161 lines
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161 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.714.11503" ID-GBIF-Dataset="82cad059-f2e2-473c-9905-f2bcd7727c2a" ID-PMC="PMC5740425" ID-Pensoft-Pub="1313-2970-714-129" ID-PubMed="29290702" ID-ZBK="82FF5425D41047E5AABC2C9FCE561713" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-714-129" ModsDocOrigin="ZooKeys 714" ModsDocTitle="New species of Metachela Coquillett (Diptera, Empididae) from the Atlantic Forest, Brazil and a key to the Neotropical species" checkinTime="1510066980661" checkinUser="pensoft" docAuthor="Camara, Josenir Teixeira & Rafael, Jose Albertino" docDate="2017" docId="AA24706F621F76ACC85CF4B8C84D0603" docLanguage="en" docName="ZooKeys 714: 129-140" docOrigin="ZooKeys 714" docSource="http://dx.doi.org/10.3897/zookeys.714.11503" docTitle="Metachela danitakiyae Camara & Rafael, 2017, sp. n." docType="treatment" docUuid="D957CD88-DFBC-4544-A465-1BB95F63082F" docUuidSource="ZooBank" docVersion="5" lastPageNumber="133" masterDocId="FFAC1421FFF74A65DC20F416FFB0DA5E" masterDocTitle="New species of Metachela Coquillett (Diptera, Empididae) from the Atlantic Forest, Brazil and a key to the Neotropical species" masterLastPageNumber="140" masterPageNumber="129" pageNumber="132" updateTime="1668165098669" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of Metachela Coquillett (Diptera, Empididae) from the Atlantic Forest, Brazil and a key to the Neotropical species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Camara, Josenir Teixeira</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rafael, Jose Albertino</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>714</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>140</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.714.11503</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.714.11503</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-714-129</mods:identifier>
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<mods:identifier type="ZBK">82FF5425D41047E5AABC2C9FCE561713</mods:identifier>
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<mods:identifier type="ZooBank">82FF5425D41047E5AABC2C9FCE561713</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="137252914" LSID="urn:lsid:zoobank.org:act:D957CD88-DFBC-4544-A465-1BB95F63082F" httpUri="http://treatment.plazi.org/id/AA24706F621F76ACC85CF4B8C84D0603" lastPageId="4" lastPageNumber="133" pageId="3" pageNumber="132">
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<subSubSection pageId="3" pageNumber="132" type="nomenclature">
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<paragraph pageId="3" pageNumber="132">
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<taxonomicName LSID="http://zoobank.org/D957CD88-DFBC-4544-A465-1BB95F63082F" class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela danitakiyae" order="Diptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="danitakiyae">Metachela danitakiyae</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="132">sp. n.</taxonomicNameLabel>
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Figs 1-7, 8-11
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="132" type="type-locality">
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<paragraph pageId="3" pageNumber="132">Type-locality.</paragraph>
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<paragraph pageId="3" pageNumber="132">
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BRAZIL, Rio de Janeiro: Itatiaia, Parque Nacional de Itatiaia,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.42739">
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22°25'38.6"
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<normalizedToken originalValue="S–">S-</normalizedToken>
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</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-44.619358">44°37'9.7"W</geoCoordinate>
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, 1140 m,
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="132" type="type-specimen">
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<paragraph pageId="3" pageNumber="132">Type-specimen.</paragraph>
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<paragraph pageId="3" pageNumber="132">
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Holotype male, pinned, not dissected: "BRAZIL, RJ [Rio de Janeiro], Itatiaia, Parque Nacional de Itatiaia.
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<normalizedToken originalValue="Córrego">Corrego</normalizedToken>
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Maromba, abaixo da Cachoeira
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<normalizedToken originalValue="Véu">Veu</normalizedToken>
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de Noiva. Malaise trap,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.42739">
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22°25'38.6"
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<normalizedToken originalValue="S–">S-</normalizedToken>
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</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-44.619358">44°37'9.7"W</geoCoordinate>
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, 1140 m. 10.
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<normalizedToken originalValue="i–">i-</normalizedToken>
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02.II.2015. D.M. Takiya, A.P.M. Santos & M.F.
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<normalizedToken originalValue="Monné”">Monne"</normalizedToken>
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(INPA). Paratypes. Same data as holotype (8 males, 2 females, INPA, 5 males, 2 females, MNRJ, 3 males, 4 females, MZUSP). BRAZIL, MG[Minas Gerais], Alto
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<normalizedToken originalValue="Caparaó">Caparao</normalizedToken>
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, Parque Nacional do
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<normalizedToken originalValue="Caparaó">Caparao</normalizedToken>
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, Vale Verde. Malaise, 17-20.i.2014.
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-20.419361">
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20°25'09.7"
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<normalizedToken originalValue="S–">S-</normalizedToken>
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</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-41.846386">41°50'47"W</geoCoordinate>
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, 1364m. J.L. Nissimian & A.P.M. Santos. (2 males, 1 female, INPA).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="133" pageId="3" pageNumber="132" type="diagnosis">
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<paragraph pageId="3" pageNumber="132">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="132">
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Scutum with a mid-longitudinal brown stripe; vein R2+3 fused to R1 (Figs 2 and 9); male cercus arched, in dorsal view, acute at extreme apex (Figs 3, 4); epandrium with a dorsoapical sinus (Fig. 5); hypandrium membranous midventrally on basal
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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and sclerotized apically (Fig. 7); female tergite 10 with bilobate posterior margin (Fig. 11); female sternite 8 elongate, concave on anterior margin, membranous on apical half (Fig. 11).
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</paragraph>
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<paragraph pageId="3" pageNumber="132">
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Male (Fig. 1). Head (Fig. 1): Dark brown to black, setae whitish. Ocellar triangle with two pairs of proclinate bristles, anterior pair stouter. Eyes iridescent black, separated on face. Occiput with scattered fine setae. Mouth parts yellow; proboscis short, slightly curved and with yellow setae. Antenna yellow, with scape and pedicel bearing distinct short ventral setulae; postpedicel nearly 2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; stylus very short, ~ 0.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as postpedicel.
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</paragraph>
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<caption pageId="3" pageNumber="132">
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<paragraph pageId="3" pageNumber="132">
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Figures 1-7.
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<taxonomicName class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela danitakiyae" order="Diptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="danitakiyae">Metachela danitakiyae</taxonomicName>
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sp. n., 1-2 holotype male 3-7 paratype ♂ 1 Habitus, lateral view 2 Wing 3 Cercus, dorsal view 4 Subepandrial sclerite, epandrium and cercus, anteroventral view 5 Cercus, epandrium and hypandrium, lateral view 6 Phallus, lateral view 7 Hypandrium, ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="3" pageNumber="132">Thorax (Fig. 1): Elongate, slightly arched dorsally; scutum yellowish except for mid-longitudinal brown stripe, wider posteriorly, and brownish posterolateral spot above wing base; scutum with very small and fine yellow setae except for one notopleural, one postalar, and two pairs of small parallel scutellars; scutellum and mediotergite brown.</paragraph>
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<paragraph pageId="3" pageNumber="132">
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Legs (Fig. 1): Yellow, except fore tibia with narrow anteroventral brown stripe distally, and hind femur at distal 2/3 and tarsomeres 4-5 brown. Fore coxa as long as distance between fore- and mid coxae, 4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, with some dorso-apical pale setae. Fore femur approximately 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as fore coxa, 4.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, with anteroventral row of 3-5 spines, anteroventral row of 6-8 denticles, and posteroventral row of 18-20 denticles, with basal ones stouter; anteroventral row of denticles placed on distal half and posteroventral row restricted to distal 0.9, and both rows without apical discontinuity and diverging at apex. Fore tibia approximately 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as fore femur; with decumbent short pale bristles dorsally, more densely distributed apically. Mid femur with two anteroventral and 12 posteroventral spines, basal pair stouter. Hind legs slender, with fine setae, except hind tibia with dorsoapical comb of short setae.
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</paragraph>
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<paragraph pageId="3" pageNumber="132">
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Wings (Fig. 2): Membranous, veins yellowish; vein R2+3 short, fused to vein R1; R4+5 fork angle around 70°; R5 and M1 slightly divergent at extreme apex; cell bm+dm ending beyond apex of R1, ~1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as cell br; cup cell closed. Halter whitish yellow.
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</paragraph>
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<paragraph pageId="4" pageNumber="133">
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<pageBreakToken pageId="4" pageNumber="133" start="start">Abdomen</pageBreakToken>
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(Fig. 1): Tergites and sternites membranous, yellow, except anterior margin of all tergites and sternite 8 brownish; sternite 8 strongly sclerotized, U-shaped posteriorly.
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</paragraph>
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<paragraph pageId="4" pageNumber="133">Male terminalia: Brown. Cercus arched, in dorsal view (Figs 3, 4) narrower on basal 1/4, expanded apically, extending beyond epandrium apex (Fig. 5); left and right cerci closely approximated anterodorsally (Fig. 3); distinctly setose. Epandrium subrectangular, with a posterodorsal sinus (Fig. 5), with distinct strong setae on outer face. Hypandrium membranous midventrally on basal 3/4, sclerotized and fused posteriorly (Fig. 7); gonocoxal apodeme projecting anteriorly as a small protuberance (Fig. 5). Subepandrial sclerite subrectangular, more sclerotized basally and laterally (Fig. 4). Phallus strongly sclerotized, except less sclerotized apically (Fig. 6), abruptly pointed apically. Ejaculatory apodeme present. Holotype: body length. 3.3 mm; wing length. 2.7 mm.</paragraph>
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<paragraph pageId="4" pageNumber="133">Female (Figs 8, 9). Similar to male. Terminalia: Tergite 7 brown, shorter than tergite 8 (Figs 10, 11); tergite 8 brown, subtrapezoidal, with anterior margin slightly concave in dorsal view (Fig. 11); tergite 10 slightly light brown, bilobate on posterior margin (Fig. 11). Cercus brown, apex pale (Fig. 10). Sternite 7 brown, with small median projection on anterior margin (Fig. 11); sternite 8 dark brown on basal 2/3 and with pale apex, elongate, concave on anterior margin (Fig. 11); sternite 10 very narrow, v-shaped.</paragraph>
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<caption pageId="4" pageNumber="133">
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<paragraph pageId="4" pageNumber="133">
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Figures 8-11.
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<taxonomicName class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela danitakiyae" order="Diptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="danitakiyae">Metachela danitakiyae</taxonomicName>
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sp. n., paratype female from Itatiaia. 8 Habitus, lateral view 9 Wing 10 Segments 7-11, lateral view 11 Tergites 7, 8, and 10, dorsal view, sternites 7 and 8, ventral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="133" type="geographical records">
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<paragraph pageId="4" pageNumber="133">Geographical records.</paragraph>
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<paragraph pageId="4" pageNumber="133">Brazil (Minas Gerais and Rio de Janeiro states).</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="133" type="etymology">
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<paragraph pageId="4" pageNumber="133">Etymology.</paragraph>
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<paragraph pageId="4" pageNumber="133">The specific epithet is a tribute to Daniela Maeda Takiya, friend of the authors and collector of the specimens.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="133" type="remarks">
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<paragraph pageId="4" pageNumber="133">Remarks.</paragraph>
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<paragraph pageId="4" pageNumber="133">
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<taxonomicName class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela danitakiyae" order="Diptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="danitakiyae">Metachela danitakiyae</taxonomicName>
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sp. n. differs from other species by the elongate male cercus, narrower on the apical 1/4 (usually short in other species, if elongate then with broad apex); epandrium with a posterodorsal sinus (without sinus in other species).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |