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<document id="A84E8F5BF65EC64B870EA19F8A967326" ID-DOI="10.5852/ejt.2024.923.2445" ID-ISSN="2118-9773" ID-Zenodo-Dep="10727741" ID-ZooBank="AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1709215085450" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2024" docId="E54587AB380E0B4FFD08F9AF5613FE2B" docLanguage="en" docName="EJT.2024.923.1-119.pdf.imf" docOrigin="European Journal of Taxonomy 923" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Mikro crassus Herbert 2024, sp. nov." docType="treatment" docUuid="3CE83169-0B5E-4678-85EA-1150A6EA8C60" docUuidSource="ZooBank" docVersion="4" lastPageNumber="83" masterDocId="197CFFD3385E0B1CFF9BFFC25776FFD3" masterDocTitle="The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species" masterLastPageNumber="119" masterPageNumber="1" pageNumber="81" updateTime="1710414732225" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D92D55054B457A5600A80FC0C32FBA3F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="48A64C852425A7BD42C75D233950789B">The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="72D48EB5F666DCC081BFA1F2780AC97A">Herbert, David G.</mods:namePart>
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<mods:title id="D70EC8A5C34524D884B8792E6763E465">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7733C0862DDED29A242BCE5F20118ECD">
<mods:date id="DDC4FC5EF8B1AE443EFB5301FCBEB522">2024</mods:date>
<mods:detail id="6A31F68FE1979C648DE047455BB11125" type="pubDate">
<mods:number id="E8D402CED843F963ED182847C8FEB7A7">2024-02-28</mods:number>
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<mods:detail id="2B5212C20A6D22CA67DE33762D4B06C2" type="volume">
<mods:number id="EBE40416903130079EF85E5A2AD0BA37">923</mods:number>
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<mods:url id="5FF7ACF3DDEA35B850A388EB335B72EF">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835</mods:url>
</mods:location>
<mods:classification id="6D5FE63883A749D0A04C8C5C1C62D550">journal article</mods:classification>
<mods:identifier id="1805B88F467C45E4F88AD31A1EA0B25D" type="DOI">10.5852/ejt.2024.923.2445</mods:identifier>
<mods:identifier id="A20C816E1BC4CC2B2A4F793B2C072928" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="DBB5B308896B8ADA5D7800B5F55335E8" type="Zenodo-Dep">10727741</mods:identifier>
<mods:identifier id="AF50251100E2053FFFC810D1A376B874" type="ZooBank">AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3</mods:identifier>
</mods:mods>
<treatment id="E54587AB380E0B4FFD08F9AF5613FE2B" LSID="urn:lsid:zoobank.org:act:3CE83169-0B5E-4678-85EA-1150A6EA8C60" httpUri="http://treatment.plazi.org/id/E54587AB380E0B4FFD08F9AF5613FE2B" lastPageId="83" lastPageNumber="83" pageId="80" pageNumber="81">
<subSubSection id="25F66536380E0B4CFD08F9AF54D7F95B" box="[659,929,1645,1672]" pageId="80" pageNumber="81" type="nomenclature">
<paragraph id="6D5336BD380E0B4CFD08F9AF54D7F95B" blockId="80.[370,1218,1645,1741]" box="[659,929,1645,1672]" pageId="80" pageNumber="81">
<heading id="361B81D1380E0B4CFD08F9AF54D7F95B" box="[659,929,1645,1672]" centered="true" fontSize="11" level="2" pageId="80" pageNumber="81" reason="2">
<taxonomicName id="AAEC4D3E380E0B4CFD08F9AF5449F95B" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[659,831,1645,1672]" class="Gastropoda" family="Skeneidae" genus="Mikro" kingdom="Animalia" order="Trochida" pageId="80" pageNumber="81" phylum="Mollusca" rank="species" species="crassus" status="sp. nov.">
<emphasis id="5F98EAAF380E0B4CFD08F9AF5449F95B" bold="true" box="[659,831,1645,1672]" italics="true" pageId="80" pageNumber="81">Mikro crassus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4380E0B4CFCDDF9AC54D7F95B" box="[838,929,1646,1672]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25F66536380E0B4CFEE9F9525435F91E" pageId="80" pageNumber="81" type="description">
<paragraph id="6D5336BD380E0B4CFEE9F95253B4F978" blockId="80.[370,1218,1645,1741]" box="[370,1218,1680,1707]" pageId="80" pageNumber="81">
<uri id="197D3ABF380E0B4CFEE9F95253B4F978" box="[370,1218,1680,1707]" pageId="80" pageNumber="81">
urn:lsid:zoobank.org:act:
<uuid id="194A0C68380E0B4CFD0AF95253B4F978" box="[657,1218,1680,1707]" pageId="80" pageNumber="81">3CE83169-0B5E-4678-85EA-1150A6EA8C60</uuid>
</uri>
</paragraph>
<paragraph id="6D5336BD380E0B4CFD6BF9715435F91E" blockId="80.[370,1218,1645,1741]" box="[752,835,1715,1741]" pageId="80" pageNumber="81">
<figureCitation id="F5D72A38380E0B4CFD6BF9715435F91E" box="[752,835,1715,1741]" captionStart="Fig" captionStartId="82.[189,232,1854,1880]" captionTargetBox="[189,1399,480,1818]" captionTargetId="figure-163@82.[189,1399,480,1818]" captionTargetPageId="82" captionText="Fig. 33. Mikro crassussp. nov. A–B. Holotype, apertural and oblique basal views (apex missing), diameter 2.9 mm (MNHN-IM-2000-38066). C. Apical view of juvenile specimen with intact apex, diameter 1.5 mm (paratype, MNHN-IM-2000-38067). D–E. Specimen with brown superficial deposit, diameter 1.7 mm (paratype, MNHN-IM-2000-38067). F. Protoconch of C, scale bar =100 μm. G. Composite image of holotype superimposed on apex of C, diameter 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.10727838" httpUri="https://zenodo.org/record/10727838/files/figure.png" pageId="80" pageNumber="81">Fig. 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="25F66536380E0B4CFF26F93154B0F83B" pageId="80" pageNumber="81" type="diagnosis">
<paragraph id="6D5336BD380E0B4CFF26F9315640F8DE" blockId="80.[189,310,1779,1805]" box="[189,310,1779,1805]" pageId="80" pageNumber="81">
<heading id="361B81D1380E0B4CFF26F9315640F8DE" bold="true" box="[189,310,1779,1805]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="5F98EAAF380E0B4CFF26F9315640F8DE" bold="true" box="[189,310,1779,1805]" pageId="80" pageNumber="81">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD380E0B4CFF26F8E354B0F83B" blockId="80.[189,1399,1825,2025]" pageId="80" pageNumber="81">Shell globose-turbiniform, white, thick, H/D ± 1.0, whorls rounded with distinct shoulder delineated by rounded spiral cord, beginning immediately after protoconch, becoming less prominent on last adult whorl; umbilicus narrow, bordered by two well-defined spiral cords, with two finer spiral cords within it; whorls otherwise smooth and glossy; aperture subcircular; basal lip weakly notched by cords bordering umbilicus; outer lip shallowly concave between suture and shoulder; interior of aperture somewhat thickened with opaque, chalky layer (?nacre); protoconch smooth.</paragraph>
</subSubSection>
<subSubSection id="25F66536380F0B4DFF26FECB534DFE81" box="[189,1083,265,338]" pageId="81" pageNumber="82" type="etymology">
<paragraph id="6D5336BD380F0B4DFF26FECB5632FEF0" blockId="81.[189,324,265,291]" box="[189,324,265,291]" pageId="81" pageNumber="82">
<heading id="361B81D1380F0B4DFF26FECB5632FEF0" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
<emphasis id="5F98EAAF380F0B4DFF26FECB5632FEF0" bold="true" box="[189,324,265,291]" pageId="81" pageNumber="82">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26FEFA534DFE81" blockId="81.[189,1083,312,338]" box="[189,1083,312,338]" pageId="81" pageNumber="82">
From the Latin
<taxonomicName id="AAEC4D3E380F0B4DFEEBFEFA56A4FE82" authorityName="Herbert" authorityYear="2024" box="[368,466,312,338]" class="Gastropoda" family="Skeneidae" genus="Mikro" kingdom="Animalia" order="Trochida" pageId="81" pageNumber="82" phylum="Mollusca" rank="species" species="crassus">
”
<emphasis id="5F98EAAF380F0B4DFEE1FEFA56A4FE82" box="[378,466,312,337]" italics="true" pageId="81" pageNumber="82">crassus</emphasis>
</taxonomicName>
” – “thick”; in reference to the thickness of the shell.
</paragraph>
</subSubSection>
<subSubSection id="25F66536380F0B4DFF26FEB856F6FD46" pageId="81" pageNumber="82" type="materials_examined">
<paragraph id="6D5336BD380F0B4DFF26FEB856DCFE47" blockId="81.[189,426,378,404]" box="[189,426,378,404]" pageId="81" pageNumber="82">
<heading id="361B81D1380F0B4DFF26FEB856DCFE47" bold="true" box="[189,426,378,404]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
<emphasis id="5F98EAAF380F0B4DFF26FEB856DCFE47" bold="true" box="[189,426,378,404]" pageId="81" pageNumber="82">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="DD843CE0380F0B4DFF71FE6B55F9FDD8" collectingDate="2017-05-11" collectionCode="MNHN" elevation="1577" latitude="-33.866665" location="WALTERS SHOAL" longLatPrecision="1202" longitude="44.083332" pageId="81" pageNumber="82" specimenCode="MNHN-IM-2000-38066" specimenCount="1" typeStatus="holotype">
<paragraph id="6D5336BD380F0B4DFF71FE6B562AFE10" blockId="81.[189,1399,425,523]" box="[234,348,425,451]" pageId="81" pageNumber="82">
<emphasis id="5F98EAAF380F0B4DFF71FE6B562AFE10" bold="true" box="[234,348,425,451]" pageId="81" pageNumber="82">
<typeStatus id="B257881F380F0B4DFF71FE6B562AFE10" box="[234,348,425,451]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26FE0F55F9FDD8" blockId="81.[189,1399,425,523]" pageId="81" pageNumber="82">
<location id="68336066380F0B4DFF26FE0F56DBFE34" LSID="urn:lsid:plazi:treatment:E54587AB380E0B4FFD08F9AF5613FE2B:68336066380F0B4DFF26FE0F56DBFE34" box="[189,429,461,487]" latitude="-33.866665" longLatPrecision="1202" longitude="44.083332" name="WALTERS SHOAL" pageId="81" pageNumber="82">WALTERS SHOAL</location>
• empty shell; south plain, stn CP4913;
<geoCoordinate id="08D8507A380F0B4DFC0AFE09548FFE34" box="[913,1017,459,487]" degrees="33" direction="south" minutes="52" orientation="latitude" pageId="81" pageNumber="82" precision="925" value="-33.866665">33°52′ S</geoCoordinate>
,
<geoCoordinate id="08D8507A380F0B4DFB9CFE095304FE34" box="[1031,1138,459,487]" degrees="44" direction="east" minutes="05" orientation="longitude" pageId="81" pageNumber="82" precision="925" value="44.083332">44°05′ E</geoCoordinate>
; depth
<quantity id="AA149B58380F0B4DFB51FE0F5207FE34" box="[1226,1393,461,488]" metricMagnitude="3" metricUnit="m" metricValue="1.577" metricValueMax="1.6149999999999998" metricValueMin="1.539" pageId="81" pageNumber="82" unit="m" value="1577.0" valueMax="1615.0" valueMin="1539.0">
<elevation id="E6C1D18E380F0B4DFB51FE0F5207FE34" box="[1226,1393,461,488]" metricMagnitude="3" metricUnit="m" metricValue="1.577" metricValueMax="1.6149999999999998" metricValueMin="1.539" pageId="81" pageNumber="82" unit="m" value="1577.0" valueMax="1615.0" valueMin="1539.0">1539–1615 m</elevation>
</quantity>
;
<date id="1952107D380F0B4DFF26FE33562AFDD8" box="[189,348,497,523]" pageId="81" pageNumber="82" value="2017-05-11">
<collectingDate id="0916E995380F0B4DFF26FE33562AFDD8" box="[189,348,497,523]" pageId="81" pageNumber="82" value="2017-05-11">11 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC6380F0B4DFEF3FE3355FCFDD8" box="[360,650,497,523]" pageId="81" pageNumber="82">MNHN-IM-2000-38066</specimenCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="DD843CE0380F0B4DFF71FDF156F6FD46" collectingDate="2017-05-11" collectionCode="MNHN" elevation="1577" latitude="-33.866665" location="WALTERS SHOAL" longLatPrecision="1202" longitude="44.083332" pageId="81" pageNumber="82" specimenCode="MNHN- IM-2000-38067" specimenCount="14" typeStatus="paratype">
<paragraph id="6D5336BD380F0B4DFF71FDF15611FD9E" blockId="81.[189,1398,563,661]" box="[234,359,563,589]" pageId="81" pageNumber="82">
<emphasis id="5F98EAAF380F0B4DFF71FDF15611FD9E" bold="true" box="[234,359,563,589]" pageId="81" pageNumber="82">
<typeStatus id="B257881F380F0B4DFF71FDF15611FD9E" box="[234,359,563,589]" pageId="81" pageNumber="82" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26FD9556F6FD46" blockId="81.[189,1398,563,661]" pageId="81" pageNumber="82">
<location id="68336066380F0B4DFF26FD9556C6FDA2" LSID="urn:lsid:plazi:treatment:E54587AB380E0B4FFD08F9AF5613FE2B:68336066380F0B4DFF26FD9556C6FDA2" box="[189,432,599,625]" name="WALTERS SHOAL" pageId="81" pageNumber="82">WALTERS SHOAL</location>
•
<specimenCount id="7BEAFD34380F0B4DFE49FD955699FDA2" box="[466,495,599,625]" pageId="81" pageNumber="82">14</specimenCount>
empty shells; same collection data as for holotype; SEM shell;
<specimenCode id="3D4A9EC6380F0B4DFA88FD95560AFD46" pageId="81" pageNumber="82">MNHN- IM-2000-38067</specimenCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="25F66536380F0B4DFF26FD7F5668F9D8" pageId="81" pageNumber="82" type="description">
<paragraph id="6D5336BD380F0B4DFF26FD7F5638FD04" blockId="81.[189,334,701,727]" box="[189,334,701,727]" pageId="81" pageNumber="82">
<heading id="361B81D1380F0B4DFF26FD7F5638FD04" bold="true" box="[189,334,701,727]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
<emphasis id="5F98EAAF380F0B4DFF26FD7F5638FD04" bold="true" box="[189,334,701,727]" pageId="81" pageNumber="82">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26FD2E53FAFB0B" blockId="81.[187,1401,748,1241]" pageId="81" pageNumber="82">
<smallCapsWord id="6BB5A061380F0B4DFF26FD2E5677FCD6" baselines="767,768" box="[189,257,748,774]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="81" pageNumber="82">SHELL</smallCapsWord>
. Small (diameter up to
<quantity id="AA149B58380F0B4DFD84FD2E5509FCD5" box="[543,639,748,775]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="81" pageNumber="82" unit="mm" value="3.5">3.5 mm</quantity>
), relatively thick, globose-turbiniform, height and width similar (apex missing in most specimens); whorls rounded with distinct shoulder delineated by a rounded spiral cord, middle of shoulder shallowly sunken; teleoconch of up to 3.5 whorls; shoulder cord begins immediately after protoconch as low, broad rib and persists throughout, but becomes broader and less prominent on last adult whorl; periphery and base evenly rounded, periphery at mid-whorl; umbilicus narrow, bordered by two well-defined spiral cords, inner of which forms umbilical rim and ends at junction of columella and basal lips; outer one peri-umbilical, ending at lowest point of basal lip; interval between these cords distinctly concave; two additional, finer spiral cords descending steeply within umbilicus, ending on columella lip, inner one weaker; whorls otherwise smooth and glossy, sculptured only by extremely fine collabral growth-lines. Aperture subcircular, prosocline, peristome interrupted in parietal region; columella lip duplex, with thin outer layer protruding slightly beyond thicker inner layer; basal lip thickened and weakly notched at termination of outer two cords bordering umbilicus; outer lip simple, notched at end of shoulder cord and shallowly concave between this and its insertion on penultimate whorl; interior somewhat thickened, inner layer opaque and chalky.
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26FAC35201FAED" blockId="81.[189,1399,1281,1343]" pageId="81" pageNumber="82">
<smallCapsWord id="6BB5A061380F0B4DFF26FAC3563AFACA" baselines="1300,1300" box="[189,332,1281,1307]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Protoconch" pageId="81" pageNumber="82">PROTOCONCH</smallCapsWord>
(
<figureCitation id="F5D72A38380F0B4DFEF9FAC356B8FAC8" box="[354,462,1281,1307]" captionStart="Fig" captionStartId="82.[189,232,1854,1880]" captionTargetBox="[189,1399,480,1818]" captionTargetId="figure-163@82.[189,1399,480,1818]" captionTargetPageId="82" captionText="Fig. 33. Mikro crassussp. nov. A–B. Holotype, apertural and oblique basal views (apex missing), diameter 2.9 mm (MNHN-IM-2000-38066). C. Apical view of juvenile specimen with intact apex, diameter 1.5 mm (paratype, MNHN-IM-2000-38067). D–E. Specimen with brown superficial deposit, diameter 1.7 mm (paratype, MNHN-IM-2000-38067). F. Protoconch of C, scale bar =100 μm. G. Composite image of holotype superimposed on apex of C, diameter 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.10727838" httpUri="https://zenodo.org/record/10727838/files/figure.png" pageId="81" pageNumber="82">Fig. 33F</figureCitation>
). Comprising just less than one whorl, globose and slightly exsert, diameter ± 330 μm; smooth save for an extremely fine superficial granulation; terminal lip straight, slightly flared.
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26FAA5549FFA76" blockId="81.[188,1398,1383,1445]" pageId="81" pageNumber="82">
<smallCapsWord id="6BB5A061380F0B4DFF26FAA5566EFAAC" baselines="1402,1402" box="[189,280,1383,1409]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="81" pageNumber="82">COLOUR</smallCapsWord>
. Uniformly milky-white; inner shell layer opaque and chalky. Some individuals partly or almost completely covered with lustreless dark brown coating (
<figureCitation id="F5D72A38380F0B4DFCD5FA4854AAFA76" box="[846,988,1418,1445]" captionStart="Fig" captionStartId="82.[189,232,1854,1880]" captionTargetBox="[189,1399,480,1818]" captionTargetId="figure-163@82.[189,1399,480,1818]" captionTargetPageId="82" captionText="Fig. 33. Mikro crassussp. nov. A–B. Holotype, apertural and oblique basal views (apex missing), diameter 2.9 mm (MNHN-IM-2000-38066). C. Apical view of juvenile specimen with intact apex, diameter 1.5 mm (paratype, MNHN-IM-2000-38067). D–E. Specimen with brown superficial deposit, diameter 1.7 mm (paratype, MNHN-IM-2000-38067). F. Protoconch of C, scale bar =100 μm. G. Composite image of holotype superimposed on apex of C, diameter 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.10727838" httpUri="https://zenodo.org/record/10727838/files/figure.png" pageId="81" pageNumber="82">Fig. 33D–E</figureCitation>
).
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26FA0F5668F9D8" blockId="81.[189,1399,1484,1547]" pageId="81" pageNumber="82">
<smallCapsWord id="6BB5A061380F0B4DFF26FA0F5634FA36" baselines="1504,1504" box="[189,322,1485,1511]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dimensions" pageId="81" pageNumber="82">DIMENSIONS</smallCapsWord>
.
<typeStatus id="B257881F380F0B4DFECFFA0F56B2FA34" box="[340,452,1485,1511]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
, height
<quantity id="AA149B58380F0B4DFDB3FA0E55F0FA34" box="[552,646,1484,1511]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="81" pageNumber="82" unit="mm" value="2.6">2.6 mm</quantity>
(apex missing), diameter
<quantity id="AA149B58380F0B4DFC5DFA0F5350FA34" box="[966,1062,1485,1511]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="81" pageNumber="82" unit="mm" value="2.9">2.9 mm</quantity>
; largest specimen, diameter
<quantity id="AA149B58380F0B4DFF26FA32566CF9D8" box="[189,282,1520,1547]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="81" pageNumber="82" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="25F66536380F0B4DFF26F9F055B2F973" pageId="81" pageNumber="82" type="distribution">
<paragraph id="6D5336BD380F0B4DFF26F9F05620F99F" blockId="81.[189,342,1586,1612]" box="[189,342,1586,1612]" pageId="81" pageNumber="82">
<heading id="361B81D1380F0B4DFF26F9F05620F99F" bold="true" box="[189,342,1586,1612]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
<emphasis id="5F98EAAF380F0B4DFF26F9F05620F99F" bold="true" box="[189,342,1586,1612]" pageId="81" pageNumber="82">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26F9A055B2F973" blockId="81.[189,1399,1634,1696]" pageId="81" pageNumber="82">
Known only from the plain surrounding Walters Shoal, at a depth of
<quantity id="AA149B58380F0B4DFC76F9A053E5F9AF" box="[1005,1171,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.577" metricValueMax="1.6149999999999998" metricValueMin="1.539" pageId="81" pageNumber="82" unit="m" value="1577.0" valueMax="1615.0" valueMin="1539.0">1539–1615 m</quantity>
; dredged on coarse sandy substrata; living specimens unknown.
</paragraph>
</subSubSection>
<subSubSection id="25F66536380F0B4FFF26F90A5613FE2B" lastPageId="83" lastPageNumber="84" pageId="81" pageNumber="82" type="discussion">
<paragraph id="6D5336BD380F0B4DFF26F90A5659F931" blockId="81.[189,303,1736,1762]" box="[189,303,1736,1762]" pageId="81" pageNumber="82">
<heading id="361B81D1380F0B4DFF26F90A5659F931" bold="true" box="[189,303,1736,1762]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
<emphasis id="5F98EAAF380F0B4DFF26F90A5659F931" bold="true" box="[189,303,1736,1762]" pageId="81" pageNumber="82">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD380F0B4DFF26F93554B5F83B" blockId="81.[189,1399,1783,2025]" pageId="81" pageNumber="82">
<taxonomicName id="AAEC4D3E380F0B4DFF26F9355611F8C2" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[189,359,1783,1809]" class="Gastropoda" family="Skeneidae" genus="Mikro" kingdom="Animalia" order="Trochida" pageId="81" pageNumber="82" phylum="Mollusca" rank="species" species="crassus" status="sp. nov.">
<emphasis id="5F98EAAF380F0B4DFF26F9355611F8C2" box="[189,359,1783,1809]" italics="true" pageId="81" pageNumber="82">Mikro crassus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4380F0B4DFEEAF93A56A6F8C1" box="[369,464,1784,1810]" pageId="81" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
is considerably larger than any of the other species currently referred to the genus (most with height &lt;
<quantity id="AA149B58380F0B4DFE62F8D9552EF8E6" box="[505,600,1819,1846]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="81" pageNumber="82" unit="mm" value="1.5">1.5 mm</quantity>
). The most similar species is
<taxonomicName id="AAEC4D3E380F0B4DFC29F8DE56E0F88A" authority="Hoffman, Van Heugten &amp; Lavaleye, 2010" authorityName="Hoffman, Van Heugten &amp; Lavaleye" authorityYear="2010" class="Gastropoda" family="Skeneidae" genus="Mikro" kingdom="Animalia" order="Trochida" pageId="81" pageNumber="82" phylum="Mollusca" rank="species" species="hattonensis">
<emphasis id="5F98EAAF380F0B4DFC29F8DE5329F8E6" box="[946,1119,1819,1845]" italics="true" pageId="81" pageNumber="82">M. hattonensis</emphasis>
Hoffman, Van Heugten &amp; Lavaleye, 2010
</taxonomicName>
from the north Atlantic, but that species has deeper spire whorls and thus a more elevated spire, and it has an unusual pitted (shagreened) micro-sculpture on the shoulder and in the umbilicus. There is also some resemblance to an unidentified species of
<taxonomicName id="AAEC4D3E380F0B4DFB92F8445241F872" authority="Bush, 1897" authorityName="Bush" authorityYear="1897" box="[1033,1335,1926,1953]" class="Gastropoda" family="Skeneidae" genus="Moelleriopsis" kingdom="Animalia" order="Seguenziida" pageId="81" pageNumber="82" phylum="Mollusca" rank="genus">
<emphasis id="5F98EAAF380F0B4DFB92F84453DCF873" box="[1033,1194,1926,1952]" italics="true" pageId="81" pageNumber="82">Moelleriopsis</emphasis>
Bush, 1897
</taxonomicName>
from
<collectingCountry id="15FB762D380F0B4DFF26F8685656F817" box="[189,288,1962,1988]" name="Réunion" pageId="81" pageNumber="82">Réunion</collectingCountry>
figured by
<bibRefCitation id="097D4B4C380F0B4DFE34F8695538F816" author="Waren A." box="[431,590,1963,1989]" pageId="81" pageNumber="82" pagination="149 - 248" refId="ref64085" refString="Waren A. 1992. New and little known &quot; skeneimorph &quot; gastropods from the Mediterranean Sea and the adjacent Atlantic Ocean. Bollettino Malacologico 27 (10 - 12): 149 - 248." type="journal article" year="1992">Warén (1992</bibRefCitation>
: fig. 31a), but that has a less exsert protoconch with distinct spiral sculpture and a strongly channelled suture (
<bibRefCitation id="097D4B4C380F0B4DFD21F80C5439F83B" author="Waren A." box="[698,847,1998,2024]" pageId="81" pageNumber="82" pagination="149 - 248" refId="ref64085" refString="Waren A. 1992. New and little known &quot; skeneimorph &quot; gastropods from the Mediterranean Sea and the adjacent Atlantic Ocean. Bollettino Malacologico 27 (10 - 12): 149 - 248." type="journal article" year="1992">Warén 1992</bibRefCitation>
: fig. 26c).
</paragraph>
<paragraph id="6D5336BD380C0B4FFF26FECB5613FE2B" blockId="82.[189,1399,265,432]" lastBlockId="83.[189,1399,265,504]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
The 15 shells of
<taxonomicName id="AAEC4D3E380C0B4EFE1BFECB5550FEF0" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[384,550,265,291]" class="Gastropoda" family="Skeneidae" genus="Mikro" kingdom="Animalia" order="Trochida" pageId="82" pageNumber="83" phylum="Mollusca" rank="species" species="crassus" status="sp. nov.">
<emphasis id="5F98EAAF380C0B4EFE1BFECB5550FEF0" box="[384,550,265,291]" italics="true" pageId="82" pageNumber="83">Mikro crassus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4380C0B4EFDB5FEC855FEFEF7" box="[558,648,266,292]" pageId="82" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
available are all damaged to some extent and most are juvenile. They all came from a single station and many evince the remains of a dark brown periostracum-like superficial deposit (
<figureCitation id="F5D72A38380C0B4EFE3CFE8D5545FEB9" box="[423,563,335,362]" captionStart="Fig" captionStartId="82.[189,232,1854,1880]" captionTargetBox="[189,1399,480,1818]" captionTargetId="figure-163@82.[189,1399,480,1818]" captionTargetPageId="82" captionText="Fig. 33. Mikro crassussp. nov. A–B. Holotype, apertural and oblique basal views (apex missing), diameter 2.9 mm (MNHN-IM-2000-38066). C. Apical view of juvenile specimen with intact apex, diameter 1.5 mm (paratype, MNHN-IM-2000-38067). D–E. Specimen with brown superficial deposit, diameter 1.7 mm (paratype, MNHN-IM-2000-38067). F. Protoconch of C, scale bar =100 μm. G. Composite image of holotype superimposed on apex of C, diameter 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.10727838" httpUri="https://zenodo.org/record/10727838/files/figure.png" pageId="82" pageNumber="83">Fig. 33D–E</figureCitation>
), possibly of a ferruginous nature, suggesting that the species may live in an unusual and highly specialised microhabitat associated with anoxic conditions (
<bibRefCitation id="097D4B4C380C0B4EFB3AFEB1578DFE63" author="Waren A. &amp; Bouchet P." pageId="82" pageNumber="83" pagination="67 - 102" refId="ref64156" refString="Waren A. &amp; Bouchet P. 1989. New gastropods from East Pacific hydrothermal vents. Zoologica Scripta 18 (1): 67 - 102. https: // doi. org / 10.1111 / j. 1463 - 6409.1989. tb 00124. x" type="journal article" year="1989">Warén &amp; Bouchet 1989</bibRefCitation>
;
<bibRefCitation id="097D4B4C380C0B4EFE9CFE5456ECFE63" author="Waren A." box="[263,410,406,432]" pageId="82" pageNumber="83" pagination="53 - 124" refId="ref64053" refString="Waren A. 1991. New and little known Mollusca from Iceland and Scandinavia. Sarsia 76: 53 - 124. https: // doi. org / 10.1080 / 00364827.1991.10413466" type="journal article" year="1991">Warén 1991</bibRefCitation>
). A similar superficial layer is evident in the Antarctic
<taxonomicName id="AAEC4D3E380C0B4FFB85FE55561CFEF0" authority="Melvill &amp; Standen, 1912" authorityName="Melvill &amp; Standen" authorityYear="1912" class="Gastropoda" family="Littorinidae" genus="Lacuna" kingdom="Animalia" lastPageId="83" lastPageNumber="84" order="Littorinimorpha" pageId="82" pageNumber="83" phylum="Mollusca" rank="species" species="abyssicola">
<emphasis id="5F98EAAF380C0B4EFB85FE55538CFE63" box="[1054,1274,406,432]" italics="true" pageId="82" pageNumber="83">Lacuna abyssicola</emphasis>
Melvill &amp; Standen, 1912
</taxonomicName>
, a species which
<bibRefCitation id="097D4B4C380D0B4FFDD8FECB55A5FEF0" author="Engl" box="[579,723,265,291]" pageId="83" pageNumber="84" refId="ref59972" refString="Engl W. 2012. Shells of Antarctica. Conchbooks, Hackenheim, Germany." type="book" year="2012">Engl (2012)</bibRefCitation>
referred to
<taxonomicName id="AAEC4D3E380D0B4FFCFCFECB537EFEF0" authorityName="Bush" authorityYear="1897" box="[871,1032,265,291]" class="Gastropoda" family="Skeneidae" genus="Moelleriopsis" kingdom="Animalia" order="Seguenziida" pageId="83" pageNumber="84" phylum="Mollusca" rank="genus">
<emphasis id="5F98EAAF380D0B4FFCFCFECB537EFEF0" box="[871,1032,265,291]" italics="true" pageId="83" pageNumber="84">Moelleriopsis</emphasis>
</taxonomicName>
on account of “the shape and sculpture of the shell”, at the same time renaming it
<taxonomicName id="AAEC4D3E380D0B4FFCA5FEEE53B1FE94" authority="Engl, 2012" authorityName="Engl" authorityYear="2012" box="[830,1223,300,327]" class="Gastropoda" family="Skeneidae" genus="Moelleriopsis" kingdom="Animalia" order="Seguenziida" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="poppei">
<emphasis id="5F98EAAF380D0B4FFCA5FEEE534CFE95" box="[830,1082,300,326]" italics="true" pageId="83" pageNumber="84">Moelleriopsis poppei</emphasis>
<bibRefCitation id="097D4B4C380D0B4FFBDFFEEE53B1FE94" author="Engl" box="[1092,1223,300,327]" pageId="83" pageNumber="84" refId="ref59972" refString="Engl W. 2012. Shells of Antarctica. Conchbooks, Hackenheim, Germany." type="book" year="2012">Engl, 2012</bibRefCitation>
</taxonomicName>
on account of secondary homonymy with the
<typeStatus id="B257881F380D0B4FFDA8FE925510FEB9" box="[563,614,336,362]" pageId="83" pageNumber="84">type</typeStatus>
species of the genus. However, details of protoconch morphology were not provided and so a comparison with the characteristic protoconch of
<taxonomicName id="AAEC4D3E380D0B4FFBD7FEB1539BFE5E" authorityName="Bush" authorityYear="1897" box="[1100,1261,371,397]" class="Gastropoda" family="Skeneidae" genus="Moelleriopsis" kingdom="Animalia" order="Seguenziida" pageId="83" pageNumber="84" phylum="Mollusca" rank="genus">
<emphasis id="5F98EAAF380D0B4FFBD7FEB1539BFE5E" box="[1100,1261,371,397]" italics="true" pageId="83" pageNumber="84">Moelleriopsis</emphasis>
</taxonomicName>
detailed by
<bibRefCitation id="097D4B4C380D0B4FFF26FE555616FE62" author="Waren A." box="[189,352,407,433]" pageId="83" pageNumber="84" pagination="149 - 248" refId="ref64085" refString="Waren A. 1992. New and little known &quot; skeneimorph &quot; gastropods from the Mediterranean Sea and the adjacent Atlantic Ocean. Bollettino Malacologico 27 (10 - 12): 149 - 248." type="journal article" year="1992">Warén (1992)</bibRefCitation>
is not possible. Further confirmation of this generic transfer is needed. The protoconch of
<taxonomicName id="AAEC4D3E380D0B4FFF44FE7856FEFE07" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[223,392,442,468]" class="Gastropoda" family="Skeneidae" genus="Mikro" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="crassus" status="sp. nov.">
<emphasis id="5F98EAAF380D0B4FFF44FE7856FEFE07" box="[223,392,442,468]" italics="true" pageId="83" pageNumber="84">Mikro crassus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4380D0B4FFE09FE795699FE06" box="[402,495,443,469]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
does not exhibit the spiral ribs and channelled suture distinctive of that of
<taxonomicName id="AAEC4D3E380D0B4FFF26FE1F5628FE24" authorityName="Bush" authorityYear="1897" box="[189,350,477,503]" class="Gastropoda" family="Skeneidae" genus="Moelleriopsis" kingdom="Animalia" order="Seguenziida" pageId="83" pageNumber="84" phylum="Mollusca" rank="genus">
<emphasis id="5F98EAAF380D0B4FFF26FE1F5628FE24" box="[189,350,477,503]" italics="true" pageId="83" pageNumber="84">Moelleriopsis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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