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<document id="9CD318ADCBB66B23E91D4FFBC65ABD71" ID-CLB-Dataset="25474" ID-DOI="10.11646/zootaxa.4668.3.7" ID-GBIF-Dataset="ddf01fc8-a165-492e-aa9a-25316f7f4ab6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3449692" ID-ZooBank="AA9BB1DA-8E27-48E2-9A4A-7428C45F6291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568897931647" checkinUser="plazi" docAuthor="Volynkin, Anton V. &amp; László, Gyula M." docDate="2019" docId="102387C1FFE9426266F0FB40A946FD66" docLanguage="en" docName="zootaxa.4668.3.7.pdf" docOrigin="Zootaxa 4668 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tumicla smithi Volynkin &amp; László 2019, sp. nov." docType="treatment" docUuid="E41D7C53-A9DC-4075-8EA5-D2AA5113ECFD" docUuidSource="ZooBank" docVersion="11" lastPageNumber="434" masterDocId="EC1AFFB9FFE5426F6667FFC8AD35FFCD" masterDocTitle="Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini)" masterLastPageNumber="434" masterPageNumber="421" pageNumber="433" updateTime="1698755699668" updateUser="ExternalLinkService">
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<mods:title id="CC2DF280E84A2AE64DC4206CD36DCAFB">Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini)</mods:title>
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<mods:namePart id="86604086BD354C2A658BA227B02B5C4D">Volynkin, Anton V.</mods:namePart>
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<mods:namePart id="D0D7CC092FB738BB5B00555D3984DE67">László, Gyula M.</mods:namePart>
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<mods:date id="3C0DCAD1E2A5D13A79F53E3206B083A1">2019</mods:date>
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<mods:number id="878F7B2FFEDB9B9DDA03C1C574A3F92F">2019-09-11</mods:number>
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<subSubSection id="D090655CFFE9426366F0FB40AC87FB6E" box="[151,434,1160,1187]" pageId="12" pageNumber="433" type="nomenclature">
<paragraph id="983536D7FFE9426366F0FB40AC87FB6E" blockId="12.[151,434,1160,1223]" box="[151,434,1160,1187]" pageId="12" pageNumber="433">
<heading id="C37D81BBFFE9426366F0FB40AC87FB6E" bold="true" box="[151,434,1160,1187]" fontSize="11" level="1" pageId="12" pageNumber="433" reason="1">
<emphasis id="AAFEEAC5FFE9426366F0FB40AC87FB6E" bold="true" box="[151,434,1160,1187]" pageId="12" pageNumber="433">
<taxonomicName id="5F8A4D54FFE9426366F0FB40AC78FB6E" ID-CoL="BFXRH" authority="Volynkin &amp; László, 2019" authorityName="Volynkin &amp; László" authorityYear="2019" box="[151,333,1160,1187]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="AAFEEAC5FFE9426366F0FB40AC78FB6E" bold="true" box="[151,333,1160,1187]" italics="true" pageId="12" pageNumber="433">Tumicla smithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B1CD57BEFFE942636732FB41AC87FB6E" box="[341,434,1161,1187]" pageId="12" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="D090655CFFE9426366F0FB64AC64FB0A" box="[151,337,1196,1223]" pageId="12" pageNumber="433" type="description">
<paragraph id="983536D7FFE9426366F0FB64AC64FB0A" blockId="12.[151,434,1160,1223]" box="[151,337,1196,1223]" pageId="12" pageNumber="433">
(
<figureCitation id="00B12A52FFE9426366F8FB64AC15FB0A" box="[159,288,1196,1223]" captionStart="FIGURES 1926" captionStartId="5.[151,264,1899,1924]" captionTargetBox="[156,1435,181,1860]" captionTargetId="figure@5.[151,1435,181,1873]" captionTargetPageId="5" captionText="FIGURES 1926. Tumicla spp.: adults. 19, T. admiranda, holotype ♂, DR Congo (©NHMUK); 20, T. admiranda, paratype ♀, DR Congo (©NHMUK); 21, T. fasciata, holotype ♀, CentralAfrican Republic (©NHMUK); 22, T. fasciata, paratype ♀, Central African Republic (©NHMUK); 23, T. smithi, holotype ♀, Zambia (ANHRT); 24, T. smithi, paratype ♀, Zambia (ANHRT); 25, T. sagenaria, ♂, South Africa (©NHMUK); 26, T. sagenaria, ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449704" httpUri="https://zenodo.org/record/3449704/files/figure.png" pageId="12" pageNumber="433">Figs 23, 24</figureCitation>
,
<figureCitation id="00B12A52FFE94263674BFB65AC7CFB0A" box="[300,329,1197,1223]" captionStart="FIGURES 4145" captionStartId="11.[151,264,1898,1923]" captionTargetBox="[189,1394,242,1857]" captionTargetId="figure@11.[189,1398,237,1874]" captionTargetPageId="11" captionText="FIGURES 4145. Tumicla spp.: female genitalia. 41, T. similis, paratype, Kenya, Kibwezi, slide NHMUK010314047 Volynkin (©NHMUK); 42, T. admiranda, paratype, DR Congo, slide NHMUK 010314600 Volynkin (©NHMUK); 43, T. fasciata, paratype, CentralAfrican Republic, slide NHMUK010314046 Volynkin (©NHMUK); 44, T. smithi, holotype, Zambia, slideAV5243 (ANHRT); 45, T. sagenaria, lectotype, [South Africa] Caffraria, slide NHRS 000011009 (photo by T. Malm, ©2017NHRS)." figureDoi="http://doi.org/10.5281/zenodo.3449718" httpUri="https://zenodo.org/record/3449718/files/figure.png" pageId="12" pageNumber="433">44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE9426366F0FB3DACF1FA52" pageId="12" pageNumber="433" type="materials_examined">
<paragraph id="983536D7FFE9426366F0FB3DA9F0FA9A" blockId="12.[151,1437,1268,2015]" pageId="12" pageNumber="433">
<emphasis id="AAFEEAC5FFE9426366F0FB3DAC74FAC2" bold="true" box="[151,321,1268,1295]" pageId="12" pageNumber="433">
<typeStatus id="47318875FFE9426366F0FB3DADE7FAC2" box="[151,210,1269,1295]" pageId="12" pageNumber="433">Type</typeStatus>
material
</emphasis>
.
<materialsCitation id="28E23C8AFFE942636734FB3DA9F7FA9A" ID-GBIF-Occurrence="2465044339" collectingDate="2019-01-27" collectingDateMax="2019-01-30" collectingDateMin="2019-01-27" collectingMethod="Light Trap" collectionCode="ANHRT" collectorName="Derozier, V. &amp; Mulvaney, L. &amp; Takano, H." country="Zambia" elevation="1456" latitude="-13.38" location="Changwena Falls, N. Swaka F. R." longLatPrecision="21" longitude="29.554998" municipality="Central Prov" pageId="12" pageNumber="433" specimenCode="ANHRTUK 00073348" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="AAFEEAC5FFE942636734FB3DACF5FAC2" bold="true" box="[339,448,1269,1295]" pageId="12" pageNumber="433">
<typeStatus id="47318875FFE942636734FB3DACF5FAC2" box="[339,448,1269,1295]" pageId="12" pageNumber="433" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="00B12A52FFE9426367B5FB3CAF19FAC2" box="[466,556,1268,1295]" captionStart="FIGURES 1926" captionStartId="5.[151,264,1899,1924]" captionTargetBox="[156,1435,181,1860]" captionTargetId="figure@5.[151,1435,181,1873]" captionTargetPageId="5" captionText="FIGURES 1926. Tumicla spp.: adults. 19, T. admiranda, holotype ♂, DR Congo (©NHMUK); 20, T. admiranda, paratype ♀, DR Congo (©NHMUK); 21, T. fasciata, holotype ♀, CentralAfrican Republic (©NHMUK); 22, T. fasciata, paratype ♀, Central African Republic (©NHMUK); 23, T. smithi, holotype ♀, Zambia (ANHRT); 24, T. smithi, paratype ♀, Zambia (ANHRT); 25, T. sagenaria, ♂, South Africa (©NHMUK); 26, T. sagenaria, ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449704" httpUri="https://zenodo.org/record/3449704/files/figure.png" pageId="12" pageNumber="433">Figs 23</figureCitation>
,
<figureCitation id="00B12A52FFE94263645BFB3DAF68FAC2" box="[572,605,1269,1295]" captionStart="FIGURES 4145" captionStartId="11.[151,264,1898,1923]" captionTargetBox="[189,1394,242,1857]" captionTargetId="figure@11.[189,1398,237,1874]" captionTargetPageId="11" captionText="FIGURES 4145. Tumicla spp.: female genitalia. 41, T. similis, paratype, Kenya, Kibwezi, slide NHMUK010314047 Volynkin (©NHMUK); 42, T. admiranda, paratype, DR Congo, slide NHMUK 010314600 Volynkin (©NHMUK); 43, T. fasciata, paratype, CentralAfrican Republic, slide NHMUK010314046 Volynkin (©NHMUK); 44, T. smithi, holotype, Zambia, slideAV5243 (ANHRT); 45, T. sagenaria, lectotype, [South Africa] Caffraria, slide NHRS 000011009 (photo by T. Malm, ©2017NHRS)." figureDoi="http://doi.org/10.5281/zenodo.3449718" httpUri="https://zenodo.org/record/3449718/files/figure.png" pageId="12" pageNumber="433">44</figureCitation>
):
<specimenCount id="8E8CFD5EFFE942636413FB3DAFBCFAC2" box="[628,649,1269,1295]" pageId="12" pageNumber="433" type="female"></specimenCount>
, “
<collectingCountry id="E09D7647FFE9426364C3FB3CAFCAFAC3" box="[676,767,1268,1294]" name="Zambia" pageId="12" pageNumber="433" value="Zambia">Zambia</collectingCountry>
,
<quantity id="5F729B32FFE942636569FB3CAE6BFAC2" box="[782,862,1268,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.456" pageId="12" pageNumber="433" unit="m" value="1456.0">
<elevation id="13A7D1E4FFE942636569FB3CAE6BFAC2" box="[782,862,1268,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.456" pageId="12" pageNumber="433" unit="m" value="1456.0">1456m</elevation>
</quantity>
,
<location id="9D55600CFFE94263650AFB3CA9DAFAC2" LSID="urn:lsid:plazi:treatment:102387C1FFE9426266F0FB40A946FD66:9D55600CFFE94263650AFB3CA9DAFAC2" box="[877,1263,1268,1295]" country="Zambia" latitude="-13.38" longLatPrecision="21" longitude="29.554998" municipality="Central Prov" name="Changwena Falls, N. Swaka F. R." pageId="12" pageNumber="433">Changwena Falls, N. Swaka F.R.</location>
,
<collectingMunicipality id="7851ACADFFE942636299FB3CA8A1FAC2" box="[1278,1428,1268,1295]" pageId="12" pageNumber="433">Central Prov</collectingMunicipality>
.,
<geoCoordinate id="FDBE5010FFE9426366F0FAD1AC17FAFE" box="[151,290,1305,1331]" degrees="13" direction="south" minutes="22" orientation="latitude" pageId="12" pageNumber="433" precision="15" seconds="48" value="-13.38">13°2248S</geoCoordinate>
,
<geoCoordinate id="FDBE5010FFE942636756FAD1AC89FAFE" box="[305,444,1305,1331]" degrees="29" direction="east" minutes="33" orientation="longitude" pageId="12" pageNumber="433" precision="15" seconds="18" value="29.554998">29°3318E</geoCoordinate>
,
<date id="EC341017FFE9426367ABFAD1AF56FAFE" box="[460,611,1305,1331]" pageId="12" pageNumber="433" value="2019-01-27" valueMax="2019-01-30" valueMin="2019-01-27">
<collectingDate id="FC70E9FFFFE9426367ABFAD1AF56FAFE" box="[460,611,1305,1331]" pageId="12" pageNumber="433" value="2019-01-27" valueMax="2019-01-30" valueMin="2019-01-27">2730.I.2019</collectingDate>
</date>
, MV
<collectingMethod id="41CB4EC0FFE9426364CFFAD0AE10FAFE" box="[680,805,1304,1331]" pageId="12" pageNumber="433">Light Trap</collectingMethod>
,
<collectorName id="357F5301FFE942636555FAD0AE8BFAFE" box="[818,958,1304,1331]" pageId="12" pageNumber="433">Dérozier, V.</collectorName>
,
<collectorName id="357F5301FFE9426365ABFAD0A953FAFE" box="[972,1126,1304,1331]" pageId="12" pageNumber="433">Mulvaney, L.</collectorName>
,
<collectorName id="357F5301FFE942636212FAD0A9C6FAFE" box="[1141,1267,1304,1331]" pageId="12" pageNumber="433">Takano, H.</collectorName>
Leg.
<collectionCode id="FE9BAE12FFE942636350FAD1A8ACFAFE" box="[1335,1433,1305,1331]" collectionName="ANHRT" pageId="12" pageNumber="433">ANHRT</collectionCode>
: 2019.4”,
<collectionCode id="FE9BAE12FFE94263669AFAF5AC6EFA9A" box="[253,347,1341,1367]" collectionName="ANHRT" pageId="12" pageNumber="433">ANHRT</collectionCode>
unique number:
<specimenCode id="C82C9EACFFE94263647FFAF5AE2CFA9A" box="[536,793,1341,1367]" pageId="12" pageNumber="433">ANHRTUK 00073348</specimenCode>
, slide No.: AV5243♀ (Coll.
<collectionCode id="FE9BAE12FFE942636231FAF5A98FFA9A" box="[1110,1210,1341,1367]" collectionName="ANHRT" pageId="12" pageNumber="433">ANHRT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="983536D7FFE9426366A0FAA9ACF1FA52" blockId="12.[151,1437,1268,2015]" pageId="12" pageNumber="433">
<materialsCitation id="28E23C8AFFE9426366A0FAA9ACF5FA52" ID-GBIF-Occurrence="2465044338" collectingDate="2019-01-27" collectingDateMax="2019-01-30" collectingDateMin="2019-01-27" collectingMethod="Light Trap" collectionCode="ANHRT" collectorName="Derozier, V. &amp; Mulvaney, L. &amp; Takano, H." country="Zambia" elevation="1456" latitude="-13.38" location="Changwena Falls, N. Swaka F. R." longLatPrecision="21" longitude="29.554998" municipality="Central Prov" pageId="12" pageNumber="433" specimenCode="ANHRTUK 00072417, 00072418, 00072419" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="AAFEEAC5FFE9426366A0FAA9AC0AFAB6" bold="true" box="[199,319,1377,1403]" pageId="12" pageNumber="433">
<typeStatus id="47318875FFE9426366A0FAA9AC0AFAB6" box="[199,319,1377,1403]" pageId="12" pageNumber="433" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="8E8CFD5EFFE94263672BFAA9AC46FAB6" box="[332,371,1377,1403]" pageId="12" pageNumber="433" type="female">3 ♀</specimenCount>
, with the same data as the holotype,
<collectionCode id="FE9BAE12FFE942636499FAA9AE69FAB6" box="[766,860,1377,1403]" collectionName="ANHRT" pageId="12" pageNumber="433">ANHRT</collectionCode>
unique numbers:
<specimenCode id="C82C9EACFFE94263627AFAA9AC32FA52" pageId="12" pageNumber="433">ANHRTUK 00072417, 00072418, 00072419</specimenCode>
(Coll.
<collectionCode id="FE9BAE12FFE942636733FA4DAC8DFA52" box="[340,440,1413,1439]" collectionName="ANHRT" pageId="12" pageNumber="433">ANHRT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE9426366A0FA61AF5BF9E2" pageId="12" pageNumber="433" type="discussion">
<paragraph id="983536D7FFE9426366A0FA61AF5BF9E2" blockId="12.[151,1437,1268,2015]" pageId="12" pageNumber="433">
<emphasis id="AAFEEAC5FFE9426366A0FA61AC05FA0E" bold="true" box="[199,304,1449,1475]" pageId="12" pageNumber="433">Remark.</emphasis>
<taxonomicName id="5F8A4D54FFE942636751FA60ACEFFA0F" authorityName="Volynkin &amp; László" authorityYear="2019" box="[310,474,1448,1474]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="AAFEEAC5FFE942636751FA60ACEFFA0F" box="[310,474,1448,1474]" italics="true" pageId="12" pageNumber="433">Tumicla smithi</emphasis>
</taxonomicName>
is to date, only known by females. However, despite its unusual wing pattern,
<taxonomicName id="5F8A4D54FFE94263635AFA61A8A9FA0F" authorityName="Volynkin &amp; László" authorityYear="2019" box="[1341,1436,1448,1474]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="AAFEEAC5FFE94263635AFA61A8A9FA0F" box="[1341,1436,1448,1474]" italics="true" pageId="12" pageNumber="433">T. smithi</emphasis>
</taxonomicName>
certainly belongs to the genus
<taxonomicName id="5F8A4D54FFE94263678CFA04AF70FA2B" authorityName="Wallengren" authorityYear="1863" box="[491,581,1484,1510]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="AAFEEAC5FFE94263678CFA04AF70FA2B" box="[491,581,1484,1510]" italics="true" pageId="12" pageNumber="433">Tumicla</emphasis>
</taxonomicName>
due to its legs tibial spur formula 0-2-2 characteristic for all
<taxonomicName id="5F8A4D54FFE94263628CFA04A870FA2B" authorityName="Wallengren" authorityYear="1863" box="[1259,1349,1484,1510]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="AAFEEAC5FFE94263628CFA04A870FA2B" box="[1259,1349,1484,1510]" italics="true" pageId="12" pageNumber="433">Tumicla</emphasis>
</taxonomicName>
species (
<bibRefCitation id="FC1B4B26FFE9426366F8FA39AC0AF9C6" author="Durante, A." box="[159,319,1521,1547]" pageId="12" pageNumber="433" pagination="53 - 68" refId="ref6776" refString="Durante, A. (2008) Asuroides, a new genus of lithosiine moths (Lepidoptera, Arctiidae, Lithosiinae). Zootaxa, 1713, 53 - 68." type="journal article" year="2008">Durante 2008</bibRefCitation>
), forewing shape, and female genitalia being similar to those of some other species of
<taxonomicName id="5F8A4D54FFE94263637AFA38A842F9C7" authorityName="Wallengren" authorityYear="1863" box="[1309,1399,1520,1546]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="AAFEEAC5FFE94263637AFA38A842F9C7" box="[1309,1399,1520,1546]" italics="true" pageId="12" pageNumber="433">Tumicla</emphasis>
</taxonomicName>
, in particular to
<taxonomicName id="5F8A4D54FFE942636741F9DDACAEF9E2" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[294,411,1557,1583]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="AAFEEAC5FFE942636741F9DDACAEF9E2" box="[294,411,1557,1583]" italics="true" pageId="12" pageNumber="433">T. fasciata</emphasis>
</taxonomicName>
and
<taxonomicName id="5F8A4D54FFE9426367B6F9DDAF52F9E3" authorityName="Volynkin &amp; László" authorityYear="2019" box="[465,615,1556,1582]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="admiranda">
<emphasis id="AAFEEAC5FFE9426367B6F9DDAF52F9E3" box="[465,615,1556,1582]" italics="true" pageId="12" pageNumber="433">T. admiranda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE9426366A0F9F0AC35F882" pageId="12" pageNumber="433" type="diagnosis">
<paragraph id="983536D7FFE9426366A0F9F0AC35F882" blockId="12.[151,1437,1268,2015]" pageId="12" pageNumber="433">
<emphasis id="AAFEEAC5FFE9426366A0F9F0AC0FF99E" bold="true" box="[199,314,1592,1619]" pageId="12" pageNumber="433">Diagnosis</emphasis>
.
<taxonomicName id="5F8A4D54FFE94263672FF9F0ACDBF99F" authorityName="Volynkin &amp; László" authorityYear="2019" box="[328,494,1592,1618]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="AAFEEAC5FFE94263672FF9F0ACDBF99F" box="[328,494,1592,1618]" italics="true" pageId="12" pageNumber="433">Tumicla smithi</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE94263679AF9F0AF48F99E" box="[509,637,1592,1619]" captionStart="FIGURES 1926" captionStartId="5.[151,264,1899,1924]" captionTargetBox="[156,1435,181,1860]" captionTargetId="figure@5.[151,1435,181,1873]" captionTargetPageId="5" captionText="FIGURES 1926. Tumicla spp.: adults. 19, T. admiranda, holotype ♂, DR Congo (©NHMUK); 20, T. admiranda, paratype ♀, DR Congo (©NHMUK); 21, T. fasciata, holotype ♀, CentralAfrican Republic (©NHMUK); 22, T. fasciata, paratype ♀, Central African Republic (©NHMUK); 23, T. smithi, holotype ♀, Zambia (ANHRT); 24, T. smithi, paratype ♀, Zambia (ANHRT); 25, T. sagenaria, ♂, South Africa (©NHMUK); 26, T. sagenaria, ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449704" httpUri="https://zenodo.org/record/3449704/files/figure.png" pageId="12" pageNumber="433">Figs 23, 24</figureCitation>
) is also an unmistakable species differing clearly from other congeners by its orange forewing lacking transverse lines and having a broad unicolorous black distal band. The female genitalia of
<taxonomicName id="5F8A4D54FFE94263668BF949AC7EF957" authorityName="Volynkin &amp; László" authorityYear="2019" box="[236,331,1664,1690]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="AAFEEAC5FFE94263668BF949AC7EF957" box="[236,331,1664,1690]" italics="true" pageId="12" pageNumber="433">T. smithi</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE94263673DF948AC9EF956" box="[346,427,1664,1691]" captionStart="FIGURES 4145" captionStartId="11.[151,264,1898,1923]" captionTargetBox="[189,1394,242,1857]" captionTargetId="figure@11.[189,1398,237,1874]" captionTargetPageId="11" captionText="FIGURES 4145. Tumicla spp.: female genitalia. 41, T. similis, paratype, Kenya, Kibwezi, slide NHMUK010314047 Volynkin (©NHMUK); 42, T. admiranda, paratype, DR Congo, slide NHMUK 010314600 Volynkin (©NHMUK); 43, T. fasciata, paratype, CentralAfrican Republic, slide NHMUK010314046 Volynkin (©NHMUK); 44, T. smithi, holotype, Zambia, slideAV5243 (ANHRT); 45, T. sagenaria, lectotype, [South Africa] Caffraria, slide NHRS 000011009 (photo by T. Malm, ©2017NHRS)." figureDoi="http://doi.org/10.5281/zenodo.3449718" httpUri="https://zenodo.org/record/3449718/files/figure.png" pageId="12" pageNumber="433">Fig. 44</figureCitation>
) are most similar to those of
<taxonomicName id="5F8A4D54FFE942636497F949AE51F956" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[752,868,1665,1691]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="AAFEEAC5FFE942636497F949AE51F956" box="[752,868,1665,1691]" italics="true" pageId="12" pageNumber="433">T. fasciata</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE942636515F948AEF0F956" box="[882,965,1664,1691]" captionStart="FIGURES 4145" captionStartId="11.[151,264,1898,1923]" captionTargetBox="[189,1394,242,1857]" captionTargetId="figure@11.[189,1398,237,1874]" captionTargetPageId="11" captionText="FIGURES 4145. Tumicla spp.: female genitalia. 41, T. similis, paratype, Kenya, Kibwezi, slide NHMUK010314047 Volynkin (©NHMUK); 42, T. admiranda, paratype, DR Congo, slide NHMUK 010314600 Volynkin (©NHMUK); 43, T. fasciata, paratype, CentralAfrican Republic, slide NHMUK010314046 Volynkin (©NHMUK); 44, T. smithi, holotype, Zambia, slideAV5243 (ANHRT); 45, T. sagenaria, lectotype, [South Africa] Caffraria, slide NHRS 000011009 (photo by T. Malm, ©2017NHRS)." figureDoi="http://doi.org/10.5281/zenodo.3449718" httpUri="https://zenodo.org/record/3449718/files/figure.png" pageId="12" pageNumber="433">Fig. 43</figureCitation>
), but differ from them by their shorter and less curved ductus bursae without a lateral protrusion (that of
<taxonomicName id="5F8A4D54FFE94263653AF96DAEE1F972" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[861,980,1701,1727]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="AAFEEAC5FFE94263653AF96DAEE1F972" box="[861,980,1701,1727]" italics="true" pageId="12" pageNumber="433">T. fasciata</emphasis>
</taxonomicName>
bears a well-developed, rounded lateral bulge), more heavily sclerotised posterior section of corpus bursae lacking a lateral protrusion (that is less strongly sclerotised, rugose having a long and broad lateral protrusion in
<taxonomicName id="5F8A4D54FFE94263651EF925AEDAF8CA" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[889,1007,1773,1799]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="AAFEEAC5FFE94263651EF925AEDAF8CA" box="[889,1007,1773,1799]" italics="true" pageId="12" pageNumber="433">T. fasciata</emphasis>
</taxonomicName>
), and shorter appendix bursae having sclerotised basal section and membranous distal section with granulation (that is larger and evenly sclerotised in
<taxonomicName id="5F8A4D54FFE9426363EFF8D9ADC5F882" baseAuthorityName="Durante" baseAuthorityYear="2008" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="AAFEEAC5FFE9426363EFF8D9ADC5F882" italics="true" pageId="12" pageNumber="433">T. fasciata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE9426266A0F890AC7DFDD6" lastPageId="13" lastPageNumber="434" pageId="12" pageNumber="433" type="description">
<paragraph id="983536D7FFE9426266A0F890ACBFFE3A" blockId="12.[151,1437,1268,2015]" lastBlockId="13.[151,1437,153,683]" lastPageId="13" lastPageNumber="434" pageId="12" pageNumber="433">
<emphasis id="AAFEEAC5FFE9426366A0F890AC64F8BE" bold="true" box="[199,337,1880,1907]" pageId="12" pageNumber="433">Description</emphasis>
.
<emphasis id="AAFEEAC5FFE942636706F891AFE0F8BE" bold="true" box="[353,725,1880,1907]" pageId="12" pageNumber="433">External morphology of adults</emphasis>
(
<figureCitation id="00B12A52FFE942636481F890AE59F8BE" box="[742,876,1880,1907]" captionStart="FIGURES 1926" captionStartId="5.[151,264,1899,1924]" captionTargetBox="[156,1435,181,1860]" captionTargetId="figure@5.[151,1435,181,1873]" captionTargetPageId="5" captionText="FIGURES 1926. Tumicla spp.: adults. 19, T. admiranda, holotype ♂, DR Congo (©NHMUK); 20, T. admiranda, paratype ♀, DR Congo (©NHMUK); 21, T. fasciata, holotype ♀, CentralAfrican Republic (©NHMUK); 22, T. fasciata, paratype ♀, Central African Republic (©NHMUK); 23, T. smithi, holotype ♀, Zambia (ANHRT); 24, T. smithi, paratype ♀, Zambia (ANHRT); 25, T. sagenaria, ♂, South Africa (©NHMUK); 26, T. sagenaria, ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449704" httpUri="https://zenodo.org/record/3449704/files/figure.png" pageId="12" pageNumber="433">Figs 23, 24</figureCitation>
). Forewing length 14.5
<quantity id="5F729B32FFE9426362E2F890A9E1F8BE" box="[1157,1236,1880,1907]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="433" unit="mm" value="15.0">15 mm</quantity>
in females. Head black; antenna black, filiform in female. Thorax black; patagium orange; tegula orange with large black basal spot. Forewing moderately broad, with rounded apex, outer margin and tornus evenly arcuate. Forewing ground colour bright orange; pattern black, represented by a short basal dash on costa, short V-like subbasal streak, narrow semilu- nar discal spot which may be absent, and broad black terminal area with S-curved inner margin filling in postmedial, subterminal and terminal areas unicolorously; cilia black. Hindwing paler than forewing, orange, with intense black suffusion on subterminal and terminal areas, varying widely in width; cilia black, pale orange along anal margin. Abdomen orange with admixture of brown scales.
<emphasis id="AAFEEAC5FFE8426264BFFECDAEAAFED2" bold="true" box="[728,927,260,287]" pageId="13" pageNumber="434">Female genitalia</emphasis>
(
<figureCitation id="00B12A52FFE8426265C8FECDA930FED2" box="[943,1029,261,287]" captionStart="FIGURES 4145" captionStartId="11.[151,264,1898,1923]" captionTargetBox="[189,1394,242,1857]" captionTargetId="figure@11.[189,1398,237,1874]" captionTargetPageId="11" captionText="FIGURES 4145. Tumicla spp.: female genitalia. 41, T. similis, paratype, Kenya, Kibwezi, slide NHMUK010314047 Volynkin (©NHMUK); 42, T. admiranda, paratype, DR Congo, slide NHMUK 010314600 Volynkin (©NHMUK); 43, T. fasciata, paratype, CentralAfrican Republic, slide NHMUK010314046 Volynkin (©NHMUK); 44, T. smithi, holotype, Zambia, slideAV5243 (ANHRT); 45, T. sagenaria, lectotype, [South Africa] Caffraria, slide NHRS 000011009 (photo by T. Malm, ©2017NHRS)." figureDoi="http://doi.org/10.5281/zenodo.3449718" httpUri="https://zenodo.org/record/3449718/files/figure.png" pageId="13" pageNumber="434">Fig. 44</figureCitation>
). Ovipositor short and broad; papillae anales rectangular, with rounded angles; apophyses moderately long and thin, equal in length. Ostium bursae relatively narrow; ductus bursae moderately long, dorso-ventrally flattened, heavily sclerotised, slightly broadened posteriorly, gently curved to the left, with convex right margin. Posterior section of corpus bursae conically tapered posteriorly, sclerotised; anterior section of corpus bursae globular, moderately sclerotised, rugose. Appendix bursae short, conical, sclerotised basally and membranous with granulation distally, situated postero-laterally on the left side of ductus bursae.
</paragraph>
<paragraph id="983536D7FFE8426266F0FDC9AC7DFDD6" blockId="13.[151,1437,153,683]" box="[151,328,513,539]" pageId="13" pageNumber="434">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE8426266A0FDECAC7BFDAE" pageId="13" pageNumber="434" type="distribution">
<paragraph id="983536D7FFE8426266A0FDECAC7BFDAE" blockId="13.[151,1437,153,683]" pageId="13" pageNumber="434">
<emphasis id="AAFEEAC5FFE8426266A0FDECAC6CFDF2" bold="true" box="[199,345,548,575]" pageId="13" pageNumber="434">Distribution</emphasis>
.
<taxonomicName id="5F8A4D54FFE84262670EFDEDAF3AFDF2" authorityName="Volynkin &amp; László" authorityYear="2019" box="[361,527,549,575]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="434" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="AAFEEAC5FFE84262670EFDEDAF3AFDF2" box="[361,527,549,575]" italics="true" pageId="13" pageNumber="434">Tumicla smithi</emphasis>
</taxonomicName>
is to date, only known from its
<typeStatus id="47318875FFE8426265E7FDEDAE85FDF2" box="[896,944,549,575]" pageId="13" pageNumber="434">type</typeStatus>
locality collected near Changwena Falls in Central Zambia.
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE8426266A0FDA5A946FD66" pageId="13" pageNumber="434" type="etymology">
<paragraph id="983536D7FFE8426266A0FDA5A946FD66" blockId="13.[151,1437,153,683]" pageId="13" pageNumber="434">
<emphasis id="AAFEEAC5FFE8426266A0FDA5AC7DFD4A" bold="true" box="[199,328,621,647]" pageId="13" pageNumber="434">Etymology</emphasis>
. The species is dedicated to Mr Richard Smith, founder of the African Natural History Research Trust, organiser of extensive entomological exploratory program in Sub-Saharan Africa.
</paragraph>
</subSubSection>
</treatment>
</document>