valueMin="52.4">52.464.3mm</quantity>
SL, río
<collectingRegion box="[1334,1452,868,889]" country="Uruguay" name="Tacuarembo" pageId="1" pageNumber="20">Tacuarembó</collectingRegion>
y ruta 26,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990304" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
Pueblo Ansina, Dpto
<collectingRegion box="[1168,1301,897,918]" country="Uruguay" name="Tacuarembo" pageId="1" pageNumber="20">Tacuarembó</collectingRegion>
,
<geoCoordinate box="[1317,1447,897,918]" degrees="31" direction="south" minutes="53" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="01" value="-31.883612">31º5301S</geoCoordinate>
,
<geoCoordinate box="[809,936,926,947]" degrees="55" direction="west" minutes="28" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="38" value="-55.477222">55º2838W</geoCoordinate>
,
<date box="[946,1046,926,947]" pageId="1" pageNumber="20" value="2005-11">
<collectingDate box="[946,1046,926,947]" pageId="1" pageNumber="20" value="2005-11">Nov 2005</collectingDate>
</date>
,
<collectorName box="[1056,1179,926,947]" pageId="1" pageNumber="20">M. Loureiro</collectorName>
, A. Danatro,
<collectorName pageId="1" pageNumber="20">
F.
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD0385CFB537506E8" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Teixeira de Mello" pageId="1" pageNumber="20" stateProvince="Durazno">Teixeira de Mello</location>
</collectorName>
,
<collectorName box="[880,981,955,976]" pageId="1" pageNumber="20">S. Oviedo</collectorName>
,
<collectorName box="[992,1119,955,976]" pageId="1" pageNumber="20">I. González.</collectorName>
<specimenCode box="[1125,1273,955,976]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="20">UFRGS 10204</specimenCode>
,
<specimenCode box="[1284,1298,955,976]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="1" pageNumber="20">2</specimenCode>
,
<quantity box="[1309,1452,955,976]" metricMagnitude="-2" metricUnit="m" metricValue="6.544999999999999" metricValueMax="7.859999999999999" metricValueMin="5.2299999999999995" pageId="1" pageNumber="20" unit="mm" value="65.44999999999999" valueMax="78.6" valueMin="52.3">52.378.6 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD6525CBD538206D5" box="[852,913,984,1005]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="rio Yi" pageId="1" pageNumber="20" stateProvince="Durazno">río Yí</location>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3313990307" collectingDate="1963-01" collectingDateMax="2008-10" collectingDateMin="1963-01" collectionCode="MNHN, UFRGS" country="Uruguay" county="All" latitude="-33.397224" location="Arroyo Cordobes" longLatPrecision="20" longitude="-56.40278" municipality="Lower" pageId="1" pageNumber="20" specimenCode="MNHN 3277, 6, UFRGS 10203, 3, UFRGS 10204, 2" specimenCount="2" stateProvince="Durazno" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:A91CAC3FFFE6FFD9D5D95C1F54F602B0:246A4BF2FFE6FFDAD69A5CBD542B06D5" box="[924,1080,984,1005]" country="Uruguay" county="All" latitude="-33.397224" longLatPrecision="20" longitude="-56.40278" municipality="Lower" name="Paso San Borja" pageId="1" pageNumber="20" stateProvince="Durazno">Paso San Borja</location>
,
<collectingRegion box="[1092,1178,984,1005]" country="Uruguay" name="Durazno" pageId="1" pageNumber="20">Durazno</collectingRegion>
,
<geoCoordinate box="[1190,1310,984,1005]" degrees="33" direction="south" minutes="23" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="50" value="-33.397224">33º2350S</geoCoordinate>
,
<geoCoordinate box="[1321,1447,984,1005]" degrees="56" direction="west" minutes="24" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="10" value="-56.40278">56º2410W</geoCoordinate>
,
<date box="[809,908,1012,1033]" pageId="1" pageNumber="20" value="2005-11">
<collectingDate box="[809,908,1012,1033]" pageId="1" pageNumber="20" value="2005-11">Nov 2005</collectingDate>
</date>
,
<collectorName box="[919,1039,1012,1033]" pageId="1" pageNumber="20">I. González</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="diagnosis">
<paragraph blockId="1.[809,1452,1074,1353]" lastBlockId="2.[136,779,178,585]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<emphasis bold="true" box="[809,927,1074,1097]" pageId="1" pageNumber="20">Diagnosis.</emphasis>
The new species can be distinguished from all others
<taxonomicName box="[882,1076,1106,1129]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis box="[882,1076,1106,1129]" italics="true" pageId="1" pageNumber="20">Gymnogeophagus</emphasis>
</taxonomicName>
species by the presence of the following unique characters: adipose hump on head deeper than the dorsal-fin upper border, adipose hump with anterior profile vertical, extending from the upper lip to the dorsal-fin origin, absence of transversal bands on body, two horizontal series of moderately elongated light blue dots between dorsal-fin spines, and a series of light blue stripes between soft rays, sometimes merged with the second series of elongated dots, and always with a red ground color between series of dots, and caudal fin with dots vertically aligned on its distal border.
<taxonomicName authorityName="González-Bergonzoni &amp; Loureiro &amp; Oviedo" authorityYear="2009" box="[223,530,242,265]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="tiraparae">
<emphasis box="[223,530,242,265]" italics="true" pageId="2" pageNumber="21">Gymnogeophagus tiraparae</emphasis>
</taxonomicName>
can be further distinguished from
<taxonomicName authorityName="Hensel" baseAuthorityName="Malabarba &amp; Reis" baseAuthorityYear="1988" box="[287,450,274,297]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="gymnogenys">
<emphasis box="[287,450,274,297]" italics="true" pageId="2" pageNumber="21">G. gymnogenys</emphasis>
</taxonomicName>
by the E1 scale count (27-30, 96% with 28-30
<emphasis box="[310,340,306,329]" italics="true" pageId="2" pageNumber="21">vs.</emphasis>
26-29, 74% with
<quantity box="[526,614,306,329]" metricMagnitude="-1" metricUnit="m" metricValue="6.731" metricValueMax="6.858" metricValueMin="6.604" pageId="2" pageNumber="21" unit="in" value="26.5" valueMax="27.0" valueMin="26.0">26-27 in</quantity>
<taxonomicName authority="(Malabarba &amp; Reis, 1988))" authorityName="Hensel" baseAuthorityName="Malabarba &amp; Reis" baseAuthorityYear="1988" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="gymnogenys">
<emphasis box="[619,778,306,329]" italics="true" pageId="2" pageNumber="21">G. gymnogenys</emphasis>
(Malabarba &amp;
<bibRefCitation author="Reis, R" box="[305,429,338,361]" pageId="2" pageNumber="21" pagination="259 - 305" refId="ref5301" refString="Reis, R. E. &amp; L. R. Malabarba. 1988. Revision of the Neotropical genus Gymnogeophagus Ribeiro, 1918 with descriptions of two new species (Pisces, Perciformes). Revista Brasileira de Zoologia, 4: 259 - 305." type="journal article" year="1988">Reis, 1988</bibRefCitation>
))
</taxonomicName>
; from
<taxonomicName authorityName="Hensel" baseAuthorityName="Malabarba &amp; Reis" baseAuthorityYear="1988" box="[526,695,338,361]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="gymnogenys">
<emphasis box="[526,695,338,361]" italics="true" pageId="2" pageNumber="21">G. gymnogenys</emphasis>
</taxonomicName>
and
<taxonomicName class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="caguazuensis">
<emphasis italics="true" pageId="2" pageNumber="21">G. caguazuensis</emphasis>
</taxonomicName>
by the absence of a black line anterior to eye (
<emphasis box="[145,167,402,425]" italics="true" pageId="2" pageNumber="21">vs</emphasis>
. present); from
<taxonomicName authorityName="Hensel" baseAuthorityName="Malabarba &amp; Reis" baseAuthorityYear="1988" box="[350,515,402,425]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="gymnogenys">
<emphasis box="[350,515,402,425]" italics="true" pageId="2" pageNumber="21">G. gymnogenys</emphasis>
</taxonomicName>
and
<taxonomicName box="[573,700,402,425]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="australis">
<emphasis box="[573,700,402,425]" italics="true" pageId="2" pageNumber="21">G. australis</emphasis>
</taxonomicName>
by the body depth (33.6-41.4% SL
<emphasis box="[422,450,434,457]" italics="true" pageId="2" pageNumber="21">vs.</emphasis>
23-29% SL and 41.2-43.3% SL, respectively); from
<taxonomicName box="[350,475,466,489]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="australis">
<emphasis box="[350,475,466,489]" italics="true" pageId="2" pageNumber="21">G. australis</emphasis>
</taxonomicName>
by the presence six to eight parallel light green bands on body (
<emphasis box="[517,546,498,521]" italics="true" pageId="2" pageNumber="21">vs.</emphasis>
absence), by snout to dorsal-fin origin length (30.5-38.4% SL
<emphasis box="[566,595,530,553]" italics="true" pageId="2" pageNumber="21">vs.</emphasis>
23.6-30.7% SL), and by the head depth (81-104% HL
<emphasis box="[516,545,562,585]" italics="true" pageId="2" pageNumber="21">vs.</emphasis>
109.7-116.2% HL).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4567201" ID-Zenodo-Dep="4567201" httpUri="https://zenodo.org/record/4567201/files/figure.png" pageId="1" pageNumber="20" startId="1.[140,179,1879,1902]" targetBox="[127,1451,1372,1862]" targetPageId="1">
<paragraph blockId="1.[140,1463,1879,1934]" pageId="1" pageNumber="20">
<emphasis bold="true" box="[140,210,1879,1902]" pageId="1" pageNumber="20">Fig. 1.</emphasis>
<taxonomicName authorityName="González-Bergonzoni &amp; Loureiro &amp; Oviedo" authorityYear="2009" box="[216,518,1879,1902]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="tiraparae">
<emphasis box="[216,518,1879,1902]" italics="true" pageId="1" pageNumber="20">Gymnogeophagus tiraparae</emphasis>
</taxonomicName>
, holotype, ZVC-P 7870, 99.7 mm SL, male, río Tacuarembó on Road 26, Pueblo Ansina, tributary to the río Negro (lower río Uruguay drainage).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4567203" ID-Zenodo-Dep="4567203" httpUri="https://zenodo.org/record/4567203/files/figure.png" pageId="2" pageNumber="21" startId="2.[816,855,510,533]" targetBox="[816,1459,178,496]" targetPageId="2">
<paragraph blockId="2.[816,1459,510,597]" pageId="2" pageNumber="21">
<emphasis bold="true" box="[816,885,510,533]" pageId="2" pageNumber="21">Fig. 2.</emphasis>
<taxonomicName authorityName="González-Bergonzoni &amp; Loureiro &amp; Oviedo" authorityYear="2009" box="[891,1190,510,533]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="tiraparae">
<emphasis box="[891,1190,510,533]" italics="true" pageId="2" pageNumber="21">Gymnogeophagus tiraparae</emphasis>
</taxonomicName>
, male not preserved from the río Yí, tributary to the río Negro (lower río Uruguay drainage).
</paragraph>
</caption>
<subSubSection pageId="2" pageNumber="21" type="description">
<paragraph blockId="2.[136,779,626,1481]" pageId="2" pageNumber="21">
<emphasis bold="true" box="[136,276,626,649]" pageId="2" pageNumber="21">Description.</emphasis>
Standard length of specimens examined
<quantity metricMagnitude="-2" metricUnit="m" metricValue="8.285" metricValueMax="13.05" metricValueMin="3.52" pageId="2" pageNumber="21" unit="mm" value="82.85" valueMax="130.5" valueMin="35.2">35.2 to 130.5 mm</quantity>
; meristic and morphometric data in
<tableCitation box="[629,708,658,681]" captionStart="Table 1" captionStartId="3.[129,192,177,200]" captionTargetBox="[219,1364,233,646]" captionText="Table 1. Frequency distribution of meristic characters of Gymnogeophagus tiraparae Paratypes and Holotype (*) n = 38." httpUri="http://table.plazi.org/id/75CA4DA1FFE4FFD8D5875FD4558005F0" pageId="2" pageNumber="21" tableUuid="75CA4DA1FFE4FFD8D5875FD4558005F0">Table 1</tableCitation>
. Body elongated, laterally compressed. Predorsal contour convex. Reproductive males with adipose hump on top of head from dorsal-fin origin to upper lip, deeper than dorsal-fin upper border and with anterior profile vertical. Mouth terminal and jaws isognathus. Body contour at dorsal-fin base slightly arched, decreasing from anterior part to caudal peduncle. Caudal peduncle rectangular, longer than deeper, dorsal and ventral profiles slightly concave. Body contour slightly convex between lower lip and last anal-fin ray, with straight segment between pelvic and anal fins.
</paragraph>
<paragraph blockId="2.[136,779,626,1481]" pageId="2" pageNumber="21">Body scales moderately large and ctenoid except for small cycloid and ctenoid scales in preventral area. Small ctenoid scales on opercle; scales on preopercle, when present, small and cycloid. Small ctenoid scales on base of hump (in adult reproductive males only) to approximately vertical line passing on center of eye.</paragraph>
<paragraph blockId="2.[136,779,626,1481]" pageId="2" pageNumber="21">Proximal third to half of caudal fin with small scales in single series between rays. Dorsal-fin without scales and its origin posterior to vertical line through posterior bony margin of opercle. Large males with sixth dorsal-fin soft ray longest, reaching proximal third of caudal fin. Pectoral fin reaching anal-fin origin. Anal fin reaching caudal-fin base. Caudal fin truncated or slightly concave; lyrate in large reproductive males; very deep distally, from distal edge of extended dorsal fin to distal edge of extended anal fin.</paragraph>
<paragraph blockId="2.[136,779,1522,1929]" lastBlockId="2.[816,1459,625,1160]" pageId="2" pageNumber="21">
<emphasis bold="true" box="[136,276,1522,1545]" pageId="2" pageNumber="21">Color in life.</emphasis>
Ground color of dorsal region of body in adults, brownish to light olivaceous, with one large dark spot on nape and two large dark spots on body flank, just below dorsal-fin base, first between eighth and twelfth dorsal-fin spine and second from first to sixth dorsal-fin soft ray. Well defined dark, circular, lateral spot just below upper portion of lateral line; often with three or four dark blotches on sides of body, anterior dark blotch larger and extending through lateral spot, other blotches smaller and aligned on sides, reaching caudal peduncle. Ventral portion of body yellowish in adult males and light olivaceus in adult females. Six or more lateral, parallel, bright green to pale blue bands from posterior part of pectoral-fin base to caudal-fin base. Numerous small bright blue dots usually present on cheeks, bright green colored lips usually present in reproductive males. Adipose hump, when present, with brownish ground coloration without dark line anterior to eye, suborbital stripe usually absent or diffuse in reproductive males. Dorsal fin yellowish on base and reddish between two horizontal series of elongated, hyaline to light blue dots, and third distal series in posterior part of fin, often fused with proximal series; small proximal circular dots very close to dorsal-fin base often present, from the fifth or sixth dorsal-fin soft ray, to its distal end. Caudal fin with red ground color in adults and entirely covered with numerous circular hyaline dots, arranged into horizontal series between rays. Distal dots smaller, vertically aligned. Pectoral fin hyaline, pelvic fin with light blue stripes between fin rays; anal fin yellowish in proximal portion and reddish distally between many small light blue dots spread along fin.
</paragraph>
<paragraph blockId="2.[816,1460,1201,1320]" pageId="2" pageNumber="21">
<emphasis bold="true" box="[816,1004,1201,1224]" pageId="2" pageNumber="21">Color in alcohol.</emphasis>
Ground color turns paler. Reddish and yellowish pigmentations turn light brown or gray and light blue fin pigmentation turns hyaline. Dark spots turn darker and lateral parallel light green bands turn clearer, almost invisible.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="distribution">
<paragraph blockId="2.[816,1459,1361,1480]" pageId="2" pageNumber="21">
<emphasis bold="true" box="[816,964,1361,1384]" pageId="2" pageNumber="21">Distribution.</emphasis>
<taxonomicName authorityName="González-Bergonzoni &amp; Loureiro &amp; Oviedo" authorityYear="2009" box="[973,1278,1361,1384]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="tiraparae">
<emphasis box="[973,1278,1361,1384]" italics="true" pageId="2" pageNumber="21">Gymnogeophagus tiraparae</emphasis>
</taxonomicName>
is distributed in the middle
<collectingRegion box="[934,1036,1393,1416]" country="Uruguay" name="Rio Negro" pageId="2" pageNumber="21">rio Negro</collectingRegion>
basin, including main tributaries (lower río
<collectingCountry box="[854,948,1425,1448]" name="Uruguay" pageId="2" pageNumber="21">Uruguay</collectingCountry>
basin) and in río Tacuarí (laguna Mirim basin) (
<figureCitation box="[824,884,1457,1480]" captionStart="Fig" captionStartId="3.[129,168,1905,1928]" captionTargetBox="[132,771,1281,1870]" captionTargetId="figure-842@3.[129,772,1278,1873]" captionTargetPageId="3" captionText="Fig. 3. Distribution of Gymnogeophagus tiraparae. Star = type locality." figureDoi="http://doi.org/10.5281/zenodo.4567207" httpUri="https://zenodo.org/record/4567207/files/figure.png" pageId="2" pageNumber="21">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="biology_ecology">
<paragraph blockId="2.[816,1459,1521,1704]" pageId="2" pageNumber="21">
<emphasis bold="true" box="[816,913,1521,1544]" pageId="2" pageNumber="21">Ecology.</emphasis>
Localities where the new species was collected are large rivers, higher than stream order 6, with clear water, sandy or rocky bottoms, and little vegetation. Specimens were collected using cast nets, seine and electro fishing device. Females holding juveniles in their mouths were collected at the end of spring and summer.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="etymology">
<paragraph blockId="2.[816,1459,1745,1928]" pageId="2" pageNumber="21">
<emphasis bold="true" box="[816,945,1745,1768]" pageId="2" pageNumber="21">Etymology.</emphasis>
<taxonomicName authorityName="González-Bergonzoni &amp; Loureiro &amp; Oviedo" authorityYear="2009" box="[950,1249,1745,1768]" class="Actinopterygii" family="Cichlidae" genus="Gymnogeophagus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="tiraparae">
<emphasis box="[950,1249,1745,1768]" italics="true" pageId="2" pageNumber="21">Gymnogeophagus tiraparae</emphasis>
</taxonomicName>
takes its name from María Luisa Tirapare, a Guaraní woman who founded the now disappeared town of San Borja del Yí (close to the first locality where the new species was found), the last native town in Uruguayan land, where natives, fugitive African slaves, gauchos, and other outsiders lived together.
</paragraph>
</subSubSection>
</treatment>
</document>