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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.895.38576" ID-GBIF-Dataset="f9f1c191-c386-4c8c-9abd-4b4e6fecd2a6" ID-PMC="PMC6906171" ID-Pensoft-Pub="1313-2970-895-1" ID-Pensoft-UUID="99AB66C9ACF058A581D9036697D415F3" ID-PubMed="31844411" ID-ZooBank="11350FE921E64D7AB1F916CE4639F406" ModsDocID="1313-2970-895-1" checkinTime="1575565024213" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas &amp; Gattolliat, Jean-Luc" docDate="2019" docId="1A70DFCF68FC59AB8AF68B6C28A4E5CF" docLanguage="en" docName="ZooKeys 895: 1-117" docOrigin="ZooKeys 895" docSource="http://dx.doi.org/10.3897/zookeys.895.38576" docTitle="Labiobaetis paraoperosus Kaltenbach &amp; Gattolliat, 2019, sp. nov." docType="treatment" docUuid="7CC2BD0A-1543-4570-A81D-CE4189B8A945" docUuidSource="ZooBank" docVersion="5" id="99AB66C9ACF058A581D9036697D415F3" lastPageNumber="1" masterDocId="99AB66C9ACF058A581D9036697D415F3" masterDocTitle="The tremendous diversity of Labiobaetis Novikova &amp; Kluge in Indonesia (Ephemeroptera, Baetidae)" masterLastPageNumber="117" masterPageNumber="1" pageNumber="1" updateTime="1668168188677" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The tremendous diversity of Labiobaetis Novikova &amp; Kluge in Indonesia (Ephemeroptera, Baetidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kaltenbach, Thomas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>895</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>117</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.895.38576</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.895.38576</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-895-1</mods:identifier>
<mods:identifier type="ZooBank">11350FE921E64D7AB1F916CE4639F406</mods:identifier>
<mods:identifier type="Pensoft-UUID">99AB66C9ACF058A581D9036697D415F3</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="160631157" LSID="urn:lsid:zoobank.org:act:7CC2BD0A-1543-4570-A81D-CE4189B8A945" httpUri="http://treatment.plazi.org/id/1A70DFCF68FC59AB8AF68B6C28A4E5CF" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
21.
<taxonomicName LSID="1A70DFCF-68FC-59AB-8AF6-8B6C28A4E5CF" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis paraoperosus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="paraoperosus">Labiobaetis paraoperosus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis paraoperosus sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure36">Figures 36</figureCitation>
,
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis paraoperosus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape g Metanotum." figureDoi="10.3897/zookeys.895.38576.figure37" httpUri="https://binary.pensoft.net/fig/361927" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure37">37</figureCitation>
,
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Habitus, larvae, dorsal view: a Labiobaetis multus b Labiobaetis jonasi sp. nov. c Labiobaetis jonasi sp. nov. d Labiobaetis paraoperosus sp. nov." figureDoi="10.3897/zookeys.895.38576.figure51" httpUri="https://binary.pensoft.net/fig/361942" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure51">51d</figureCitation>
,
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Distribution of Labiobaetis in Indonesia." figureDoi="10.3897/zookeys.895.38576.figure55" httpUri="https://binary.pensoft.net/fig/361946" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure55">55b</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
Following combination of characters: A) dorsal surface of labrum with submarginal arc of 1 + 9 or 10 long, feathered setae; B) labial palp segment II with a large, thumb-like distomedial protuberance, segment III oblong; C) fore femur rather broad, length ca. 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width, dorsal margin with a row of 11-18 spine-like setae; D) hindwing pad well developed; E) paraproct distally not expanded, with ca. 30 stout marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva</emphasis>
(
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis paraoperosus sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure36">Figs 36</figureCitation>
,
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis paraoperosus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape g Metanotum." figureDoi="10.3897/zookeys.895.38576.figure37" httpUri="https://binary.pensoft.net/fig/361927" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure37">37</figureCitation>
,
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Habitus, larvae, dorsal view: a Labiobaetis multus b Labiobaetis jonasi sp. nov. c Labiobaetis jonasi sp. nov. d Labiobaetis paraoperosus sp. nov." figureDoi="10.3897/zookeys.895.38576.figure51" httpUri="https://binary.pensoft.net/fig/361942" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure51">51d</figureCitation>
). Body length 5.5-6.0 mm; antenna approximately 3
<normalizedToken originalValue="×">x</normalizedToken>
as long as head length; cerci ca. 0.7
<normalizedToken originalValue="×">x</normalizedToken>
body length.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
. Head, thorax, and abdomen dorsally brown, head and thorax with bright median, dorsal suture, thorax and abdomen with pattern as in
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Habitus, larvae, dorsal view: a Labiobaetis multus b Labiobaetis jonasi sp. nov. c Labiobaetis jonasi sp. nov. d Labiobaetis paraoperosus sp. nov." figureDoi="10.3897/zookeys.895.38576.figure51" httpUri="https://binary.pensoft.net/fig/361942" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure51">Fig. 51d</figureCitation>
. Forewing pads light brown at base and with light brown distal spots. Head, thorax, and abdomen ventrally very light brown, abdominal segments VIII and IX brown. Legs transparent, femur with a distomedial, brown spot and brown apex, caudal filaments transparent with a dark brown band at ca. 1/3 of cerci.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
with scape and pedicel subcylindrical, with well-developed distolateral process at scape; flagellum with lanceolate spines and fine, simple setae on apex of each segment.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
(
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis paraoperosus sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure36">Fig. 36a</figureCitation>
). Rectangular, length 0.7
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae; submarginal arc of setae composed of 1 + 9 or10 long, feathered setae. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid, pectinate setae; ventral surface with three short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
(
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis paraoperosus sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure36">Fig. 36b, c</figureCitation>
). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles and one minute intermediate denticle. Inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola straight. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
(
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis paraoperosus sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure36">Fig. 36d, e</figureCitation>
). Incisors fused. Outer and inner sets of denticles with 4 + 3 denticles. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight, with minute denticles towards subtriangular process. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="1">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hypopharynx</emphasis>
</taxonomicName>
(
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis paraoperosus sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure36">Fig. 36f</figureCitation>
). Lingua approx. as long as superlingua. Lingua broader than long; medial tuft of stout setae poorly developed; distal half not expanded. Superlingua rounded; lateral margin rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
(
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis paraoperosus sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure36">Fig. 36g</figureCitation>
). Galea-lacinia with two simple, robust apical setae under crown. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and 4-5 long, simple setae. Maxillary palp 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of galea-lacinia; two segmented; palp segment II 1.3
<normalizedToken originalValue="×">x</normalizedToken>
length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment rounded, with slight excavation at inner distolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Labium</emphasis>
(
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Labiobaetis paraoperosus sp. nov., larva morphology: a Labrum b Right mandible c Right prostheca d Left mandible e Left prostheca f Hypopharynx g Maxilla h Labium i Apex of paraglossa." figureDoi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure36">Fig. 36h, i</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with eight or nine spine-like setae increasing in length distally; apex with two long and one medium, robust, pectinate setae and one short, robust seta; outer margin with 5-7 spine-like setae increasing in length distally; ventral surface with short, fine, simple, scattered setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and two or three medium, simple setae in anteromedial area and partly one short, simple seta in proximomedial area; dorsally with a row of four or five long, spine-like setae near inner margin. Labial palp with segment I 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with broad thumb-like distomedial protuberance; distomedial protuberance 0.9
<normalizedToken originalValue="×">x</normalizedToken>
width of base at segment III; inner and outer margin with short, fine, simple setae; dorsally with two long, spine-like, simple setae near outer margin. Segment III oblong; apex rounded; length 1.5
<normalizedToken originalValue="×">x</normalizedToken>
width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<caption doi="10.3897/zookeys.895.38576.figure36" httpUri="https://binary.pensoft.net/fig/361926" pageId="0" pageNumber="1" start="Figure 36" startId="F36">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
<taxonomicName class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis paraoperosus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="paraoperosus">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis paraoperosus</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
Labrum
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
Right mandible
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
Right prostheca
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
Left mandible
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
Left prostheca
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
<taxonomicName genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="1" rank="genus">Hypopharynx</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
Maxilla
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
Labium
<emphasis bold="true" pageId="0" pageNumber="1">i</emphasis>
Apex of paraglossa.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Hind wing pads</emphasis>
well developed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
(
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis paraoperosus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape g Metanotum." figureDoi="10.3897/zookeys.895.38576.figure37" httpUri="https://binary.pensoft.net/fig/361927" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure37">Fig. 37a, b</figureCitation>
). Ratio of foreleg segments 1.2:1.0:0.5:0.2.
<emphasis italics="true" pageId="0" pageNumber="1">Femur</emphasis>
. Length ca. 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with a row of 11-18 spine-like setae and one or two spine-like setae near margin; length of setae 0.25
<normalizedToken originalValue="×">x</normalizedToken>
maximum width of femur. Apex rounded; with one pair of curved, spine-like setae and some short, stout setae. Many short, stout, lanceolate setae scattered along the ventral margin; femoral patch absent.
<emphasis italics="true" pageId="0" pageNumber="1">Tibia</emphasis>
. Dorsal margin with a row of short, stout setae; on apex one longer seta, and a row of short, stout setae close to dorsal margin. Ventral margin with a row of curved, spine-like setae, distally longer, on apex one bipectinate, spine-like seta and a tuft of long, fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal 1/2.
<emphasis italics="true" pageId="0" pageNumber="1">Tarsus</emphasis>
. Dorsal margin with a row of short, stout setae and long, fine, simple setae. Ventral margin with a row of curved, spine-like setae. Tarsal claw with one row of 9-11 denticles; distally pointed; with 3-5 stripes; subapical setae absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Tergum</emphasis>
(
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis paraoperosus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape g Metanotum." figureDoi="10.3897/zookeys.895.38576.figure37" httpUri="https://binary.pensoft.net/fig/361927" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure37">Fig. 37c</figureCitation>
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae. Posterior margin of tergum IV with triangular spines, wider than long.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gills</emphasis>
(
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis paraoperosus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape g Metanotum." figureDoi="10.3897/zookeys.895.38576.figure37" httpUri="https://binary.pensoft.net/fig/361927" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure37">Fig. 37d</figureCitation>
). Present on segments
<normalizedToken originalValue="IVII">I-VII</normalizedToken>
. Margin with small denticles intercalating fine simple setae. Tracheae extending from main trunk to inner and outer margins. Gill I ca. 2/3 length of segment II. Gill IV as long as length of segments V and 1/3 VI combined. Gill VII as long as length of segment VIII.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
(
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Labiobaetis paraoperosus sp. nov., larva morphology: a Foreleg b Fore claw c Tergum IV d Gill IV e Paraproct f Scape g Metanotum." figureDoi="10.3897/zookeys.895.38576.figure37" httpUri="https://binary.pensoft.net/fig/361927" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.895.38576.figure37">Fig. 37e</figureCitation>
). Distally not expanded with 29-32 stout marginal spines. Surface scattered with U-shaped scale bases and fine, simple setae. Cercotractor with numerous small marginal spines.
</paragraph>
<caption doi="10.3897/zookeys.895.38576.figure37" httpUri="https://binary.pensoft.net/fig/361927" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
<taxonomicName class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis paraoperosus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="paraoperosus">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis paraoperosus</emphasis>
</taxonomicName>
sp. nov., larva morphology:
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
Foreleg
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
Fore claw
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
Tergum IV
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
Gill IV
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
Paraproct
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
Scape
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
Metanotum.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Refers to the morphological similarity with
<taxonomicName lsidName="L. operosus" pageId="0" pageNumber="1" rank="species" species="operosus">
<emphasis italics="true" pageId="0" pageNumber="1">L. operosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Indonesia: Sumatra.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biological aspects">
<paragraph pageId="0" pageNumber="1">Biological aspects.</paragraph>
<paragraph pageId="0" pageNumber="1">The specimens were collected at altitudes between sea level and 890 m, one of them in a small (2 m wide), shallow (25 cm deep) and slow (velocity 0.3 m/s) stream in a highly disturbed area (agriculture, livestock, waste). The substrate was predominantly stone and gravel without patches of leaf litter or dead wood.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type-material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Holotype.</emphasis>
Larva (on slide, GBIFCH 00592254), Indonesia, Sumatra, Aceh, Ketambe, Alas River at Cinta Alam Lodge,
<geoCoordinate degrees="03" direction="north" minutes="40.73" orientation="latitude" precision="9" value="3.6788332">03°40.73'N</geoCoordinate>
,
<geoCoordinate degrees="97" direction="east" minutes="39.37" orientation="longitude" precision="9" value="97.656166">97°39.37'E</geoCoordinate>
, 400 m, 14.X.2013, M. Balke leg. (SUM 45). Temporary deposited in MZL before definitely housed in MZB.
<emphasis bold="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
11 larvae (1 on slide, GBIFCH 00592253, 5 in alcohol, GBIFCH 00515329, GBIFCH 00515338, deposited in MZL; 5 in alcohol, GBIFCH 00515328, deposited in ZSM), same data as holotype; 1 larva (on slide, GBIFCH 00592249, deposited in MZL), Indonesia, Sumatra Barat, Sawahlunto, stream, 275 m, 10.XI.2011,
<geoCoordinate degrees="00" direction="south" minutes="41.33" orientation="latitude" precision="9" value="-0.68883336">00°41.33'S</geoCoordinate>
,
<geoCoordinate degrees="100" direction="east" minutes="46.72" orientation="longitude" precision="9" value="100.77866">100°46.72'E</geoCoordinate>
, M. Balke leg. (UN5); 1 larva (on slide, GBIFCH 00422165, deposited in MZL), Indonesia, Sumatra, volcano Marapi,
<geoCoordinate degrees="00" direction="south" minutes="21.97" orientation="latitude" precision="9" value="-0.36616665">00°21.97'S</geoCoordinate>
,
<geoCoordinate degrees="100" direction="east" minutes="33.30" orientation="longitude" precision="9" value="100.555">100°33.30'E</geoCoordinate>
, 890 m, 02.IV.2014, M. Gueuning leg.; 2 larvae (on slides, GBIFCH 00592248, GBIFCH 00592247, deposited in MZL), Indonesia, Sumatra Barat, Tarusan, upstream Tarusan,
<geoCoordinate degrees="01" direction="south" minutes="13.62" orientation="latitude" precision="9" value="-1.227">01°13.62'S</geoCoordinate>
,
<geoCoordinate degrees="100" direction="east" minutes="29.83" orientation="longitude" precision="9" value="100.49717">100°29.83E</geoCoordinate>
, 10 m, 24.V.2010, J.-M. Elouard leg.
</paragraph>
</subSubSection>
</treatment>
</document>