130 lines
19 KiB
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130 lines
19 KiB
XML
<document id="A1EA235D6A8536387CD20B3E269B5B99" ID-DOI="http://dx.doi.org/10.3897/zookeys.482.8343" ID-GBIF-Dataset="4fdb2d6d-b69c-4e87-9fb3-1de661126876" ID-PMC="PMC4337222" ID-Pensoft-Pub="1313-2970-482-91" ID-PubMed="25709532" ID-ZooBank="21C3B7D0B18743EABB38175C704D7550" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-482-91" ModsDocOrigin="ZooKeys 482" ModsDocTitle="Review of the genera Anelaphinis Kolbe, 1892 and Atrichelaphinis Kraatz, 1898 (Coleoptera, Scarabaeidae, Cetoniinae)" checkinTime="1451244653051" checkinUser="pensoft" docAuthor="Rojkoff, Sebastien & Perissinotto, Renzo" docDate="2015" docId="20C1B89FBD7ECBBAE8B2EA7D429CC836" docLanguage="en" docName="ZooKeys 482: 91-142" docOrigin="ZooKeys 482" docSource="http://dx.doi.org/10.3897/zookeys.482.8343" docTitle="Atrichelaphinis (Heterelaphinis) quadripunctata Lansberge 1882" docType="treatment" docVersion="5" lastPageNumber="102" masterDocId="822C54210D6A7F60FFE2FFC9FFA1A039" masterDocTitle="Review of the genera Anelaphinis Kolbe, 1892 and Atrichelaphinis Kraatz, 1898 (Coleoptera, Scarabaeidae, Cetoniinae)" masterLastPageNumber="142" masterPageNumber="91" pageNumber="102" updateTime="1701316131199" updateUser="plazi">
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<mods:title id="6ACFB4AC01D73920E100842CFF8FFF68">Review of the genera Anelaphinis Kolbe, 1892 and Atrichelaphinis Kraatz, 1898 (Coleoptera, Scarabaeidae, Cetoniinae)</mods:title>
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<mods:namePart id="848EB448CCB4E6089D97F34A88D9ED6A">Rojkoff, Sebastien</mods:namePart>
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<mods:namePart id="AD180E5F3D932F838B56A6808742ED44">Perissinotto, Renzo</mods:namePart>
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<mods:typeOfResource id="3A0177609A10F79D87E6117C84B1AB99">text</mods:typeOfResource>
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<mods:date id="351ADA97501434A01A5DE03056DB6219">2015</mods:date>
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<mods:number id="883494813E84636AA5F8D6ED1EBBF176">482</mods:number>
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<mods:start id="E36215ADCCCD4F50680886BDF200B6D5">91</mods:start>
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<mods:end id="7ACBE681E140E1224B6620F86A99B2E1">142</mods:end>
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<mods:classification id="EDC34B7C67994553FC925D8735BDBF93">journal article</mods:classification>
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<mods:identifier id="E186E5110CB07409B780DA1FF6979BE4" type="DOI">http://dx.doi.org/10.3897/zookeys.482.8343</mods:identifier>
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<mods:identifier id="3D26CF9DB733486C694C15844292425B" type="Pensoft-Pub">1313-2970-482-91</mods:identifier>
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<treatment id="20C1B89FBD7ECBBAE8B2EA7D429CC836" ID-GBIF-Taxon="152058286" LSID="urn:lsid:plazi:treatment:20C1B89FBD7ECBBAE8B2EA7D429CC836" httpUri="http://treatment.plazi.org/id/20C1B89FBD7ECBBAE8B2EA7D429CC836" lastPageNumber="102" pageId="11" pageNumber="102">
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<subSubSection id="7D0AB5DE60BE51B7F73E5FFE29E42A30" pageId="11" pageNumber="102" type="multiple">
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<paragraph id="E3053C2F2E44F67111A53B362C90D7F1" pageId="11" pageNumber="102">
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<pageBreakToken id="540624C0D6F56F8C3A6CFFDBAE59019A" pageId="11" pageNumber="102" start="start">Taxon</pageBreakToken>
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classification Animalia Coleoptera Scarabaeidae
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</paragraph>
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</subSubSection>
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<subSubSection id="47BB307DF8744921FDBDB98000AEC51D" pageId="11" pageNumber="102" type="nomenclature">
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<paragraph id="C6C8E3A626F4F01508B5437DBAE1DF79" pageId="11" pageNumber="102">
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<taxonomicName id="13053B20E0F6C01845C0976475F29883" ID-CoL="JKG4" authority="Lansberge, 1882" authorityName="Lansberge" authorityYear="1882" class="Insecta" family="Scarabaeidae" genus="Atrichelaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichelaphinis (Heterelaphinis) quadripunctata" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="quadripunctata" subGenus="Heterelaphinis">Atrichelaphinis (Heterelaphinis) quadripunctata (Lansberge, 1882)</taxonomicName>
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Figures 5and 6
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</paragraph>
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</subSubSection>
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<subSubSection id="E4C93288DA34098C056F7402B4FBD830" pageId="11" pageNumber="102" type="reference_group">
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<paragraph id="E585832EFE7A9C7D49186343B80BA87F" pageId="11" pageNumber="102">
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<taxonomicName id="3AAD12615804729EB171557D0EEB3651" class="Insecta" family="Scarabaeidae" genus="Elaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elaphinis quadripunctata" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="quadripunctata">Elaphinis quadripunctata</taxonomicName>
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Lansberge, 1882: 24;
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<bibRefCitation id="B779183384D519DAA613B69FB7C46E32" author="Ritsema, C" journalOrPublisher="Tijdschrift voor Entomologie" pageId="35" pageNumber="126" pagination="201 - 234" title="Lijst der entomologische Geschriften van Mr. J. W. van Lansberge, gevolgd door eenen opgawe der daarin beschreven nieuwe geslachten, ondergeslachten en soorten." volume="31" year="1888">Ritsema 1888</bibRefCitation>
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: 225;
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<bibRefCitation id="930E4E74C71B598A71CB2EFF71EBE727" author="Kolbe, HJ" editor="Moebius, K" journalOrPublisher="Dietrich Reimer, Berlin" pageId="34" pageNumber="125" title="Kaefer und Netzflueger Ost-Afrikas." volumeTitle="Deutsch Ostafrika - Wissenschaftliche Forschungsresultate ueber Land und Leute unseres ostafrikanischen Schutzgebietes und der Angrenzenden Laender (Band IV) - Die Thierwelt Ost-Afrikas und der Nachbargebiete - Wirbellose Thiere, Coleopteren" year="1897">Kolbe 1897</bibRefCitation>
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: 180;
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<bibRefCitation id="EDF7D6FC2FE4AF5CF7D2267EF023728E" author="Antoine, P" journalOrPublisher="Revue francaise d'Entomologie (NS)" pageId="32" pageNumber="123" pagination="1 - 12" title="Revision des genres Dolichosthetus Kolbe et Dulcinea Peringuey (Coleoptera, Cetoniidae)." volume="13" year="1991">Antoine 1991</bibRefCitation>
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: 2;
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<bibRefCitation id="9C7F42326618584BB13154DCCDD9B319" author="Marais, E" journalOrPublisher="Cimbebasia Memoir" pageId="34" pageNumber="125" pagination="1 - 117" title="Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa." volume="8" year="1992">Marais and Holm 1992</bibRefCitation>
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: 11.
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</paragraph>
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<paragraph id="27315D2B1494FE86D6171F427D080AA5" pageId="11" pageNumber="102">
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<taxonomicName id="28B3227D2B36ED0460715D6B7DA15CBB" class="Insecta" family="Scarabaeidae" genus="Atrichelaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichelaphinis quadripunctata" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="quadripunctata">Atrichelaphinis quadripunctata</taxonomicName>
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(Lansberge) Marais & Holm, 1989: 30;
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<bibRefCitation id="E6902A1E4EB084F79CF1D72D98C7BF38" author="Marais, E" journalOrPublisher="Cimbebasia Memoir" pageId="34" pageNumber="125" pagination="1 - 117" title="Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa." volume="8" year="1992">Marais and Holm 1992</bibRefCitation>
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: 11;
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<bibRefCitation id="CFD3C1044710C7B6CC2C06FCEE06EBFD" author="Krajcik, M" journalOrPublisher="Typos Studio Most" pageId="34" pageNumber="125" title="Cetoniidae of the World - Catalogue, Part I" year="1998">Krajcik 1998</bibRefCitation>
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: 50.
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</paragraph>
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<paragraph id="C67CBB858FA54642128F4E50208FAFED" pageId="11" pageNumber="102">
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<taxonomicName id="B9EE8B7637D8D5F3ABC3307CC6E9636E" class="Insecta" family="Scarabaeidae" genus="Cetonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cetonia quadripunctata" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="quadripunctata">Cetonia quadripunctata</taxonomicName>
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(Lansberge) Antoine, 2002: 185.
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</paragraph>
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</subSubSection>
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<subSubSection id="9AFD16500016AC96E87161975DC12FFB" pageId="11" pageNumber="102" type="type specimen">
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<paragraph id="A6DA4AF0B4E0762EF4F0A5A94724CA25" pageId="11" pageNumber="102">Type specimen.</paragraph>
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<paragraph id="E0FA0273D10E06FE2BBEF8C9475E8250" pageId="11" pageNumber="102">
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<bibRefCitation id="FA19C217569FE87AB7010674F944CE6B" author="Marais, E" journalOrPublisher="Cimbebasia Memoir" pageId="34" pageNumber="125" pagination="1 - 117" title="Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa." volume="8" year="1992">Marais and Holm (1992)</bibRefCitation>
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mentioned two paralectotypes: one in the BMNH collections and one in the MNHN. The male specimen housed in the MNHN shows the following labels: "Somali, Ouarsangueli, Revoil 1881, Museum Paris/1598 81"; and "Lectotype,
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<taxonomicName id="17041C00DBFBF19985874357D31ABC2A" class="Insecta" family="Scarabaeidae" genus="Elaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elaphinis quadripunctata" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="quadripunctata">Elaphinis quadripunctata</taxonomicName>
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van Lansberge, Ph. Antoine det 88". There is, however no reference to this designation in the publications of
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<bibRefCitation id="D82345E3CD400CB4486BE488A8516451" author="Antoine, P" journalOrPublisher="Revue francaise d'Entomologie (NS)" pageId="32" pageNumber="123" pagination="1 - 12" title="Revision des genres Dolichosthetus Kolbe et Dulcinea Peringuey (Coleoptera, Cetoniidae)." volume="13" year="1991">Antoine (1991</bibRefCitation>
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,
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<bibRefCitation id="D1A7939CCF33D267C39B183F0DFBA2ED" author="Antoine, P" journalOrPublisher="Coleopteres" pageId="32" pageNumber="123" pagination="175 - 190" title="Quelques especes nouvelles ou peu connues de la famille des Cetoniidae - X (ColeopteraScarabaeoidea)." volume="8" year="2002">2002</bibRefCitation>
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), apart from a mention of the lectotype in the legend to Figure 21 of
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<bibRefCitation id="77CEE2E5F588008D9B51D265BF7CF636" author="Antoine, P" journalOrPublisher="Coleopteres" pageId="32" pageNumber="123" pagination="175 - 190" title="Quelques especes nouvelles ou peu connues de la famille des Cetoniidae - X (ColeopteraScarabaeoidea)." volume="8" year="2002">Antoine (2002)</bibRefCitation>
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. Consequently, in order to settle the status of the species, the male illustrated in Figure 5 is here designated as Lectotype and a new label is added to the two described above, reading: "Lectotype,
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<taxonomicName id="2BE9F867B9DAE11EB80708963445F82B" class="Insecta" family="Scarabaeidae" genus="Elaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elaphinis quadripunctata" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="quadripunctata">Elaphinis quadripunctata</taxonomicName>
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van Lansberge, Rojkoff & Perissinotto 2014". Four other specimens, identified as
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<taxonomicName id="2E50C28BBF4689C03760E1C65B768C6B" class="Insecta" family="Scarabaeidae" genus="Elaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elaphinis quadripunctata" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="quadripunctata">Elaphinis quadripunctata</taxonomicName>
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by Antoine in 1994, were found in the MNHN collections. Two females have the same label as the lectotype and are here designated as paralectotypes. The last specimens, one male missing pronotum and head and a female are only labelled "Ex-Musaeo Van Lansberge" and "Museum Paris, ex. Coll. R. Oberthur". It is possible that these specimens belong to the type series, but as this could not be confirmed during this study, they cannot be designated as paralectotypes here.
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</paragraph>
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</subSubSection>
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<subSubSection id="06189FFC562CD4B1F52B2B0175B214E8" pageId="11" pageNumber="102" type="redescription">
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<paragraph id="BB85CE04FB67E679ABB79CD1682ED623" pageId="11" pageNumber="102">Redescription</paragraph>
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<paragraph id="9504A2C792B2795D0A4D11CEEE69CCD0" pageId="11" pageNumber="102">(n = 7). Size: length LT ♂, 11 mm; width 5.5 mm.</paragraph>
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<paragraph id="5CDE2A456E042011F4EF63C43DA17A12" pageId="11" pageNumber="102">Head. Dark brown with blackish areas, strongly sculpted, converging points forming deep striae; clypeus longer than wide, lateral and anterior margins strongly reborded, anterior slightly upturned and very slightly bilobed, lateral margins almost carinate in the basal part, then curved downwards, depressed in the middle and reborded in the apical part as the anterior margin, clypeal disc convex; frons with large striated protuberance between eyes, vertex with few smooth jointed areas between striae in apical part, posterior part only punctate; antennae brown with long clubs (as long as the flagellum in male).</paragraph>
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<paragraph id="60D6BFCDF4769706CB4678BD68DC3DB0" pageId="11" pageNumber="102">Pronotum. Transverse, dark brown with transverse points of sculpture, disc poorly punctate, sculpture becoming more dense and confluent to striae in front and laterally; anterior margin slightly wider than head, medially slightly tuberculate; lateral margins reborded with very smooth lateral angles in posterior third; posterior margin bisinuate (concave in front of scutellum), posterior angles rounded.</paragraph>
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<paragraph id="881470AD04E5AF590E51DCFB54E362E9" pageId="11" pageNumber="102">Scutellum. Dark brown, longer than wide, apex acute, smooth, only a few setigerous points on lateral angles (scale pilosity); laterally grooved.</paragraph>
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<paragraph id="05CDACBA8C8F10D3053E1145EAB652E1" pageId="11" pageNumber="102">Elytra. Orange with four black markings, one on side of scutellum, one at middle split on each costa (discal and humeral), one on apical quarter near the suture and last on spiny apex; costae convex, smooth with only few points, discal costa incomplete, humeral costa concave to suture with concavity reaching elytral disc; sculpture of small longitudinal lines (near scutellum) and of horseshoe points in anterior half, becoming confluent posteriorly and leading to two formations: 1) laterally (i.e. between humeral costa and lateral margin) transverse lines becoming longer and denser toward apex; 2) longitunal lines between sutural and humeral costae becoming more numerous and strigillate towards apex; few minute and very short setae near apex.</paragraph>
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<paragraph id="3D9D61A4369DC00096824A8E0D7679A2" pageId="11" pageNumber="102">Pygidium. Transverse, chestnut brown; sculpture horseshoe-like to annulate points drawing large irregular circles towards apex, some transverse striae along apex; few minute and very short setae.</paragraph>
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<paragraph id="AE8DDE2859A79558CA1B9FE518BAAAAF" pageId="11" pageNumber="102">Underside. From dark brown to chestnut brown, sculpture setigerous with long whitish pilosity, not dense except on femora and laterally on sternites 2-5; sculpture sparce, crescent on metasternum, denser to confluent laterally, disc poorly sculpted; abdomen with horseshoe sculpture, median part almost smooth, denser laterally near the margin; posterior coxae reborded laterally, setigerous sculpture made of transversal to backward-curved striae; mesosternal apophysis glabrous, transverse with minute points, strongly compressed between mesocoxae but not protruding; male abdomen concave with visible groove on sternites 2-5.</paragraph>
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<paragraph id="F9DA1A0B8828127FAA0AE3C8EC67122C" pageId="11" pageNumber="102">Legs. From dark brown to chestnut brown, with whitish pilosity; protibiae bidentate, meso- and metatibiae with median carina; profemora strigillate, mesofemora with crescent punctures or small striae, long setigerous stria along internal margin; metafemora slightly dilated with crescent punctures or small striae; all tarsal segments longer than first, metatarsi spiny, claws normal.</paragraph>
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<paragraph id="5A653FF368E842937606A5BDCAECFF98" pageId="11" pageNumber="102">Aedeagus. Parameres fused and short, with two carinated lateral spines at apex, apical centre with short protrusion.</paragraph>
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</subSubSection>
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<subSubSection id="21DC960FBC5992965F5B19478EFE1721" pageId="11" pageNumber="102" type="remarks">
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<paragraph id="10982294B3E8CEE441FEDF2CFDBEB682" pageId="11" pageNumber="102">Remarks.</paragraph>
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<paragraph id="9D2EA90C0041B2C40FDDEE0F3A609A82" pageId="11" pageNumber="102">
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Only the MNHN type specimens are known. No recent material was found in the collections examined. Unfortunately,
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<bibRefCitation id="80AFDD7C3B8DB708E35A1763A03711F7" author="Lansberge, JW" journalOrPublisher="Bulletin de la Societe entomologique de Belgique" pageId="34" pageNumber="125" pagination="21 - 31" title="Enumeration des Scarabaeides rapportes du pays des Somalis (Afrique equatoriale) par M. Revoil avec diagnoses des especes nouvelles." volume="26" year="1882">Lansberge (1882)</bibRefCitation>
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did not specify the number of specimens used for his description. The specimen length indicated in his work does not match the measurements reported above. This difference cannot be explained at this stage, but it is possible that
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<bibRefCitation id="1CA3EA6094990CC4EDFBE5A888404B6F" author="Lansberge, JW" journalOrPublisher="Bulletin de la Societe entomologique de Belgique" pageId="34" pageNumber="125" pagination="21 - 31" title="Enumeration des Scarabaeides rapportes du pays des Somalis (Afrique equatoriale) par M. Revoil avec diagnoses des especes nouvelles." volume="26" year="1882">Lansberge (1882)</bibRefCitation>
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may have only provided a coarse estimate, without accurate measurement. The female is larger than the male; its abdomen does not exhibit a deep groove but there are occasional sligth depressions on sternites III and IV, otherwise it varies from flat to slightly convex. The main difference between the two sexes lies in the metatibial spurs, which are strongly enlarged in the female (especially the upper one, spatuliform when thin), but acute and curved at the apex in the male. Nothing is known about the biology of this species, but the adult is probably a flower visitor.
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</paragraph>
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<caption id="8FB8A670DFFDF115B855D106BF1E8A5D" pageId="11" pageNumber="102">
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<paragraph id="8E1D1748742DC70268250737B08650E4" pageId="11" pageNumber="102">
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Figure 5.
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<taxonomicName id="52991DDC7585A46D019C43C48D25CB33" class="Insecta" family="Scarabaeidae" genus="Atrichelaphinis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichelaphinis (Heterelaphinis) quadripunctata" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="quadripunctata" subGenus="Heterelaphinis">Atrichelaphinis (Heterelaphinis) quadripunctata</taxonomicName>
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(Lansberge, 1882), lectotype male, Somalia (MNHN). A Dorsal view B ventral view C parameres in dorsal view D parameres in lateral view E apex of parameres.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |