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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e90570" ID-Pensoft-Pub="1864-8312-80-649" ID-Pensoft-UUID="FAE6DFEF69735531A4CBA275E078320D" ID-ZooBank="AD0B1CC262414460B6B888E6B95F20D7" ModsDocID="1864-8312-80-649" checkinTime="1671114540108" checkinUser="pensoft" docAuthor="Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S. &amp; Harms, Danilo" docDate="2022" docId="6A3BBE243EC9550B9814DF9A24D5867C" docLanguage="en" docName="ArthropodSystPhyl 80: 649-691" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-12-14" docSource="http://dx.doi.org/10.3897/asp.80.e90570" docTitle="Antsirananaella faulstichi Lorenz &amp; Loria &amp; Harvey &amp; Harms 2022, sp. nov." docType="treatment" docVersion="1" id="FAE6DFEF69735531A4CBA275E078320D" lastPageNumber="649" masterDocId="FAE6DFEF69735531A4CBA275E078320D" masterDocTitle="The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the &quot; hottest &quot; biodiversity hotspots" masterLastPageNumber="691" masterPageNumber="649" pageNumber="649" updateTime="1671114540108" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the &quot; hottest &quot; biodiversity hotspots</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lorenz, Michelle</mods:namePart>
<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany &amp; University of Hamburg, Biology Department, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">michellelorenz@gmx.net</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Loria, Stephanie F.</mods:namePart>
<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
<mods:affiliation>Western Australian Museum, Collections &amp; Research, 49 Kew Street, Welshpool, WA 6106, Australia &amp; University of Western Australia, School of Biological Sciences, Crawley WA 6009, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harms, Danilo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier>
<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>649</mods:start>
<mods:end>691</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e90570</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e90570</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-649</mods:identifier>
<mods:identifier type="ZooBank">AD0B1CC262414460B6B888E6B95F20D7</mods:identifier>
<mods:identifier type="Pensoft-UUID">FAE6DFEF69735531A4CBA275E078320D</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:NomEnClAturAlACts/ABF2723E-BF6D-4D6E-861C-3D31B73DD563" httpUri="http://treatment.plazi.org/id/6A3BBE243EC9550B9814DF9A24D5867C" lastPageNumber="649" pageId="0" pageNumber="649">
<subSubSection pageId="0" pageNumber="649" type="nomenclature">
<paragraph pageId="0" pageNumber="649">
<taxonomicName LSID="https://zoobank.org/NomenclaturalActs/ABF2723E-BF6D-4D6E-861C-3D31B73DD563" authority="Lorenz &amp; Loria &amp; Harvey &amp; Harms, 2022" authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Antsirananaella faulstichi" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="faulstichi" status="sp. nov.">Antsirananaella faulstichi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="649">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="type material">
<paragraph pageId="0" pageNumber="649">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="649">
MADAGASCAR:
<taxonomicName genus="Antsiranana" lsidName="Antsiranana" pageId="0" pageNumber="649" rank="genus">Antsiranana</taxonomicName>
Prov. [Diana Region]: Holotype ♂ (CAS 9071843 [BLF3012]), allotype ♀ (ZMH-A0016680), paratypes: 7 ♂, 1 ♀ (CAS 9071843 [BLF3012]), 3 ♂ (ZMH-A0016681 - ZMH-A0016683), Reserve special
<normalizedToken originalValue="dAnkarana">d'Ankarana</normalizedToken>
,
<geoCoordinate degrees="12" direction="south" minutes="51" orientation="latitude" precision="15" seconds="49" value="-12.863611">12°5149″S</geoCoordinate>
<geoCoordinate degrees="49" direction="east" minutes="13" orientation="longitude" precision="15" seconds="33" value="49.225834">49°1333″E</geoCoordinate>
, December 2003, B.L. Fisher.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="materials_examined">
<paragraph pageId="0" pageNumber="649">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="649">
<materialsCitation accessionNumber="BLF11400" collectingDate="2001-02-10" collectingDateMax="2004-12-05" collectingDateMin="2001-02-10" collectorName="B. L. Fisher, Fisher, Griswold" country="MADAGASCAR" county="Reserve Analamerana" latitude="-12.908889" location="Reserve Speciale de l'Ankarana" longLatPrecision="21" longitude="49.109722" municipality="Reserve Speciale de l'Ankarana" specimenCount="14" specimenCount-female="5" specimenCount-male="9" stateProvince="Antsiranana Prov.">
<collectingCountry name="Madagascar">MADAGASCAR</collectingCountry>
:
<collectingRegion country="Madagascar" name="Antsiranana">
<taxonomicName genus="Antsiranana" lsidName="Antsiranana" pageId="0" pageNumber="649" rank="genus">Antsiranana</taxonomicName>
Prov.
</collectingRegion>
[Diana Region]:
<specimenCount type="male">8 ♂</specimenCount>
,
<specimenCount type="female">4 ♀</specimenCount>
, 7 nymphs (CAS 9072436 [
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/BLF11400">BLF11400</accessionNumber>
]),
<collectingCounty>Reserve Analamerana</collectingCounty>
,
<geoCoordinate degrees="12" direction="south" minutes="44" orientation="latitude" precision="15" seconds="48" value="-12.746667">12°4448″S</geoCoordinate>
<geoCoordinate degrees="49" direction="east" minutes="29" orientation="longitude" precision="15" seconds="41" value="49.494724">49°2941″E</geoCoordinate>
,
<collectingDate value="2004-12-05">5 December 2004</collectingDate>
,
<collectorName>B.L. Fisher</collectorName>
;
<specimenCount type="male">1 ♂</specimenCount>
(CAS 9008191 [BLF2972]),
<collectingMunicipality>
Reserve Speciale de
<normalizedToken originalValue="lAnkarana">l'Ankarana</normalizedToken>
</collectingMunicipality>
,
<geoCoordinate degrees="12" direction="south" minutes="54" orientation="latitude" precision="15" seconds="32" value="-12.908889">12°5432″S</geoCoordinate>
<geoCoordinate degrees="49" direction="east" minutes="06" orientation="longitude" precision="15" seconds="35" value="49.109722">49°0635″E</geoCoordinate>
,
<collectingDate value="2001-02-10" valueMax="2001-02-16" valueMin="2001-02-10">10-16 February 2001</collectingDate>
,
<collectorName>Fisher</collectorName>
,
<collectorName>Griswold</collectorName>
et al.;
<specimenCount type="female">1 ♀</specimenCount>
(CAS 9071838 [BLF2858]),
<location LSID="urn:lsid:plazi:treatment:6A3BBE243EC9550B9814DF9A24D5867C:E064865A2B4F4CC866901ECDBE2366D1" country="MADAGASCAR" county="Reserve Analamerana" latitude="-12.908889" longLatPrecision="21" longitude="49.109722" municipality="Reserve Speciale de l'Ankarana" name="Reserve Speciale de l'Ankarana" stateProvince="Antsiranana Prov.">
Reserve Speciale de
<normalizedToken originalValue="lAnkarana">l'Ankarana</normalizedToken>
</location>
;
<geoCoordinate degrees="12" direction="south" minutes="54" orientation="latitude" precision="15" seconds="32" value="-12.908889">12°5432″S</geoCoordinate>
<geoCoordinate degrees="49" direction="east" minutes="06" orientation="longitude" precision="15" seconds="35" value="49.109722">49°0635″E</geoCoordinate>
,
<collectingDate value="2001-02-10" valueMax="2001-02-16" valueMin="2001-02-10">10-16 February 2001</collectingDate>
,
<collectorName>Fisher</collectorName>
,
<collectorName>Griswold</collectorName>
et al
</materialsCitation>
.;
<materialsCitation collectingDate="2003-12-03" collectorName="de Binara, B. L. Fisher" country="Madagascar" latitude="-13.263333" location="Foret de Binara" longLatPrecision="21" longitude="49.603333" specimenCount="1" stateProvince="Antsiranana Prov.">
<collectingRegion country="Madagascar" name="Antsiranana">
<taxonomicName genus="Antsiranana" lsidName="Antsiranana" pageId="0" pageNumber="649" rank="genus">Antsiranana</taxonomicName>
Prov.
</collectingRegion>
[Sava Region]: 1 nymph (CAS 9072439 [BLF9656]),
<location LSID="urn:lsid:plazi:treatment:6A3BBE243EC9550B9814DF9A24D5867C:78589C86EA1483A456BAE1EE8DD4628B" country="Madagascar" latitude="-13.263333" longLatPrecision="21" longitude="49.603333" name="Foret de Binara" stateProvince="Antsiranana Prov.">
Foret
<collectorName>de Binara</collectorName>
</location>
,
<geoCoordinate degrees="13" direction="south" minutes="15" orientation="latitude" precision="15" seconds="48" value="-13.263333">13°1548″S</geoCoordinate>
<geoCoordinate degrees="49" direction="east" minutes="36" orientation="longitude" precision="15" seconds="12" value="49.603333">49°3612″E</geoCoordinate>
,
<collectingDate value="2003-12-03">3 December 2003</collectingDate>
,
<collectorName>B.L. Fisher</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="diagnosis">
<paragraph pageId="0" pageNumber="649">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="649">
Smallest body sizes of all species of
<taxonomicName authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
(e.g., body length 1.80/2.17 ♂/♀ versus 1.94-2.17/ 2.21-2.47 ♂/♀; carapace length 0.55/0.65 ♂/♀ versus 0.58-0.69/ 0.70-0.77 ♂/♀). Unique cheotaxy:
<emphasis italics="true" pageId="0" pageNumber="649">sb</emphasis>
not on the same sagittal level as
<emphasis italics="true" pageId="0" pageNumber="649">est</emphasis>
but slightly more distal (on the same sagittal level in
<taxonomicName lsidName="A. marlae" pageId="0" pageNumber="649" rank="species" species="marlae">
<emphasis italics="true" pageId="0" pageNumber="649">A. marlae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
and
<taxonomicName lsidName="A. leniae" pageId="0" pageNumber="649" rank="species" species="leniae">
<emphasis italics="true" pageId="0" pageNumber="649">A. leniae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
);
<emphasis italics="true" pageId="0" pageNumber="649">it</emphasis>
between
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
closer to
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
than to
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
(equidistant in
<taxonomicName lsidName="A. leniae" pageId="0" pageNumber="649" rank="species" species="leniae">
<emphasis italics="true" pageId="0" pageNumber="649">A. leniae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
);
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
in the distal third between
<emphasis italics="true" pageId="0" pageNumber="649">dt</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
on the fixed finger (midway in between them in
<taxonomicName lsidName="A. lorenzorum" pageId="0" pageNumber="649" rank="species" species="lorenzorum">
<emphasis italics="true" pageId="0" pageNumber="649">A. lorenzorum</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
and
<taxonomicName lsidName="A. leniae" pageId="0" pageNumber="649" rank="species" species="leniae">
<emphasis italics="true" pageId="0" pageNumber="649">A. leniae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="etymology">
<paragraph pageId="0" pageNumber="649">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="649">This species is named after my former Latin teacher, Gerhard Faulstich, who might be one of the wisest men ML has ever known.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="description">
<paragraph pageId="0" pageNumber="649">Description.</paragraph>
<paragraph pageId="0" pageNumber="649">
The following description is based on holotype and allotype. -
<emphasis italics="true" pageId="0" pageNumber="649">Carapace</emphasis>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawings of Antsirananaella faulstichi sp. nov.. A carapace, dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure8" httpUri="https://binary.pensoft.net/fig/782000" pageId="0" pageNumber="649">8A</figureCitation>
): 1.36-1.55 (♂), 1.43-1.50 (♀) times longer than broad. -
<emphasis italics="true" pageId="0" pageNumber="649">Pedipalps</emphasis>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawings of Antsirananaella faulstichi sp. nov.. A carapace, dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure8" httpUri="https://binary.pensoft.net/fig/782000" pageId="0" pageNumber="649">8C</figureCitation>
): Trichobothrial pattern:
<emphasis italics="true" pageId="0" pageNumber="649">sb</emphasis>
on the movable finger distinctly distal to
<emphasis italics="true" pageId="0" pageNumber="649">est</emphasis>
on the fixed finger;
<emphasis italics="true" pageId="0" pageNumber="649">it</emphasis>
between
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
on the fixed finger closer to
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
than to
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="649">et</emphasis>
situated in the distal third between
<emphasis italics="true" pageId="0" pageNumber="649">dt</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="649">eb</emphasis>
on the fixed finger. Chelal fixed finger with 12 teeth in the OR, 16 in the MR and 15 in the IR; movable finger with 13 teeth in the OR, 13 in the MR and 16 in the IR. -
<taxonomicName genus="Dimensions" lsidName="Dimensions" pageId="0" pageNumber="649" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="649">Dimensions</emphasis>
</taxonomicName>
(mm):
<emphasis italics="true" pageId="0" pageNumber="649">Holotype</emphasis>
♂: Body length 1.98; abdomen length 1.35, width (without pleura) 1.16, 1.25 (with pleura); carapace length 0.62, width 0.42; Pedipalp: trochanter 0.25; femur length 0.57, width 0.35; patella 0.46; chela (without pedicel) 0.57; hand length 0.13, width 0.15; movable finger 0.44; Leg I: trochanter 0.11, femur 0.20, patella 0.16, tibia 0.18, tarsus 0.27; Leg IV: trochanter 0.18, femur 0.12, patella 0.33, tibia 0.29, tarsus 0.32.
<emphasis italics="true" pageId="0" pageNumber="649">Allotype</emphasis>
♀: Body length 2.28; abdomen length 1.53, width (without pleura) 1.34, 1.45 (with pleura); carapace length 0.73, width 0.51; Pedipalp: trochanter 0.32; femur length 0.71, width 0.38; patella 0.56; chela 0.65; hand length 0.14, width 0.18; movable finger 0.50; Leg I: trochanter 0.13; femur 0.25; patella 0.22; tibia 0.20; tarsus 0.33; Leg IV: trochanter 0.23; femur 0.14; patella 0.36; tibia 0.36; tarsus 0.36.
</paragraph>
<caption doi="10.3897/asp.80.e90570.figure8" httpUri="https://binary.pensoft.net/fig/782000" pageId="0" pageNumber="649" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="649">
<emphasis bold="true" pageId="0" pageNumber="649">Figure 8.</emphasis>
Drawings of
<taxonomicName authorityName="Lorenz &amp; Loria &amp; Harvey &amp; Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella faulstichi" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="faulstichi">
<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella faulstichi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
.
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
carapace, dorsal;
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
leg IV;
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
left chela from dorsal with trichobothrial pattern;
<emphasis bold="true" pageId="0" pageNumber="649">D</emphasis>
right pedipalp from dorsal. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
); 0.4 mm (
<emphasis bold="true" pageId="0" pageNumber="649">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="variation">
<paragraph pageId="0" pageNumber="649">Variation.</paragraph>
<paragraph pageId="0" pageNumber="649">Body length 1.95-2.02 (♂), 2.14-2.28 (♀); abdomen length 1.29-1.37 (♂), 1.46-1.53 (♀), width (without pleura) 1.12-1.23 (♂), 1.24-1.34 (♀), 1.20-1.30 (♂), 1.33-1.45 (♀) (with pleura); 1.09-1.22 (♂), 1.14-1.18 (♀) times longer than broad; carapace length 0.58-0.66 (♂), 0.69-0.73 (♀), width 0.42-0.45 (♂), 0.46-0.51 (♀), 1.36-1.51 (♂), 1.43-1.50 (♀) times longer than broad. Pedipalp: trochanter 0.24-0.31 (♂), 0.26-0.32 (♀); femur length 0.57-0.67 (♂), 0.65-0.71 (♀), width 0.31-0.36 (♂), 0.36-0.38 (♀); 1.62-2.00 (♂), 1.81-1.87 (♀) times longer than broad; tibia 0.46-0.51 (♂), 0.52-0.56 (♀); chela (without pedicel) 0.55-0.58 (♂), 0.62-0.65 (♀); movable finger 0.44-0.47 (♂), 0.49-0.50 (♀); hand length (without pedicel) 0.10-0.13 (♂), 0.10-0.14 (♀), width 0.12-0.16 (♂), 0.16-0.18 (♀); 0.63-1.00 (♂), 0.63-0.78 (♀) times longer than broad.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="genbank code">
<paragraph pageId="0" pageNumber="649">GenBank Code.</paragraph>
<paragraph pageId="0" pageNumber="649">Genetic data is unavailable for this species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="habitat">
<paragraph pageId="0" pageNumber="649">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="649">Tropical dry forest at 290 m altitude.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="649" type="distribution">
<paragraph pageId="0" pageNumber="649">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="649">
Known from the type locality (BLF3012) and four additional locations (BLF11400, BLF2858, BLF2972 and BLF9656) in the Diana and Sava Regions (formerly
<taxonomicName genus="Antsiranana" lsidName="Antsiranana" pageId="0" pageNumber="649" rank="genus">Antsiranana</taxonomicName>
Province).
</paragraph>
</subSubSection>
</treatment>
</document>