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<document id="5BE712EB28F2EAA8D386F767CA6C64BF" ID-DOI="10.11646/zootaxa.4576.3.9" ID-GBIF-Dataset="555f42f9-9135-45a6-b7aa-c0b4c064c7db" ID-ISSN="1175-5326" ID-Zenodo-Dep="3715838" ID-ZooBank="4994B0F7-3CB4-4DDE-99CB-BCD061D5730C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584603813141" checkinUser="plazi" docAuthor="Perkovsky, Evgeny E. &amp; Makarkin, Vladimir N." docDate="2019" docId="AA30746FFF8F854CFF0BFF0BFD12FAC3" docLanguage="en" docName="zootaxa.4576.3.9.pdf" docOrigin="Zootaxa 4576 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Succinoraphidia radioni Perkovsky &amp; Makarkin 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="574" masterDocId="56090C17FF8D8548FF9CFF9CFFEDFFE2" masterDocTitle="A new species of Succinoraphidia Aspöck &amp; Aspöck, 2004 (Raphidioptera: Raphidiidae) from the late Eocene Rovno amber, with venation characteristics of the genus" masterLastPageNumber="580" masterPageNumber="570" pageNumber="572" updateTime="1698730083632" updateUser="plazi">
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<mods:title id="2ACD3C44F5411E83A470C31C0BBD98A8">A new species of Succinoraphidia Aspöck &amp; Aspöck, 2004 (Raphidioptera: Raphidiidae) from the late Eocene Rovno amber, with venation characteristics of the genus</mods:title>
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<mods:namePart id="8C906B97F23DCA05B0E6ECFCAF5B041E">Perkovsky, Evgeny E.</mods:namePart>
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<mods:namePart id="B1D41B6BEEA7E14EE08D62B6DB2BCC12">Makarkin, Vladimir N.</mods:namePart>
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<mods:date id="F4EEE4B5B4C1896918699E342311CD7F">2019</mods:date>
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<treatment id="AA30746FFF8F854CFF0BFF0BFD12FAC3" ID-DOI="http://doi.org/10.5281/zenodo.3716803" ID-GBIF-Taxon="162639272" ID-Zenodo-Dep="3716803" LSID="urn:lsid:plazi:treatment:AA30746FFF8F854CFF0BFF0BFD12FAC3" httpUri="http://treatment.plazi.org/id/AA30746FFF8F854CFF0BFF0BFD12FAC3" lastPageId="4" lastPageNumber="574" pageId="2" pageNumber="572">
<subSubSection id="6A8396F2FF8F854AFF0BFF0BFDCEFF50" box="[151,547,151,178]" pageId="2" pageNumber="572" type="nomenclature">
<paragraph id="2226C579FF8F854AFF0BFF0BFDCEFF50" blockId="2.[151,547,151,211]" box="[151,547,151,178]" pageId="2" pageNumber="572">
<heading id="796E7215FF8F854AFF0BFF0BFDCEFF50" bold="true" box="[151,547,151,178]" fontSize="11" level="1" pageId="2" pageNumber="572" reason="1">
<emphasis id="10ED196BFF8F854AFF0BFF0BFDCEFF50" bold="true" box="[151,547,151,178]" pageId="2" pageNumber="572">
<taxonomicName id="E599BEFAFF8F854AFF0BFF0BFE53FF53" ID-CoL="7Q46" authority="Perkovsky &amp; Makarkin, 2019" authorityName="Perkovsky &amp; Makarkin" authorityYear="2019" box="[151,446,151,177]" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="10ED196BFF8F854AFF0BFF0BFE53FF53" bold="true" box="[151,446,151,177]" italics="true" pageId="2" pageNumber="572">Succinoraphidia radioni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BDEA410FF8F854AFE59FF04FDCEFF50" box="[453,547,152,178]" pageId="2" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="6A8396F2FF8F854AFF0BFF26FF14FF31" box="[151,249,186,211]" pageId="2" pageNumber="572" type="description">
<paragraph id="2226C579FF8F854AFF0BFF26FF14FF31" blockId="2.[151,547,151,211]" box="[151,249,186,211]" pageId="2" pageNumber="572">
<figureCitation id="BAA2D9FCFF8F854AFF0BFF26FF14FF31" box="[151,249,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,249,1956,1978]" captionStartId-1="4.[151,249,862,884]" captionStartId-2="5.[151,249,641,663]" captionStartId-3="5.[151,249,1945,1967]" captionTargetBox-0="[155,1432,195,1932]" captionTargetBox-1="[151,1436,200,818]" captionTargetBox-2="[151,1436,193,619]" captionTargetBox-3="[362,1211,747,1927]" captionTargetId-0="figure@3.[155,1432,195,1932]" captionTargetId-1="figure@4.[151,1436,200,818]" captionTargetId-2="figure@5.[151,1436,193,619]" captionTargetId-3="figure@5.[332,1250,731,1933]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionText-0="FIGURE 1. Succinoraphidia radioni sp. nov., holotype SIZK L-105 as preserved. A, left view; B, dorsal view. Scale bars = 2 mm." captionText-1="FIGURE 2. Succinoraphidia radioni sp. nov., holotype SIZK L-105, right wings. Scale bar = 2 mm." captionText-2="FIGURE 3. Succinoraphidia radioni sp. nov., holotype SIZK L-105, apex of ovipositor. A, ventral view; B, latero-caudal view. gs9, gonistyli of 9th segment; gx9, gonocoxites of 9th segment. Scale bars = 0.1 mm." captionText-3="FIGURE 4. Wing venation of Succinoraphidia radioni sp. nov., holotype SIZK L-105. A, left forewing; B, right forewing; C, left hind wing; D, right hind wing (A and C converted to standard view, with apex to the right). Fused veins are shown by red lines. Scale bar = 2 mm (all to scale)." figureDoi-0="http://doi.org/10.5281/zenodo.3715840" figureDoi-1="http://doi.org/10.5281/zenodo.3715842" figureDoi-2="http://doi.org/10.5281/zenodo.3715844" figureDoi-3="http://doi.org/10.5281/zenodo.3715846" httpUri-0="https://zenodo.org/record/3715840/files/figure.png" httpUri-1="https://zenodo.org/record/3715842/files/figure.png" httpUri-2="https://zenodo.org/record/3715844/files/figure.png" httpUri-3="https://zenodo.org/record/3715846/files/figure.png" pageId="2" pageNumber="572">Figs 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6A8396F2FF8F854AFF0BFF61FAD9FE62" pageId="2" pageNumber="572" type="diagnosis">
<paragraph id="2226C579FF8F854AFF0BFF61FAD9FE62" blockId="2.[151,1437,253,2029]" pageId="2" pageNumber="572">
<emphasis id="10ED196BFF8F854AFF0BFF61FEFCFEF4" bold="true" box="[151,273,253,278]" pageId="2" pageNumber="572">Diagnosis.</emphasis>
In both fore- and hind wings, ScP relatively long, length between termination of ScP and proximal end of pterostigma twice shorter than length of pterostigma [this length slightly longer than length of pterostigma in
<taxonomicName id="E599BEFAFF8F854AFA1BFEBEFF08FEBF" authorityName="Perkovsky &amp; Makarkin" authorityYear="2019" baseAuthorityName="Carpenter" baseAuthorityYear="1957" class="Insecta" family="Raphidiidae" genus="Raphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="baltica">
<emphasis id="10ED196BFF8F854AFA1BFEBEFF08FEBF" italics="true" pageId="2" pageNumber="572">S. baltica</emphasis>
</taxonomicName>
]; pterostigma distinctively paler in proximal part, darker in distal part [somewhat darker proximally in
<taxonomicName id="E599BEFAFF8F854AFA1BFED9FF14FE63" baseAuthorityName="Aspock &amp; Aspock" baseAuthorityYear="2004" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="exhibens">
<emphasis id="10ED196BFF8F854AFA1BFED9FF14FE63" italics="true" pageId="2" pageNumber="572">S. exhibens</emphasis>
</taxonomicName>
]; in hind wing, anterior trace of MP with three pectinate branches [two branches in
<taxonomicName id="E599BEFAFF8F854AFB39FEF5FAC9FE63" baseAuthorityName="Aspock &amp; Aspock" baseAuthorityYear="2004" box="[1189,1316,360,385]" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="exhibens">
<emphasis id="10ED196BFF8F854AFB39FEF5FAC9FE63" box="[1189,1316,360,385]" italics="true" pageId="2" pageNumber="572">S. exhibens</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="6A8396F2FF8F854AFF5BFE10FBE4F80F" pageId="2" pageNumber="572" type="description">
<paragraph id="2226C579FF8F854AFF5BFE10FE8FFE2A" blockId="2.[151,1437,253,2029]" pageId="2" pageNumber="572">
<emphasis id="10ED196BFF8F854AFF5BFE10FEB4FE47" bold="true" box="[199,345,396,421]" pageId="2" pageNumber="572">Description.</emphasis>
Female. Body poorly discernible. Prothorax relatively broad. Ovipositor long, with oval gonostyli (
<figureCitation id="BAA2D9FCFF8F854AFE92FE33FEBEFE2A" box="[270,339,431,456]" captionStart="FIGURE 3" captionStartId="5.[151,249,641,663]" captionTargetBox="[151,1436,193,619]" captionTargetId="figure@5.[151,1436,193,619]" captionTargetPageId="5" captionText="FIGURE 3. Succinoraphidia radioni sp. nov., holotype SIZK L-105, apex of ovipositor. A, ventral view; B, latero-caudal view. gs9, gonistyli of 9th segment; gx9, gonocoxites of 9th segment. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3715844" httpUri="https://zenodo.org/record/3715844/files/figure.png" pageId="2" pageNumber="572">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="2226C579FF8F854AFF5BFE4FFD11FB39" blockId="2.[151,1437,253,2029]" pageId="2" pageNumber="572">
Forewing oval,
<quantity id="E561689CFF8F854AFE1DFE48FE36FE09" box="[385,475,468,492]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="572" unit="mm" value="8.7">8.7 mm</quantity>
long,
<quantity id="E561689CFF8F854AFDB7FE48FD6BFE09" box="[555,646,468,492]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="572" unit="mm" value="2.7">2.7 mm</quantity>
wide. Costal space moderately dilated at 1/5 wing length; all four subcostal veinlets simple, widely spaced (additional incomplete veinlet between two basal veinlets in right wing). ScP relatively long, terminating on costal margin at about 0.55 wing length; length between termination of ScP and proximal end of pterostigma twice shorter than length of pterostigma. Subcostal space broad, with two crossveins: basal crossvein 1scp-r located distad seeming origin of M; distal crossvein 2scp-r at proximal end of pterostigma. Pterostigma elongate, paler proximad proximal branch of RP (light-brown), darker distad (brown). Anterior trace of RA not forked distally, strongly bent posteriorly after origin of distal branch, terminating at margin before wing apex; with two branch (veinlet): proximal branch located within pterostigma (part of pterostigma proximad this branch longer than distad); distal one nearly straight, located at distal end of pterostigma. RA space (between RA, RP) with two crossveins: 2ra-rp long, located slightly proximad proximal end of pterostigma; 3ra-rp shorter, located well distad end of pterostigma. RP originates far from wing base (at about 0.48 complete wing length), with two branches: RP1 deeply forked at outer gradate series of crossveins; RP2 simple. Anterior trace of RP not forked distally. One crossvein between branches of RP as part of outer gradate series (irp), connecting anterior branch of RP1, RP2 at its origin. Three crossveins between RP, MA: 1r-m, 2r-m connect anterior trace of RP, MA; 3r-m connects RP1 and anterior branch of MA. M appear fused basally with R for considerable distance, then actually fused with CuA for short distance; forked distad origin of RP. MA once deeply forked. MP fused with CuA for considerable distance, then pectinately branched, with two simple branches. Two intramedian crossveins form two
<emphasis id="10ED196BFF8F854AFF0BFBAFFF56FBAE" box="[151,187,1075,1100]" italics="true" pageId="2" pageNumber="572">doi</emphasis>
. Cu dividing into CuA, CuP rather far from wing base. CuA crossvein-like before fusion with M, perpendicular to Cu, then fused with M for very short distance, not forked distally. CuP simple, aligned with Cu and nearly parallel to distal part of CuA. Crossvein between CuA, CuP (icu) long; crossvein between CuP, AA1 short. Anal veins not forked distally; no crossvein between these. AA2, AA3 fused basally; then AA2 bent to AA1 and fused with it for short distance; free distal part of AA3 short.
</paragraph>
<paragraph id="2226C579FF8F854AFF5BFB79FBE4F80F" blockId="2.[151,1437,253,2029]" pageId="2" pageNumber="572">
Hind wing ca.
<quantity id="E561689CFF8F854AFEE9FB79FE23FB1C" box="[373,462,1253,1278]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="572" unit="mm" value="7.5">7.5 mm</quantity>
long,
<quantity id="E561689CFF8F854AFD86FB79FD9EFB1C" box="[538,627,1253,1278]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="572" unit="mm" value="2.6">2.6 mm</quantity>
wide. Costal space narrow; subcostal veinlets poorly discernible. ScP relatively long, terminating on costal margin at about 0.63 wing length; length between termination of ScP and proximal end of pterostigma nearly twice shorter than length of pterostigma. Subcostal space with one distal crossvein 2scp-r at proximal end of pterostigma. Pterostigma elongate, slightly lighter than in forewing; paler proximad proximal branch of RP, darker distad. Anterior trace of RA not forked distally, strongly bent posteriorly after origin of distal branch, terminating at margin before wing apex (slightly distad than in forewing); with two branch (veinlet): proximal branch located within pterostigma (part of pterostigma proximad this branch longer than distad); distal one nearly straight, located at distal end of pterostigma. RA space with three crossveins: 1ra-rp located nearly opposite to 2r-m; 2ra-rp located somewhat proximad proximal end of pterostigma; short 3ra-rp located well distad termination of pterostigma, connecting anterior traces of RA and RP. RP originates relatively close to wing base (at ca. 0.35 complete wing length), with two branches. RP1 deeply forked outer gradate series of crossveins; RP2 simple. One crossvein between branches of RP as part of outer gradate series. Three crossveins between RP, M/MA; 1r-m long, slightly sinuate, connecting anterior trace of RP and M near its seemingly origin; 2r-m connects RP1 and anterior trace of MA; 3r-m as part of outer gradate series connecting RP1 before fork and anterior branch of MA. M appears fused basally with R, forked into MA, MP distad well origin of RP. MA deeply forked distally; MP pectinately branched, with three branches. Two crossveins between MA and MP (1im, 2im) form two cells (
<emphasis id="10ED196BFF8F854AFED3F8BDFE6DF8D8" box="[335,384,1825,1850]" italics="true" pageId="2" pageNumber="572">1doi</emphasis>
,
<emphasis id="10ED196BFF8F854AFE0DF8BDFE2FF8D8" box="[401,450,1825,1850]" italics="true" pageId="2" pageNumber="572">2doi</emphasis>
), nearly equal in length; length/width ratio of
<emphasis id="10ED196BFF8F854AFC45F8BDFBE7F8D8" box="[985,1034,1825,1850]" italics="true" pageId="2" pageNumber="572">2doi</emphasis>
1.7 (left wing) to 2.1 (right wing). Two crossveins between M and Cu: basal 1m-cu moderately long, only slightly oblique, located near wing base; 2m-cu long, connecting MP and anterior branch of CuA. CuA deeply forked. CuP simple. One long crossvein between CuA, CuP (icu), located far distad 1m-cu. Cu and AA1 fully separate; one short crossvein connecting these located between 1m-cu, 2m-cu. AA1, AA2 fused proximally; their distal (free) parts simple. AA3 short, simple (poorly discernible). One long crossvein between AA1+AA2 and AA3.
</paragraph>
</subSubSection>
<caption id="76E695F1FF8E854BFF0BF838FF2BF83B" ID-DOI="http://doi.org/10.5281/zenodo.3715840" ID-Zenodo-Dep="3715840" httpUri="https://zenodo.org/record/3715840/files/figure.png" pageId="3" pageNumber="573" startId="3.[151,249,1956,1978]" targetBox="[155,1432,195,1932]" targetPageId="3">
<paragraph id="2226C579FF8E854BFF0BF838FF2BF83B" blockId="3.[151,1435,1956,2009]" pageId="3" pageNumber="573">
<emphasis id="10ED196BFF8E854BFF0BF838FEE0F858" bold="true" box="[151,269,1956,1978]" pageId="3" pageNumber="573">FIGURE 1</emphasis>
.
<taxonomicName id="E599BEFAFF8E854BFE85F838FDFFF858" authority="Perkovsky &amp; Makarkin, 2019" authorityName="Perkovsky &amp; Makarkin" authorityYear="2019" box="[281,530,1956,1978]" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="10ED196BFF8E854BFE85F838FDFFF858" box="[281,530,1956,1978]" italics="true" pageId="3" pageNumber="573">Succinoraphidia radioni</emphasis>
</taxonomicName>
<emphasis id="10ED196BFF8E854BFD85F838FD84F858" bold="true" box="[537,617,1956,1978]" pageId="3" pageNumber="573">
<taxonomicNameLabel id="0BDEA410FF8E854BFD85F838FD84F858" box="[537,617,1956,1978]" pageId="3" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype SIZK L-105 as preserved. A, left view; B, dorsal view. Scale bars = 2 mm.
</paragraph>
</caption>
<caption id="76E695F1FF89854CFF0BFCC2FB7BFC96" ID-DOI="http://doi.org/10.5281/zenodo.3715842" ID-Zenodo-Dep="3715842" box="[151,1174,862,884]" httpUri="https://zenodo.org/record/3715842/files/figure.png" pageId="4" pageNumber="574" startId="4.[151,249,862,884]" targetBox="[151,1436,200,818]" targetPageId="4">
<paragraph id="2226C579FF89854CFF0BFCC2FB7BFC96" blockId="4.[151,1174,862,884]" box="[151,1174,862,884]" pageId="4" pageNumber="574">
<emphasis id="10ED196BFF89854CFF0BFCC2FEE1FC96" bold="true" box="[151,268,862,884]" pageId="4" pageNumber="574">FIGURE 2</emphasis>
.
<taxonomicName id="E599BEFAFF89854CFE84FCC2FDFCFC96" authority="Perkovsky &amp; Makarkin, 2019" authorityName="Perkovsky &amp; Makarkin" authorityYear="2019" box="[280,529,862,884]" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="10ED196BFF89854CFE84FCC2FDFCFC96" box="[280,529,862,884]" italics="true" pageId="4" pageNumber="574">Succinoraphidia radioni</emphasis>
</taxonomicName>
<emphasis id="10ED196BFF89854CFD8BFCC2FD8AFC96" bold="true" box="[535,615,862,884]" pageId="4" pageNumber="574">
<taxonomicNameLabel id="0BDEA410FF89854CFD8BFCC2FD8AFC96" box="[535,615,862,884]" pageId="4" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype SIZK L-105, right wings. Scale bar = 2 mm.
</paragraph>
</caption>
<subSubSection id="6A8396F2FF89854CFF5BFC3CFB1FFC5B" box="[199,1266,928,953]" pageId="4" pageNumber="574" type="etymology">
<paragraph id="2226C579FF89854CFF5BFC3CFB1FFC5B" blockId="4.[151,1437,928,1313]" box="[199,1266,928,953]" pageId="4" pageNumber="574">
<emphasis id="10ED196BFF89854CFF5BFC3CFEABFC5B" bold="true" box="[199,326,928,953]" pageId="4" pageNumber="574">Etymology</emphasis>
. From the first name of Radion F. Khomich, father of the collector of the specimen.
</paragraph>
</subSubSection>
<subSubSection id="6A8396F2FF89854CFF5BFC58FA70FBAA" pageId="4" pageNumber="574" type="materials_examined">
<paragraph id="2226C579FF89854CFF5BFC58FF06FBC7" blockId="4.[151,1437,928,1313]" pageId="4" pageNumber="574">
<emphasis id="10ED196BFF89854CFF5BFC58FE91FC3F" bold="true" box="[199,380,964,989]" pageId="4" pageNumber="574">Type material.</emphasis>
<materialsCitation id="92F1CF24FF89854CFE16FC58FF06FBC7" ID-GBIF-Occurrence="2578498299" collectionCode="SIZK" collectorName="Specimen No. &amp; Nikolai R. Khomich" pageId="4" pageNumber="574" specimenCount="1" typeStatus="holotype">
<typeStatus id="FD227BDBFF89854CFE16FC58FE1AFC3F" box="[394,503,964,989]" pageId="4" pageNumber="574" type="holotype">Holotype</typeStatus>
:
<collectorName id="8F6CA0AFFF89854CFD94FC58FD43FC3E" box="[520,686,964,989]" pageId="4" pageNumber="574">Specimen No.</collectorName>
<collectionCode id="44885DBCFF89854CFD20FC59FD17FC3F" box="[700,762,965,989]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="4" pageNumber="574">SIZK</collectionCode>
L-105, collected by
<collectorName id="8F6CA0AFFF89854CFB9CFC58FB1FFC3F" box="[1024,1266,964,989]" pageId="4" pageNumber="574">Nikolai R. Khomich</collectorName>
and currently deposited in his collection, but will be ultimately deposited in
<collectionCode id="44885DBCFF89854CFCE2FC75FC2DFBE3" box="[894,960,1001,1025]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="4" pageNumber="574">SIZK</collectionCode>
. A poorly-preserved, slightly incomplete female.
</materialsCitation>
</paragraph>
<paragraph id="2226C579FF89854CFF5BFBACFA70FBAA" blockId="4.[151,1437,928,1313]" box="[199,1437,1072,1097]" pageId="4" pageNumber="574">
<emphasis id="10ED196BFF89854CFF5BFBACFE2BFBAB" bold="true" box="[199,454,1072,1097]" pageId="4" pageNumber="574">Locality and horizon.</emphasis>
<materialsCitation id="92F1CF24FF89854CFE50FBADFA74FBAA" ID-GBIF-Occurrence="2578498298" box="[460,1433,1072,1097]" country="Ukraine" location="Voronki in Vladimirets District" pageId="4" pageNumber="574" specimenCount="1" stateProvince="Rivne">
<collectingRegion id="E05D0B9BFF89854CFE50FBADFDFAFBAA" box="[460,535,1073,1096]" country="Ukraine" name="Rivne" pageId="4" pageNumber="574">Rovno</collectingRegion>
amber (
<location id="274693A2FF89854CFDEDFBACFC23FBAB" LSID="urn:lsid:plazi:treatment:AA30746FFF8F854CFF0BFF0BFD12FAC3:274693A2FF89854CFDEDFBACFC23FBAB" box="[625,974,1072,1097]" country="Ukraine" name="Voronki in Vladimirets District" pageId="4" pageNumber="574" stateProvince="Rivne">Voronki in Vladimirets District</location>
of
<collectingRegion id="E05D0B9BFF89854CFC6EFBADFBD0FBAA" box="[1010,1085,1073,1096]" country="Ukraine" name="Rivne" pageId="4" pageNumber="574">Rovno</collectingRegion>
Region,
<collectingCountry id="5A8E85E9FF89854CFB3EFBACFAECFBAB" box="[1186,1281,1072,1097]" name="Ukraine" pageId="4" pageNumber="574">Ukraine</collectingCountry>
); late Eocene
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6A8396F2FF89854CFF5BFBC8FD12FAC3" pageId="4" pageNumber="574" type="discussion">
<paragraph id="2226C579FF89854CFF5BFBC8FD12FAC3" blockId="4.[151,1437,928,1313]" pageId="4" pageNumber="574">
<emphasis id="10ED196BFF89854CFF5BFBC8FED6FB8F" bold="true" box="[199,315,1108,1133]" pageId="4" pageNumber="574">Remarks.</emphasis>
The new species easily differs from
<taxonomicName id="E599BEFAFF89854CFD7CFBC9FCA6FB8F" authorityName="Perkovsky &amp; Makarkin" authorityYear="2019" baseAuthorityName="Carpenter" baseAuthorityYear="1957" box="[736,843,1108,1133]" class="Insecta" family="Raphidiidae" genus="Raphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="baltica">
<emphasis id="10ED196BFF89854CFD7CFBC9FCA6FB8F" box="[736,843,1108,1133]" italics="true" pageId="4" pageNumber="574">S. baltica</emphasis>
</taxonomicName>
by the longer ScP. Unfortunately, the venation of
<taxonomicName id="E599BEFAFF89854CFA1BFBC9FF14FB73" baseAuthorityName="Aspock &amp; Aspock" baseAuthorityYear="2004" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="exhibens">
<emphasis id="10ED196BFF89854CFA1BFBC9FA78FB8E" box="[1415,1429,1109,1132]" italics="true" pageId="4" pageNumber="574">S</emphasis>
.
<emphasis id="10ED196BFF89854CFF0BFBE4FF14FB73" box="[151,249,1144,1169]" italics="true" pageId="4" pageNumber="574">exhibens</emphasis>
</taxonomicName>
is incompletely described. However, the photograph of the
<typeStatus id="FD227BDBFF89854CFC03FBE4FBECFB73" box="[927,1025,1144,1169]" pageId="4" pageNumber="574" type="holotype">holotype</typeStatus>
and the drawing of the basal part of the hind wing (see
<bibRefCitation id="4608B888FF89854CFEF7FB00FD90FB57" author="Aspock, U. &amp; Aspock, H." box="[363,637,1180,1205]" pageId="4" pageNumber="574" pagination="11 - 19" refId="ref4869" refString="Aspock, U. &amp; Aspock, H. (2004) Two significant new snakeflies from Baltic amber, with discussion on autapomorphies of the order and its included taxa (Raphidioptera). Systematic Entomology, 29, 11 - 19. https: // doi. org / 10.1111 / j. 1365 - 3113.2004.00245. x" type="journal article" year="2004">Aspöck &amp; Aspöck 2004</bibRefCitation>
:
<figureCitation id="BAA2D9FCFF89854CFD1BFB00FD21FB57" box="[647,716,1180,1205]" captionStart="FIGURE 1" captionStartId="3.[151,249,1956,1978]" captionTargetBox="[155,1432,195,1932]" captionTargetId="figure@3.[155,1432,195,1932]" captionTargetPageId="3" captionText="FIGURE 1. Succinoraphidia radioni sp. nov., holotype SIZK L-105 as preserved. A, left view; B, dorsal view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3715840" httpUri="https://zenodo.org/record/3715840/files/figure.png" pageId="4" pageNumber="574">Figs 1</figureCitation>
,
<figureCitation id="BAA2D9FCFF89854CFD46FB00FD05FB57" box="[730,744,1180,1205]" captionStart="FIGURE 5" captionStartId="7.[151,249,1707,1729]" captionTargetBox="[151,1436,1119,1685]" captionTargetId="figure@7.[151,1436,1119,1685]" captionTargetPageId="7" captionText="FIGURE 5. Succinoraphidia baltica (Carpenter, 1957), holotype MCZ 5122. A, ventral view; B, dorsal view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3715850" httpUri="https://zenodo.org/record/3715850/files/figure.png" pageId="4" pageNumber="574">5</figureCitation>
) show that the venation of
<taxonomicName id="E599BEFAFF89854CFB85FB01FB7AFB57" baseAuthorityName="Aspock &amp; Aspock" baseAuthorityYear="2004" box="[1049,1175,1180,1205]" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="exhibens">
<emphasis id="10ED196BFF89854CFB85FB01FBCAFB56" box="[1049,1063,1181,1204]" italics="true" pageId="4" pageNumber="574">S</emphasis>
.
<emphasis id="10ED196BFF89854CFBA9FB00FB7AFB57" box="[1077,1175,1180,1205]" italics="true" pageId="4" pageNumber="574">exhibens</emphasis>
</taxonomicName>
differs from that of the new species by the number of MP branches and the presence of additional long crossvein between MP and CuA in the hind wing, as well as by the coloration of the pterostigma (this is “mainly yellowish, proximally somewhat darker” in
<taxonomicName id="E599BEFAFF89854CFE92FA95FD1FFAC3" authority=": Aspock &amp; Aspock 2004, p. 12" authorityName=": Aspock &amp; Aspock" authorityPageNumber="12" authorityYear="2004" box="[270,754,1288,1313]" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="exhibens">
<emphasis id="10ED196BFF89854CFE92FA95FEF1FAC2" box="[270,284,1289,1312]" italics="true" pageId="4" pageNumber="574">S</emphasis>
.
<emphasis id="10ED196BFF89854CFEB7FA94FE60FAC3" box="[299,397,1288,1313]" italics="true" pageId="4" pageNumber="574">exhibens</emphasis>
:
<bibRefCitation id="4608B888FF89854CFE07FA94FD41FAC3" author="Aspock, U. &amp; Aspock, H." box="[411,684,1288,1313]" pageId="4" pageNumber="574" pagination="11 - 19" refId="ref4869" refString="Aspock, U. &amp; Aspock, H. (2004) Two significant new snakeflies from Baltic amber, with discussion on autapomorphies of the order and its included taxa (Raphidioptera). Systematic Entomology, 29, 11 - 19. https: // doi. org / 10.1111 / j. 1365 - 3113.2004.00245. x" type="journal article" year="2004">Aspöck &amp; Aspöck 2004</bibRefCitation>
, p. 12
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>