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<mods:title id="7DCAC30B45EDF57370077828B70DBF92">Early eusuchia crocodylomorpha from the vertebrate-rich Plattenkalk of Pietraroia (Lower Albian, southern Apennines, Italy)</mods:title>
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<mods:namePart id="4B87764FB6A8158A6E83B1B619E9D3FC">Buscalioni, Angela D.</mods:namePart>
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<mods:namePart id="D025FE9E2FDE2B5C5D57CBE3CA331010">Piras, Paolo</mods:namePart>
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<mods:namePart id="991AD56F259076B4996101EEC2E3AD53">Signore, Marco</mods:namePart>
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<heading id="36088F4453302F03FCD5DDED730BAF9B" box="[894,1334,1458,1479]" centered="true" fontSize="8" level="2" pageId="4" reason="8">
GENUS
<taxonomicName id="AAFF43AB53302F03FC7CDDED7285AF9A" authority="Buscalioni &amp; Piras &amp; Vullo &amp; Signore &amp; Barbera, 2011" authorityName="Buscalioni &amp; Piras &amp; Vullo &amp; Signore &amp; Barbera" authorityYear="2011" box="[983,1208,1458,1479]" class="Reptilia" family="Hylaeochampsidae" genus="Pietraroiasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" phylum="Chordata" rank="genus" status="gen. nov.">PIETRAROIASUCHUS</taxonomicName>
<emphasis id="5F8BE43A53302F03FB69DDE9730BAF9B" box="[1218,1334,1458,1479]" italics="true" pageId="4">
<taxonomicNameLabel id="44B8594153302F03FB69DDE9730BAF9B" box="[1218,1334,1458,1479]" pageId="4" rank="genus">GEN. NOV.</taxonomicNameLabel>
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<paragraph id="6D40382853302F03FC8DDD867330AFB2" blockId="4.[806,1293,1496,1518]" box="[806,1293,1496,1518]" pageId="4">
<emphasis id="5F8BE43A53302F03FC8DDD867330AFB2" box="[806,1293,1496,1518]" italics="true" pageId="4">
<typeStatus id="B244868A53302F03FC8DDD867567AFB2" box="[806,858,1497,1518]" pageId="4">Type</typeStatus>
species:
<taxonomicName id="AAFF43AB53302F03FC69DD877330AFB2" authorityName="Buscalioni &amp; Piras &amp; Vullo &amp; Signore &amp; Barbera" authorityYear="2011" box="[962,1293,1496,1518]" class="Reptilia" family="Hylaeochampsidae" genus="Pietraroiasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" phylum="Chordata" rank="species" species="ormezzanoi">Pietraroiasuchus ormezzanoi</taxonomicName>
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<emphasis id="5F8BE43A53302F03FC8DDE78720BAC60" box="[806,1078,1575,1596]" italics="true" pageId="4">Diagnosis of the genus:</emphasis>
Small crocodylomorph about
<quantity id="AA0795CD53302F03FC8DDE19756CAC07" box="[806,849,1606,1627]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" unit="m" value="1.0">1 m</quantity>
long, with flat skull (rostrum and mandible) and round rostral contour. Rostral profile convex in front of orbits; orbits with anterior rectilinear contour; premaxillary constituting about one-third of rostral length; maxillary alveoli medially placed and laterally bordered by a ridge (shared with
<taxonomicName id="AAFF43AB53302F03FB01DE817343ACAF" baseAuthorityName="Rogers" baseAuthorityYear="2003" box="[1194,1406,1758,1779]" class="Reptilia" family="Hylaeochampsidae" genus="Pachycheilosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" phylum="Chordata" rank="genus">
<emphasis id="5F8BE43A53302F03FB01DE817343ACAF" box="[1194,1406,1758,1779]" italics="true" pageId="4">Pachycheilosuchus</emphasis>
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); three final maxillary teeth medially arranged in a platform (autapomorphy); maxillary bone with minimal sculpture and pitting aligned along lateral margins; maxillae send a short caudal process between the lachrymal and prefrontal, thus avoiding lachrymonasal contact; maxillary tooth row ends well anterior to posterior tip of maxilla; posteromedial edge of maxillae forming the margin of a depression or fossa in front of the maxillojugal suture; prefrontal and lachrymal about same length; preorbital fenestra slit-like; choana in pterygoid, anteriorly placed, and bordered by palatines; compound quadrate quadratojugal condyles (autapomorphy); furrowed quadratojugal and quadrate suture (autapomorphy); pterygoidean flange not reaching laterally the medial quadrate condyle (shared with
<taxonomicName id="AAFF43AB53312F02FDE1DE3B74C1AC25" baseAuthorityName="Osi" baseAuthorityYear="2008" box="[586,764,1636,1657]" class="Reptilia" family="Hylaeochampsidae" genus="Iharkutosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" phylum="Chordata" rank="genus">
<emphasis id="5F8BE43A53312F02FDE1DE3B74C1AC25" box="[586,764,1636,1657]" italics="true" pageId="5">Iharkutosuchus</emphasis>
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); lateroventral protuberance in exoccipitals form a vertical ridge (shared with
<taxonomicName id="AAFF43AB53312F02FE6ADEFE7448ACEA" authorityName="Owen" authorityYear="1874" box="[449,629,1697,1718]" class="Reptilia" family="Hylaeochampsidae" genus="Hylaeochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" phylum="Chordata" rank="genus">
<emphasis id="5F8BE43A53312F02FE6ADEFE7448ACEA" box="[449,629,1697,1718]" italics="true" pageId="5">Hylaeochampsa</emphasis>
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and
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<emphasis id="5F8BE43A53312F02FD1FDEFE773AAC89" italics="true" pageId="5">Iharkutosuchus</emphasis>
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); unforked and unique posterior dentary process (shared with
<taxonomicName id="AAFF43AB53312F02FE68DE8174AAACAF" baseAuthorityName="Rogers" baseAuthorityYear="2003" box="[451,663,1758,1779]" class="Reptilia" family="Hylaeochampsidae" genus="Pachycheilosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" phylum="Chordata" rank="genus">
<emphasis id="5F8BE43A53312F02FE68DE8174AAACAF" box="[451,663,1758,1779]" italics="true" pageId="5">Pachycheilosuchus</emphasis>
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); medial extension of dentary; surangular anterodorsally expanded; posterodorsal smooth dentary projection lacking alveolar setting; homodont dentition; absence of external mandibular fenestra. Postcranial skeleton is characterized by slightly procoelous centra, being more pronounced in cervicals; anteroposteriorly, expansions of transverse processes in thoracic area; tail longer than the combined length of skull and presacrum; forelimb more slender and shorter than hindlimbs, with stout femora; humerus with straight shaft and distal condyles of similar transverse extension; long anterior dorsal end of coracoid twice the length of its ventral edge; nuchal shield separated from dorsal shield and composed of a compound set of six profusely ornamented osteoderms (two parasagittal rows and one lateral) (shared with
<taxonomicName id="AAFF43AB53312F02FB53DE3B7553ACC4" baseAuthorityName="Rogers" baseAuthorityYear="2003" class="Reptilia" family="Hylaeochampsidae" genus="Pachycheilosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" phylum="Chordata" rank="genus">
<emphasis id="5F8BE43A53312F02FB53DE3B7553ACC4" italics="true" pageId="5">Pachycheilosuchus</emphasis>
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); dorsal dermal skeleton composed of tetraserial paravertebral shield and one accessory osteoderm.
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