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<document id="70B69350CDC4D2ABF0094CB516899473" ID-DOI="10.5852/ejt.2024.923.2445" ID-ISSN="2118-9773" ID-Zenodo-Dep="10727741" ID-ZooBank="AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1709215085450" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2024" docId="E54587AB38650B20FDA4FAAF53FFF967" docLanguage="en" docName="EJT.2024.923.1-119.pdf.imf" docOrigin="European Journal of Taxonomy 923" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Benthobrookula scalaroides Herbert 2024, sp. nov." docType="treatment" docUuid="8261B816-4DC9-4D46-A58F-D4F056DE1951" docUuidSource="ZooBank" docVersion="5" lastPageNumber="61" masterDocId="197CFFD3385E0B1CFF9BFFC25776FFD3" masterDocTitle="The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species" masterLastPageNumber="119" masterPageNumber="1" pageNumber="60" updateTime="1710426596983" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="10BC94DBA667E9292229A1D6E1E6382D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F18E08B8CDA688C9F1582B327E76AFC">
<mods:title id="2FCDE33AFA6F22FC136E0D4A2BE5405C">The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species</mods:title>
</mods:titleInfo>
<mods:name id="7E34D2FF14878158817406436B04B23C" type="personal">
<mods:role id="8A1AA7B4910F9B56235B880A8675463A">
<mods:roleTerm id="2156A78DE7578906E9F307ED60BB5F03">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1900E7BAD3A1A928C6C9AB40C2D1A5B">Herbert, David G.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C22FA67A3EE324542652654F0AFA7174">text</mods:typeOfResource>
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<mods:titleInfo id="4D1B894480CD69A53E7372283D4EF09B">
<mods:title id="3BA4C1AA490C9FFB3A4EFC09574E00BC">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="8E6C4731265F41AF20D15F5C66523716">
<mods:date id="D295DD5AD5527D7C0861D46609BEC434">2024</mods:date>
<mods:detail id="E9B85D8E07C88CF51539A4E8D300FDD4" type="pubDate">
<mods:number id="20341FA46321D2C76DC795399848791B">2024-02-28</mods:number>
</mods:detail>
<mods:detail id="C431E8A13C09EB228EC113CDACEBD1E6" type="volume">
<mods:number id="3B8686E04671AF9A6E594117DC0B9C49">923</mods:number>
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<mods:start id="5D7ECE90A5432435FB09A1CBEE2DDE23">1</mods:start>
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<mods:url id="2BDCC56721BB48178BBF0486062B8325">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835</mods:url>
</mods:location>
<mods:classification id="6CC870EC530B604C75DFF4856390A041">journal article</mods:classification>
<mods:identifier id="BD3E9E8FA3C94876BB2B8B446803EDEC" type="DOI">10.5852/ejt.2024.923.2445</mods:identifier>
<mods:identifier id="026563C32ECFF17787EA0391AA873096" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="4A449B2F791073D8C8D49D1F4DB79E83" type="Zenodo-Dep">10727741</mods:identifier>
<mods:identifier id="124D6B9598264BF8DEBAD41B9EDF6B66" type="ZooBank">AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3</mods:identifier>
</mods:mods>
<treatment id="E54587AB38650B20FDA4FAAF53FFF967" LSID="urn:lsid:zoobank.org:act:8261B816-4DC9-4D46-A58F-D4F056DE1951" httpUri="http://treatment.plazi.org/id/E54587AB38650B20FDA4FAAF53FFF967" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<subSubSection id="25F6653638650B27FDA4FAAF5483FA5B" box="[575,1013,1389,1416]" pageId="59" pageNumber="60" type="nomenclature">
<paragraph id="6D5336BD38650B27FDA4FAAF5483FA5B" blockId="59.[368,1219,1389,1485]" box="[575,1013,1389,1416]" pageId="59" pageNumber="60">
<heading id="361B81D138650B27FDA4FAAF5483FA5B" box="[575,1013,1389,1416]" centered="true" fontSize="11" level="2" pageId="59" pageNumber="60" reason="2">
<taxonomicName id="AAEC4D3E38650B27FDA4FAAF54E5FA5B" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[575,915,1389,1416]" class="Gastropoda" family="Unassigned" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="scalaroides" status="sp. nov." superFamily="Seguenzioidea">
<emphasis id="5F98EAAF38650B27FDA4FAAF54E5FA5B" bold="true" box="[575,915,1389,1416]" italics="true" pageId="59" pageNumber="60">Benthobrookula scalaroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438650B27FC01FAAC5483FA5B" box="[922,1013,1390,1416]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25F6653638650B27FEEBFA5254FEFA1E" pageId="59" pageNumber="60" type="description">
<paragraph id="6D5336BD38650B27FEEBFA5253B5FA78" blockId="59.[368,1219,1389,1485]" box="[368,1219,1424,1451]" pageId="59" pageNumber="60">
<uri id="197D3ABF38650B27FEEBFA5253B5FA78" box="[368,1219,1424,1451]" pageId="59" pageNumber="60">
urn:lsid:zoobank.org:act:
<uuid id="194A0C6838650B27FD0BFA5253B5FA78" box="[656,1219,1424,1451]" pageId="59" pageNumber="60">8261B816-4DC9-4D46-A58F-D4F056DE1951</uuid>
</uri>
</paragraph>
<paragraph id="6D5336BD38650B27FD30FA7054FEFA1E" blockId="59.[368,1219,1389,1485]" box="[683,904,1458,1485]" pageId="59" pageNumber="60">
<figureCitation id="F5D72A3838650B27FD30FA705444FA1F" box="[683,818,1458,1484]" captionStart="Fig" captionStartId="55.[189,232,1751,1777]" captionTargetBox="[189,1399,265,1717]" captionTargetId="figure-114@55.[189,1399,265,1717]" captionTargetPageId="55" captionText="Fig. 24. A–C. Benthobrookula araneum sp. nov. A–B. Holotype, height 1.2 mm (MNHN-IM-2000- 38068). C. Paratype, diameter 1.1 mm (MNHN-IM-2000-38069). D–F. Benthobrookula galeneae sp. nov. D–E. Holotype, height 1.25 mm (MNHN-IM-2000-38075). F. Paratype, diameter 1.2 mm (MNHN-IM-2000-38076). G–I. Benthobrookula laticostata sp. nov. G–H. Holotype, diameter 1.0 mm (MNHN-IM-2000-38080).I.Paratype,diameter1.0mm(MNHN-IM-2000-38082).J–L.Benthobrookula scalaroides sp. nov. J–K. Holotype, height 1.14 mm (MNHN-IM-2000-38084). L. Paratype, diameter 1.0 mm (MNHN-IM-2000-38085). M–O. Benthobrookula semisculpta sp. nov. M–N. Holotype, height 1.4 mm (MNHN-IM-2000-38088). O. Paratype, diameter 1.4 mm (MNHN-IM-2000-38090)." figureDoi="http://doi.org/10.5281/zenodo.10727808" httpUri="https://zenodo.org/record/10727808/files/figure.png" pageId="59" pageNumber="60">Figs 24J–L</figureCitation>
,
<figureCitation id="F5D72A3838650B27FCA6FA7054FEFA1E" box="[829,904,1458,1485]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="59" pageNumber="60">25D, I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="25F6653638650B27FF26FA3155D3F915" pageId="59" pageNumber="60" type="diagnosis">
<paragraph id="6D5336BD38650B27FF26FA315640F9DE" blockId="59.[189,310,1523,1549]" box="[189,310,1523,1549]" pageId="59" pageNumber="60">
<heading id="361B81D138650B27FF26FA315640F9DE" bold="true" box="[189,310,1523,1549]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="5F98EAAF38650B27FF26FA315640F9DE" bold="true" box="[189,310,1523,1549]" pageId="59" pageNumber="60">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38650B27FF26F9E355D3F915" blockId="59.[189,1399,1569,1734]" pageId="59" pageNumber="60">Shell elevated-turbiniform, H/D =1.1–1.2, whorls evenly rounded, suture deeply indented, base umbilicate; teleoconch sculptured with widely-spaced, sharp, orthocline, axial ribs; first whorl with ± 17 ribs, penultimate whorl with ± 20 ribs; ribs sometimes more close-set in last quarter whorl; rib intervals with microscopic spiral threads; basal sculpture similar, ribs continuing into umbilicus; peristome complete; protoconch globose and exsert.</paragraph>
</subSubSection>
<subSubSection id="25F6653638650B27FF26F92E53A2F8E7" box="[189,1236,1772,1844]" pageId="59" pageNumber="60" type="etymology">
<paragraph id="6D5336BD38650B27FF26F92E5632F8D5" blockId="59.[189,324,1772,1798]" box="[189,324,1772,1798]" pageId="59" pageNumber="60">
<heading id="361B81D138650B27FF26F92E5632F8D5" bold="true" box="[189,324,1772,1798]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="5F98EAAF38650B27FF26F92E5632F8D5" bold="true" box="[189,324,1772,1798]" pageId="59" pageNumber="60">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38650B27FF26F8D853A2F8E7" blockId="59.[189,1236,1818,1844]" box="[189,1236,1818,1844]" pageId="59" pageNumber="60">
From the Latin “
<emphasis id="5F98EAAF38650B27FE1AF8D856C9F8E7" box="[385,447,1818,1844]" italics="true" pageId="59" pageNumber="60">scala</emphasis>
” – “a flight of stairs”; in reference to the strongly ribbed sculpture.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638650B20FF26F8985558FE62" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="6D5336BD38650B27FF26F89856DCF8A7" blockId="59.[189,426,1882,1908]" box="[189,426,1882,1908]" pageId="59" pageNumber="60">
<heading id="361B81D138650B27FF26F89856DCF8A7" bold="true" box="[189,426,1882,1908]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="5F98EAAF38650B27FF26F89856DCF8A7" bold="true" box="[189,426,1882,1908]" pageId="59" pageNumber="60">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="DD843CE038650B27FF71F84A5558F83B" collectingDate="2017-05-03" elevation="578" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.933334" pageId="59" pageNumber="60" specimenCode="MNHN-IM-2000-38084" specimenCount="1" typeStatus="holotype">
<paragraph id="6D5336BD38650B27FF71F84A562AF871" blockId="59.[189,1399,1928,2024]" box="[234,348,1928,1954]" pageId="59" pageNumber="60">
<emphasis id="5F98EAAF38650B27FF71F84A562AF871" bold="true" box="[234,348,1928,1954]" pageId="59" pageNumber="60">
<typeStatus id="B257881F38650B27FF71F84A562AF871" box="[234,348,1928,1954]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD38650B27FF26F8695558F83B" blockId="59.[189,1399,1928,2024]" pageId="59" pageNumber="60">
<location id="6833606638650B27FF26F86956DAF816" LSID="urn:lsid:plazi:treatment:E54587AB38650B20FDA4FAAF53FFF967:6833606638650B27FF26F86956DAF816" box="[189,428,1963,1989]" latitude="-33.283333" longLatPrecision="1205" longitude="43.933334" name="WALTERS SHOAL" pageId="59" pageNumber="60">WALTERS SHOAL</location>
• empty shell; slopes, stn DW4886;
<geoCoordinate id="08D8507A38650B27FCC5F86B54B3F816" box="[862,965,1961,1989]" degrees="33" direction="south" minutes="17" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="-33.283333">33°17′ S</geoCoordinate>
,
<geoCoordinate id="08D8507A38650B27FC48F86B534BF816" box="[979,1085,1961,1989]" degrees="43" direction="east" minutes="56" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="43.933334">43°56′ E</geoCoordinate>
; depth
<quantity id="AA149B5838650B27FB0FF869526BF816" box="[1172,1309,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="5.775" metricValueMax="5.82" metricValueMin="5.7299999999999995" pageId="59" pageNumber="60" unit="m" value="577.5" valueMax="582.0" valueMin="573.0">
<elevation id="E6C1D18E38650B27FB0FF869526BF816" box="[1172,1309,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="5.775" metricValueMax="5.82" metricValueMin="5.7299999999999995" pageId="59" pageNumber="60" unit="m" value="577.5" valueMax="582.0" valueMin="573.0">573–582 m</elevation>
</quantity>
;
<date id="1952107D38650B27FAB1F869578DF83B" pageId="59" pageNumber="60" value="2017-05-03">
<collectingDate id="0916E99538650B27FAB1F869578DF83B" pageId="59" pageNumber="60" value="2017-05-03">3 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638650B27FE9CF80C555FF83B" box="[263,553,1998,2024]" pageId="59" pageNumber="60">MNHN-IM-2000-38084</specimenCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="DD843CE038620B20FF71FECB560EFEB9" collectingDate="2017-05-03" collectionCode="MNHN" elevation="578" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.933334" pageId="60" pageNumber="61" specimenCode="MNHN- IM-2000-38085" specimenCount="16" typeStatus="paratype">
<paragraph id="6D5336BD38620B20FF71FECB5611FEF0" blockId="60.[189,1399,265,433]" box="[234,359,265,291]" pageId="60" pageNumber="61">
<emphasis id="5F98EAAF38620B20FF71FECB5611FEF0" bold="true" box="[234,359,265,291]" pageId="60" pageNumber="61">
<typeStatus id="B257881F38620B20FF71FECB5611FEF0" box="[234,359,265,291]" pageId="60" pageNumber="61" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD38620B20FF26FEEE5558FE62" blockId="60.[189,1399,265,433]" pageId="60" pageNumber="61">
<location id="6833606638620B20FF26FEEE56C7FE95" LSID="urn:lsid:plazi:treatment:E54587AB38650B20FDA4FAAF53FFF967:6833606638620B20FF26FEEE56C7FE95" box="[189,433,300,326]" name="WALTERS SHOAL" pageId="60" pageNumber="61">WALTERS SHOAL</location>
–
<emphasis id="5F98EAAF38620B20FE45FEEE555FFE95" bold="true" box="[478,553,300,326]" pageId="60" pageNumber="61">slopes</emphasis>
•
<specimenCount id="7BEAFD3438620B20FDCAFEEE5518FE95" box="[593,622,300,326]" pageId="60" pageNumber="61">16</specimenCount>
empty shells; same collection data as for holotype;
<specimenCode id="3D4A9EC638620B20FA88FEEF560EFEB9" pageId="60" pageNumber="61">MNHN- IM-2000-38085</specimenCode>
•
<materialsCitation id="DD843CE038620B20FE0CFE9256ADFE5E" collectingDate="2017-05-03" collectionCode="MNHN" elevation="620" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.95" pageId="60" pageNumber="61" specimenCode="MNHN-IM-2000-38086" specimenCount="3" typeStatus="paratype">
<specimenCount id="7BEAFD3438620B20FE0CFE9256D0FEB9" box="[407,422,336,362]" pageId="60" pageNumber="61">3</specimenCount>
empty shells; stn DW4887;
<geoCoordinate id="08D8507A38620B20FC99FE8C541DFEB9" box="[770,875,334,362]" degrees="33" direction="south" minutes="17" orientation="latitude" pageId="60" pageNumber="61" precision="925" value="-33.283333">33°17′ S</geoCoordinate>
,
<geoCoordinate id="08D8507A38620B20FCE0FE8C5491FEB9" box="[891,999,334,362]" degrees="43" direction="east" minutes="57" orientation="longitude" pageId="60" pageNumber="61" precision="925" value="43.95">43°57′ E</geoCoordinate>
; depth
<quantity id="AA149B5838620B20FBDAFE8D53BAFEB9" box="[1089,1228,335,362]" metricMagnitude="2" metricUnit="m" metricValue="6.195" metricValueMax="6.4" metricValueMin="5.99" pageId="60" pageNumber="61" unit="m" value="619.5" valueMax="640.0" valueMin="599.0">
<elevation id="E6C1D18E38620B20FBDAFE8D53BAFEB9" box="[1089,1228,335,362]" metricMagnitude="2" metricUnit="m" metricValue="6.195" metricValueMax="6.4" metricValueMin="5.99" pageId="60" pageNumber="61" unit="m" value="619.5" valueMax="640.0" valueMin="599.0">599–640 m</elevation>
</quantity>
;
<date id="1952107D38620B20FB40FE925204FEB9" box="[1243,1394,336,362]" pageId="60" pageNumber="61" value="2017-05-03">
<collectingDate id="0916E99538620B20FB40FE925204FEB9" box="[1243,1394,336,362]" pageId="60" pageNumber="61" value="2017-05-03">3 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638620B20FF26FEB156ADFE5E" box="[189,475,371,397]" pageId="60" pageNumber="61">MNHN-IM-2000-38086</specimenCode>
</materialsCitation>
•
<materialsCitation id="DD843CE038620B20FE62FEB15558FE62" collectingDate="2017-05-06" collectionCode="MNHN" elevation="714" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="44.033333" pageId="60" pageNumber="61" specimenCode="MNHN-IM-2000-38087" specimenCount="1" typeStatus="paratype">
1 empty shell; stn DW4899;
<geoCoordinate id="08D8507A38620B20FCCEFEB054CBFE5E" box="[853,957,370,398]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="60" pageNumber="61" precision="925" value="-33.15">33°09′ S</geoCoordinate>
,
<geoCoordinate id="08D8507A38620B20FC57FEB05341FE5D" box="[972,1079,370,398]" degrees="44" direction="east" minutes="02" orientation="longitude" pageId="60" pageNumber="61" precision="925" value="44.033333">44°02′ E</geoCoordinate>
; depth
<quantity id="AA149B5838620B20FB0BFEB1526CFE5D" box="[1168,1306,371,398]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="60" pageNumber="61" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">
<elevation id="E6C1D18E38620B20FB0BFEB1526CFE5D" box="[1168,1306,371,398]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="60" pageNumber="61" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">707–720 m</elevation>
</quantity>
;
<date id="1952107D38620B20FAB2FEB1578DFE62" pageId="60" pageNumber="61" value="2017-05-06">
<collectingDate id="0916E99538620B20FAB2FEB1578DFE62" pageId="60" pageNumber="61" value="2017-05-06">6 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638620B20FE9CFE55555CFE62" box="[263,554,407,433]" pageId="60" pageNumber="61">MNHN-IM-2000-38087</specimenCode>
.
</materialsCitation>
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="25F6653638620B20FF26FE1A5388FB40" pageId="60" pageNumber="61" type="description">
<paragraph id="6D5336BD38620B20FF26FE1A5638FE21" blockId="60.[189,334,472,498]" box="[189,334,472,498]" pageId="60" pageNumber="61">
<heading id="361B81D138620B20FF26FE1A5638FE21" bold="true" box="[189,334,472,498]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="5F98EAAF38620B20FF26FE1A5638FE21" bold="true" box="[189,334,472,498]" pageId="60" pageNumber="61">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38620B20FF26FDCA5333FC55" blockId="60.[189,1400,520,902]" pageId="60" pageNumber="61">
<smallCapsWord id="6BB5A06138620B20FF26FDCA5677FDF3" baselines="539,539" box="[189,257,520,546]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="60" pageNumber="61">SHELL</smallCapsWord>
. Very small (height up to
<quantity id="AA149B5838620B20FDCCFDCA55CCFDF1" box="[599,698,520,546]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="60" pageNumber="61" unit="mm" value="1.3">1.3 mm</quantity>
), elevated-turbiniform; higher than wide (H/D = 1.1–1.2); whorls strongly rounded and suture deeply indented; periphery evenly rounded, close to mid-whorl; base rounded, deeply umbilicate; teleoconch of up to 3.25 whorls. First teleoconch whorl with ± 17 axial ribs, increasing in strength toward end of whorl; subsequent whorls with similar ribs, orthocline and progressively more widely spaced until final quarter of last adult whorl where they usually (but not always) become more close-set; penultimate whorl with ± 20 ribs; ribs narrow and blade-like, sometimes appearing weakly alate below adapical suture; intervals between ribs substantially wider than ribs, particularly on final two whorls; intervals sculptured by close-set, microscopic spiral threads (
<figureCitation id="F5D72A3838620B20FF5DFCE6565EFCEC" box="[198,296,804,831]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="60" pageNumber="61">Fig. 25I</figureCitation>
) and even finer and more close-set axial threads (visible only under SEM); basal sculpture similar, continuing into umbilicus; umbilicus relatively narrow and deep, its margin evenly rounded. Aperture circular; peristome complete, its edge simple; interior not nacreous.
</paragraph>
<paragraph id="6D5336BD38620B20FF26FC6C5587FBDC" blockId="60.[189,1399,941,1039]" pageId="60" pageNumber="61">
<smallCapsWord id="6BB5A06138620B20FF26FC6C563AFC15" baselines="961,961" box="[189,332,942,968]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Protoconch" pageId="60" pageNumber="61">PROTOCONCH</smallCapsWord>
(
<figureCitation id="F5D72A3838620B20FEFAFC6F56A4FC1B" box="[353,466,941,968]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="60" pageNumber="61">Fig. 25D</figureCitation>
). Globose and exsert; diameter 200–210 μm, ca 1.25 whorls; where unworn sculptured by fine anastomosing threads producing a network of close-set, irregularly-shaped pits; terminal lip more or less straight, not thickened.
</paragraph>
<paragraph id="6D5336BD38620B20FF26FBF55493FB83" blockId="60.[189,997,1078,1105]" box="[189,997,1078,1105]" pageId="60" pageNumber="61">
<smallCapsWord id="6BB5A06138620B20FF26FBF5566EFB9C" baselines="1098,1098" box="[189,280,1079,1105]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="60" pageNumber="61">COLOUR</smallCapsWord>
. Uniformly white; freshest specimens somewhat translucent.
</paragraph>
<paragraph id="6D5336BD38620B20FF26FBBB5388FB40" blockId="60.[189,1278,1144,1171]" box="[189,1278,1144,1171]" pageId="60" pageNumber="61">
<smallCapsWord id="6BB5A06138620B20FF26FBBB5634FB42" baselines="1164,1164" box="[189,322,1145,1171]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dimensions" pageId="60" pageNumber="61">DIMENSIONS</smallCapsWord>
.
<typeStatus id="B257881F38620B20FECAFBBA56B7FB41" box="[337,449,1144,1170]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
, height
<quantity id="AA149B5838620B20FD86FBBA55FFFB40" box="[541,649,1144,1171]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="60" pageNumber="61" unit="mm" value="1.14">1.14 mm</quantity>
, diameter
<quantity id="AA149B5838620B20FC99FBBA5418FB40" box="[770,878,1144,1171]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="60" pageNumber="61" unit="mm" value="0.98">0.98 mm</quantity>
; largest specimen, height
<quantity id="AA149B5838620B20FB06FBBA538CFB40" box="[1181,1274,1144,1171]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="60" pageNumber="61" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638620B20FF26FB785533FAF4" pageId="60" pageNumber="61" type="distribution">
<paragraph id="6D5336BD38620B20FF26FB785620FB07" blockId="60.[189,342,1210,1236]" box="[189,342,1210,1236]" pageId="60" pageNumber="61">
<heading id="361B81D138620B20FF26FB785620FB07" bold="true" box="[189,342,1210,1236]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="5F98EAAF38620B20FF26FB785620FB07" bold="true" box="[189,342,1210,1236]" pageId="60" pageNumber="61">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38620B20FF26FB285533FAF4" blockId="60.[189,1399,1257,1319]" pageId="60" pageNumber="61">
Known only from the slopes of Walters Shoal, at depths of
<quantity id="AA149B5838620B20FCE1FB2B5375FAD7" box="[890,1027,1257,1284]" metricMagnitude="2" metricUnit="m" metricValue="6.445" metricValueMax="7.07" metricValueMin="5.82" pageId="60" pageNumber="61" unit="m" value="644.5" valueMax="707.0" valueMin="582.0">582–707 m</quantity>
; dredged on substrata of coarse sand; living specimens unknown.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638620B20FF26FA8D53FFF967" pageId="60" pageNumber="61" type="discussion">
<paragraph id="6D5336BD38620B20FF26FA8D5659FABA" blockId="60.[189,303,1359,1385]" box="[189,303,1359,1385]" pageId="60" pageNumber="61">
<heading id="361B81D138620B20FF26FA8D5659FABA" bold="true" box="[189,303,1359,1385]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="5F98EAAF38620B20FF26FA8D5659FABA" bold="true" box="[189,303,1359,1385]" pageId="60" pageNumber="61">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38620B20FF26FABF53FFF967" blockId="60.[189,1399,1405,1717]" pageId="60" pageNumber="61">
Other somewhat elevated species of
<taxonomicName id="AAEC4D3E38620B20FDF4FABF545DFA44" authorityName="Clarke" authorityYear="1961" box="[623,811,1405,1431]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="60" pageNumber="61" phylum="Mollusca" rank="genus">
<emphasis id="5F98EAAF38620B20FDF4FABF545DFA44" box="[623,811,1405,1431]" italics="true" pageId="60" pageNumber="61">Benthobrookula</emphasis>
</taxonomicName>
include
<emphasis id="5F98EAAF38620B20FC0AFABC54CEFA44" box="[913,952,1406,1431]" italics="true" pageId="60" pageNumber="61">Be.</emphasis>
<taxonomicName id="AAEC4D3E38620B20FC5BFABF5204FA4B" authority="(Schwabe &amp; Engl, 2008)" baseAuthorityName="Schwabe &amp; Engl" baseAuthorityYear="2008" box="[960,1394,1405,1432]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="charleenae">
<emphasis id="5F98EAAF38620B20FC5BFABF5334FA44" box="[960,1090,1405,1431]" italics="true" pageId="60" pageNumber="61">charleenae</emphasis>
(Schwabe &amp; Engl, 2008)
</taxonomicName>
,
<emphasis id="5F98EAAF38620B20FF25FA6056F2FA68" box="[190,388,1441,1467]" italics="true" pageId="60" pageNumber="61">Be. kerguelensis</emphasis>
(
<bibRefCitation id="097D4B4C38620B20FE02FA63554CFA6F" author="Thiele J." box="[409,570,1441,1468]" pageId="60" pageNumber="61" pagination="35 - 382" refId="ref63529" refString="Thiele J. 1925. Gastropoda der deutschen Tiefsee-Expedition, II. Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot;, 1898 - 1899, 17 (2): 35 - 382 [dual pagination; also numbered 1 - 348]. Gustav Fischer, Jena." type="journal article" year="1925">Thiele, 1925</bibRefCitation>
),
<emphasis id="5F98EAAF38620B20FDCDFA60550AFA68" box="[598,636,1442,1467]" italics="true" pageId="60" pageNumber="61">Be.</emphasis>
<taxonomicName id="AAEC4D3E38620B20FD11FA6054AFFA68" authority="(Gatliff, 1906)" baseAuthorityName="Gatliff" baseAuthorityYear="1906" box="[650,985,1441,1467]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="nepeanensis">
<emphasis id="5F98EAAF38620B20FD11FA60546FFA68" box="[650,793,1442,1467]" italics="true" pageId="60" pageNumber="61">nepeanensis</emphasis>
(Gatliff, 1906)
</taxonomicName>
and
<emphasis id="5F98EAAF38620B20FBBBFA6053DAFA68" box="[1056,1196,1441,1467]" italics="true" pageId="60" pageNumber="61">Be. strebeli</emphasis>
(Powell, 1951).
<taxonomicName id="AAEC4D3E38620B20FF26FA075575FA0C" baseAuthorityName="Schwabe &amp; Engl" baseAuthorityYear="2008" box="[189,515,1477,1503]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="charleenae">
<emphasis id="5F98EAAF38620B20FF26FA075575FA0C" box="[189,515,1477,1503]" italics="true" pageId="60" pageNumber="61">Benthobrookula charleenae</emphasis>
</taxonomicName>
, from over
<quantity id="AA149B5838620B20FD14FA07559FFA0C" box="[655,745,1477,1503]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="60" pageNumber="61" unit="m" value="2900.0">2900 m</quantity>
off
<collectingCountry id="15FB762D38620B20FC81FA075315FA0C" box="[794,1123,1477,1503]" name="South Georgia and the South Sandwich Islands" pageId="60" pageNumber="61">the South Sandwich Islands</collectingCountry>
, differs in having more close-set axial ribs (27 vs 20 on penultimate whorl) and a larger protoconch (diameter 280 vs 200– 210 μm). In
<emphasis id="5F98EAAF38620B20FEC8F9CF5562F9F5" box="[339,532,1548,1574]" italics="true" pageId="60" pageNumber="61">Be. kerguelensis</emphasis>
the shell is larger (height
<quantity id="AA149B5838620B20FCD5F9CE54DDF9F5" box="[846,939,1548,1574]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="60" pageNumber="61" unit="mm" value="1.6">1.6 mm</quantity>
) and, judging from the original figure, the last adult whorl is deeper and the spire less elevated.
<taxonomicName id="AAEC4D3E38620B20FCEBF9ED53B3F99A" baseAuthorityName="Gatliff" baseAuthorityYear="1906" box="[880,1221,1583,1609]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="nepeanensis">
<emphasis id="5F98EAAF38620B20FCEBF9ED53B3F99A" box="[880,1221,1583,1609]" italics="true" pageId="60" pageNumber="61">Benthobrookula nepeanensis</emphasis>
</taxonomicName>
from southern
<collectingCountry id="15FB762D38620B20FF24F991565DF9BE" box="[191,299,1619,1645]" name="Australia" pageId="60" pageNumber="61">Australia</collectingCountry>
has an even more elevated spire and a much narrower umbilicus, whereas in
<emphasis id="5F98EAAF38620B20FB29F9965240F9BE" box="[1202,1334,1619,1645]" italics="true" pageId="60" pageNumber="61">Be. strebeli</emphasis>
from
<collectingCountry id="15FB762D38620B20FF26F9B5561EF942" box="[189,360,1655,1681]" name="South Georgia and the South Sandwich Islands" pageId="60" pageNumber="61">South Georgia</collectingCountry>
the axial ribs are more prosocline and less blade-like, and the base is imperforate with more conspicuous spiral sculpture (
<typeStatus id="B257881F38620B20FDC5F95855B1F967" box="[606,711,1690,1716]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
figured by
<bibRefCitation id="097D4B4C38620B20FCD6F9585355F967" author="Zelaya D. G. &amp; Absalao R. S. &amp; Pimenta A. D." box="[845,1059,1690,1717]" pageId="60" pageNumber="61" pagination="77 - 87" refId="ref64671" refString="Zelaya D. G., Absalao R. S. &amp; Pimenta A. D. 2006. A revision of Benthobrookula Clarke, 1961 (Gastropoda, Trochoidea) in the southwestern Atlantic Ocean. Journal of Molluscan Studies 72 (1): 77 - 87. https: // doi. org / 10.1093 / mollus / eyi 050" type="journal article" year="2006">
Zelaya
<emphasis id="5F98EAAF38620B20FC39F95954A0F967" box="[930,982,1690,1716]" italics="true" pageId="60" pageNumber="61">et al</emphasis>
. 2006
</bibRefCitation>
: fig. 6a).
</paragraph>
</subSubSection>
</treatment>
</document>
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