185 lines
19 KiB
XML
185 lines
19 KiB
XML
<document id="B999C82910F0DE844E3719F54B0D6283" ID-CLB-Dataset="30461" ID-DOI="10.11646/zootaxa.4399.1.13" ID-GBIF-Dataset="b7b33ffc-5409-49b5-a7ec-30b8db38c1cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1206434" ID-ZooBank="02E46C96-076D-4404-87CD-192B891E328A" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521898265120" checkinUser="plazi" docAuthor="Li, Qinghua, Ren, Dong & Wang, Yongjie" docDate="2018" docId="FE5082769747FFE5FF3AFD745937D90B" docLanguage="en" docName="zootaxa.4399.1.13.pdf" docOrigin="Zootaxa 4399 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nilionympha shantouensis Li & Ren & Wang 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="149" masterDocId="0269FA0E9745FFE6FFADFFC35812DB64" masterDocTitle="Revision of the gumilline genus Nilionympha with a new species from the Middle Jurassic of China (Neuroptera: Osmylidae)" masterLastPageNumber="150" masterPageNumber="146" pageNumber="148" updateTime="1698532585944" updateUser="ExternalLinkService">
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<mods:mods id="FEC9AAFD16F842C7E6E378096ACD6C7B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="07508645FA912FEF447400BB5647C996">
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<mods:title id="AAD3C3AECBE0B868EBB3C46CCA5A44CA">Revision of the gumilline genus Nilionympha with a new species from the Middle Jurassic of China (Neuroptera: Osmylidae)</mods:title>
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<mods:name id="C6B1848F4CCC53B7449EE5ADA15FD888" type="personal">
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<mods:role id="171BAFC1D61248C0E67948366D304090">
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<mods:roleTerm id="CFBBEAE142D9B1163E4F102DE4CB227B">Author</mods:roleTerm>
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<mods:namePart id="CB870E60FB70B8A6F83ED53ED0113135">Li, Qinghua</mods:namePart>
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<mods:roleTerm id="822853C4BE54580DE6951056DF219BA8">Author</mods:roleTerm>
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<mods:namePart id="DAC6815526F098FC350C4C4747F49890">Ren, Dong</mods:namePart>
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<mods:name id="8E9F77311491072D11271321B74C49AD" type="personal">
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<mods:roleTerm id="26C1EC30072EEF0A081ED0D480513E6C">Author</mods:roleTerm>
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<mods:namePart id="99DE6A452E83483D91F491CC670D0855">Wang, Yongjie</mods:namePart>
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<mods:typeOfResource id="933125458A85FC2AD28A2279BF0E01FF">text</mods:typeOfResource>
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<mods:title id="438D172F9E261F0B7301D8B0B27AD3A6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="158739552D6E4D864B5DC192B4208CAB">
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<mods:date id="ABA124A2EE0C123ED504B46D98CF9E03">2018</mods:date>
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<mods:detail id="FE328B27FC5F87D200B0B9F4083CD66E" type="pubDate">
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<mods:number id="A1EA66FCCEA58640FB02D737F6036EE3">2018-03-20</mods:number>
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<mods:detail id="9C14F37BF07C92BCA23A5AA4C6E7181F" type="volume">
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<mods:number id="63CA96F34C68B40F7B9D88C8FC216BB5">4399</mods:number>
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<mods:detail id="F575A6E13FE4877FBA9DD306AEF1A9B6" type="issue">
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<mods:start id="921A3EE462D38ED2A792974D81DDC498">146</mods:start>
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<mods:classification id="2045F23D3E6E482DDF13F8A590868937">journal article</mods:classification>
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<treatment id="FE5082769747FFE5FF3AFD745937D90B" ID-DOI="http://doi.org/10.5281/zenodo.5989240" ID-GBIF-Taxon="142524166" ID-Zenodo-Dep="5989240" LSID="urn:lsid:plazi:treatment:FE5082769747FFE5FF3AFD745937D90B" httpUri="http://treatment.plazi.org/id/FE5082769747FFE5FF3AFD745937D90B" lastPageId="3" lastPageNumber="149" pageId="2" pageNumber="148">
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<subSubSection id="3EE360EB9747FFE4FF3AFD745AB2D9B6" box="[151,672,695,722]" pageId="2" pageNumber="148" type="nomenclature">
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<paragraph id="764633609747FFE4FF3AFD745AB2D9B6" blockId="2.[151,1436,695,1137]" box="[151,672,695,722]" pageId="2" pageNumber="148">
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<heading id="2D0E840C9747FFE4FF3AFD745AB2D9B6" bold="true" box="[151,672,695,722]" fontSize="11" level="1" pageId="2" pageNumber="148" reason="1">
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<emphasis id="448DEF729747FFE4FF3AFD745AB2D9B6" bold="true" box="[151,672,695,722]" pageId="2" pageNumber="148">
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<taxonomicName id="B1F948E39747FFE4FF3AFD7459C8D9B5" ID-CoL="47F32" authority="Li & Ren & Wang, 2018" authorityName="Li & Ren & Wang" authorityYear="2018" box="[151,474,695,721]" class="Insecta" family="Osmylidae" genus="Nilionympha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="148" phylum="Arthropoda" rank="species" species="shantouensis" status="sp. nov.">
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<emphasis id="448DEF729747FFE4FF3AFD7459C8D9B5" bold="true" box="[151,474,695,721]" italics="true" pageId="2" pageNumber="148">Nilionympha shantouensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5FBE52099747FFE4FE4CFD7B5A2DD9B6" box="[481,575,696,722]" pageId="2" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="EEC22FE59747FFE4FDE2FD7B5A85D9B6" box="[591,663,696,722]" captionStart="FIGURE 2" captionStartId="2.[151,244,1947,1969]" captionTargetBox="[151,1436,1186,1927]" captionTargetId="figure@2.[151,1436,1186,1927]" captionTargetPageId="2" captionText="FIGURE 2. Enodinympha shantouensis sp. nov. (holotype CNU-NEU-NN200211). A. habitus photograph. B. line drawing of forewing.C.line drawing of hindwing.Scale bars:3 mm (A–C)." httpUri="https://zenodo.org/record/1206438/files/figure.png" pageId="2" pageNumber="148">Fig. 2</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3EE360EB9747FFE4FF3AFD195D4FD995" box="[151,1373,730,753]" pageId="2" pageNumber="148" type="etymology">
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<paragraph id="764633609747FFE4FF3AFD195D4FD995" blockId="2.[151,1436,695,1137]" box="[151,1373,730,753]" pageId="2" pageNumber="148">
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<emphasis id="448DEF729747FFE4FF3AFD195908D995" bold="true" box="[151,282,730,753]" pageId="2" pageNumber="148">Etymology:</emphasis>
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The species named after the fossil locality of Shantou Township, Ningcheng County,
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<collectingRegion id="B43DFD829747FFE4FB02FD195D4BD995" box="[1199,1369,730,753]" country="China" name="Nei Mongol" pageId="2" pageNumber="148">
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Inner
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<collectingCountry id="0EEE73F09747FFE4FB43FD195D4BD995" box="[1262,1369,730,753]" name="Mongolia" pageId="2" pageNumber="148">Mongolia</collectingCountry>
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</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="3EE360EB9747FFE4FF6AFD395A84D815" pageId="2" pageNumber="148" type="materials_examined">
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<paragraph id="764633609747FFE4FF6AFD395A1BD855" blockId="2.[151,1436,695,1137]" pageId="2" pageNumber="148">
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<emphasis id="448DEF729747FFE4FF6AFD395963D875" bold="true" box="[199,369,762,785]" pageId="2" pageNumber="148">Type material.</emphasis>
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Holotype, CNU-NEU-NN
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<date id="024715A09747FFE4FD37FD395AFAD875" box="[666,744,762,785]" pageId="2" pageNumber="148" value="11-02-20">200211</date>
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, with only well-preserved forewing and partly preserved hind wing, body fragmentary (
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<figureCitation id="EEC22FE59747FFE4FE0BFCD959EED855" box="[422,508,794,817]" captionStart="FIGURE 2" captionStartId="2.[151,244,1947,1969]" captionTargetBox="[151,1436,1186,1927]" captionTargetId="figure@2.[151,1436,1186,1927]" captionTargetPageId="2" captionText="FIGURE 2. Enodinympha shantouensis sp. nov. (holotype CNU-NEU-NN200211). A. habitus photograph. B. line drawing of forewing.C.line drawing of hindwing.Scale bars:3 mm (A–C)." httpUri="https://zenodo.org/record/1206438/files/figure.png" pageId="2" pageNumber="148">Fig. 2A</figureCitation>
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).
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</paragraph>
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<paragraph id="764633609747FFE4FF6AFCF95A84D815" blockId="2.[151,1436,695,1137]" pageId="2" pageNumber="148">
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<emphasis id="448DEF729747FFE4FF6AFCF959D0D835" bold="true" box="[199,450,826,849]" pageId="2" pageNumber="148">Locality and horizon.</emphasis>
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Daohugou Village, Shantou Township, Ningcheng County, Chifen City,
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<collectingRegion id="B43DFD829747FFE4FB46FCF95D85D835" box="[1259,1431,826,849]" country="China" name="Nei Mongol" pageId="2" pageNumber="148">
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Inner
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<collectingCountry id="0EEE73F09747FFE4FA80FCF95D85D835" box="[1325,1431,826,849]" name="Mongolia" pageId="2" pageNumber="148">Mongolia</collectingCountry>
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</collectingRegion>
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,
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<collectingCountry id="0EEE73F09747FFE4FF3AFC9958C8D815" box="[151,218,858,881]" name="China" pageId="2" pageNumber="148">China</collectingCountry>
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; Jiulongshan Formation, Middle Jurassic.
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</paragraph>
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</subSubSection>
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<subSubSection id="3EE360EB9747FFE4FF6AFCB95A85DF75" pageId="2" pageNumber="148" type="diagnosis">
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<paragraph id="764633609747FFE4FF6AFCB95A85DF75" blockId="2.[151,1436,695,1137]" pageId="2" pageNumber="148">
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<emphasis id="448DEF729747FFE4FF6AFCB9592ED8F5" bold="true" box="[199,316,890,913]" pageId="2" pageNumber="148">Diagnosis.</emphasis>
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Forewing with three rs-mp crossveins; twelve Rs branches; MA separating from Rs at basal quarter of wing length; MP forked near wing base; CuA deeply bifurcated before wing midlength; CuP simply pectinated, terminated before wing midlength. In hind wing, two rs-mp crossveins; Rs with eleven branches; MA separating from Rs before one-third of wing length; MP straight with forking apically; CuA simply forked after wing midlength; CuP slightly shorter than CuA, pectinately branched.
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</paragraph>
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</subSubSection>
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<subSubSection id="3EE360EB9747FFE5FF6AFBD95BECDAAB" lastPageId="3" lastPageNumber="149" pageId="2" pageNumber="148" type="description">
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<paragraph id="764633609747FFE4FF6AFBD95936DF15" blockId="2.[151,1436,695,1137]" pageId="2" pageNumber="148">
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<emphasis id="448DEF729747FFE4FF6AFBD95946DF55" bold="true" box="[199,340,1050,1073]" pageId="2" pageNumber="148">Description:</emphasis>
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Body length (from the apex of head to the terminalia of abdomen)
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<quantity id="B1019E859747FFE4FBE2FBD95CA7DF55" box="[1103,1205,1050,1073]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="148" unit="mm" value="12.3">12.3 mm</quantity>
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; head hypognathous, antenna extremely long, scape prominent, large and robust; pedicel small; flagellum filiform; compound eye large, but ocelli absent.
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</paragraph>
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<caption id="228663E89747FFE4FF3AF8585B1EDCAA" httpUri="https://zenodo.org/record/1206438/files/figure.png" pageId="2" pageNumber="148" startId="2.[151,244,1947,1969]" targetBox="[151,1436,1186,1927]" targetPageId="2">
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<paragraph id="764633609747FFE4FF3AF8585B1EDCAA" blockId="2.[151,1436,1947,1999]" pageId="2" pageNumber="148">
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<emphasis id="448DEF729747FFE4FF3AF8585903DCD5" bold="true" box="[151,273,1947,1969]" pageId="2" pageNumber="148">FIGURE 2.</emphasis>
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<taxonomicName id="B1F948E39747FFE4FEB1F8585A3ADCD5" authority="Li & Ren & Wang, 2018" authorityName="Li & Ren & Wang" authorityYear="2018" box="[284,552,1947,1969]" class="Insecta" family="Osmylidae" genus="Enodinympha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="148" phylum="Arthropoda" rank="species" species="shantouensis" status="sp. nov.">
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<emphasis id="448DEF729747FFE4FEB1F8585A3ADCD5" box="[284,552,1947,1969]" italics="true" pageId="2" pageNumber="148">Enodinympha shantouensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5FBE52099747FFE4FD9EF8585A97DCD5" box="[563,645,1947,1969]" pageId="2" pageNumber="148" rank="species">
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<emphasis id="448DEF729747FFE4FD9EF8585A6DDCD5" bold="true" box="[563,639,1947,1969]" pageId="2" pageNumber="148">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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<emphasis id="448DEF729747FFE4FD3DF8585A8ADCD5" bold="true" box="[656,664,1947,1969]" pageId="2" pageNumber="148">(</emphasis>
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holotype CNU-NEU-NN200211).
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<emphasis id="448DEF729747FFE4FC40F8585C17DCD5" bold="true" box="[1005,1029,1947,1969]" pageId="2" pageNumber="148">A.</emphasis>
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habitus photograph.
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<emphasis id="448DEF729747FFE4FB73F8585CE6DCD5" bold="true" box="[1246,1268,1947,1969]" pageId="2" pageNumber="148">B.</emphasis>
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line drawing of forewing.
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<emphasis id="448DEF729747FFE4FF54F87A5903DCAB" bold="true" box="[249,273,1977,1999]" pageId="2" pageNumber="148">C.</emphasis>
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line drawing of hind wing. Scale bars: 3 mm (
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<emphasis id="448DEF729747FFE4FD62F87A5AEFDCAB" bold="true" box="[719,765,1977,1999]" pageId="2" pageNumber="148">A–C</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph id="764633609746FFE5FF6AFF5B5B27DA0B" blockId="3.[151,1436,152,623]" pageId="3" pageNumber="149">
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Forewing:
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<quantity id="B1019E859746FFE5FE93FF5B59B3DBCB" box="[318,417,152,175]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="3" pageNumber="149" unit="mm" value="22.1">22.1 mm</quantity>
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long,
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<quantity id="B1019E859746FFE5FE47FF5B5A2CDBCB" box="[490,574,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="149" unit="mm" value="7.7">7.7 mm</quantity>
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wide, a row of markings along the subcostal space on membrane. Trichosors poorly developed apically; possibly one or two trichosors between each vein at apically posterior wing margin. Costal space relatively narrow. Subcostal veinlets simple, straight, rather widely spaced. Sc and R1 fused anterior to the wing apex. Rs originating nearly at wing base, zigzagged, with 11 long zigzagged branches. MA separating from Rs at one third of wing length. MP forked into MP1 and MP2 near wing base, straight for most of their length and dichotomously branched distally, with one complete row of cells between MP branches; CuA long, dichotomously branched deeply before midlength of wing; CuP pectinately branched (
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<figureCitation id="EEC22FE59746FFE5FD78FE9B5B3ADA0B" box="[725,808,344,367]" captionStart="FIGURE 2" captionStartId="2.[151,244,1947,1969]" captionTargetBox="[151,1436,1186,1927]" captionTargetId="figure@2.[151,1436,1186,1927]" captionTargetPageId="2" captionText="FIGURE 2. Enodinympha shantouensis sp. nov. (holotype CNU-NEU-NN200211). A. habitus photograph. B. line drawing of forewing.C.line drawing of hindwing.Scale bars:3 mm (A–C)." httpUri="https://zenodo.org/record/1206438/files/figure.png" pageId="3" pageNumber="149">Fig. 2B</figureCitation>
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).
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</paragraph>
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<paragraph id="764633609746FFE5FF6AFEBB5BECDAAB" blockId="3.[151,1436,152,623]" pageId="3" pageNumber="149">
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Hind wing elongated and relatively narrow,
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<quantity id="B1019E859746FFE5FD18FEBB5B0ADAEB" box="[693,792,376,399]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="3" pageNumber="149" unit="mm" value="20.2">20.2 mm</quantity>
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long,
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<quantity id="B1019E859746FFE5FCC9FEBB5BA8DAEB" box="[868,954,376,399]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="149" unit="mm" value="7.3">7.3 mm</quantity>
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wide, partly preserved, venation similar to forewing; Rs with eleven branches; MA separating from Rs before one-third of wing length; MP simple, bifurcated distally; CuA simply branched distally; CuP long, pectinately branched (
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<figureCitation id="EEC22FE59746FFE5FC30FE7B5BE0DAAB" box="[925,1010,440,463]" captionStart="FIGURE 2" captionStartId="2.[151,244,1947,1969]" captionTargetBox="[151,1436,1186,1927]" captionTargetId="figure@2.[151,1436,1186,1927]" captionTargetPageId="2" captionText="FIGURE 2. Enodinympha shantouensis sp. nov. (holotype CNU-NEU-NN200211). A. habitus photograph. B. line drawing of forewing.C.line drawing of hindwing.Scale bars:3 mm (A–C)." httpUri="https://zenodo.org/record/1206438/files/figure.png" pageId="3" pageNumber="149">Fig. 2C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="3EE360EB9746FFE5FF6AFE1B5937D90B" pageId="3" pageNumber="149" type="discussion">
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<paragraph id="764633609746FFE5FF6AFE1B5937D90B" blockId="3.[151,1436,152,623]" pageId="3" pageNumber="149">
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<emphasis id="448DEF729746FFE5FF6AFE1B5927DA8B" bold="true" box="[199,309,472,495]" pageId="3" pageNumber="149">Remarks.</emphasis>
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The new species can be attributed to
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<taxonomicName id="B1F948E39746FFE5FD79FE1B5B4CDA8B" baseAuthorityName="Ren & Engel" baseAuthorityYear="2007" box="[724,862,472,495]" class="Insecta" family="Osmylidae" genus="Nilionympha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="448DEF729746FFE5FD79FE1B5B4CDA8B" box="[724,862,472,495]" italics="true" pageId="3" pageNumber="149">Nilionympha</emphasis>
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</taxonomicName>
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for sharing the proximal divergence of MA from Rs and numerous Rs branches.
|
||
<taxonomicName id="B1F948E39746FFE5FE6BFE3B5A62D96B" authority="Li & Ren & Wang, 2018" authorityName="Li & Ren & Wang" authorityYear="2018" box="[454,624,504,527]" class="Insecta" family="Osmylidae" genus="Enodinympha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="149" phylum="Arthropoda" rank="species" species="shantouensis" status="sp. nov.">
|
||
<emphasis id="448DEF729746FFE5FE6BFE3B5A62D96B" box="[454,624,504,527]" italics="true" pageId="3" pageNumber="149">N. shantouensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5FBE52099746FFE5FDD5FE3B5ADCD96B" box="[632,718,504,527]" pageId="3" pageNumber="149" rank="species">
|
||
<emphasis id="448DEF729746FFE5FDD5FE3B5AD5D96B" bold="true" box="[632,711,504,527]" pageId="3" pageNumber="149">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is easily identified by the presence of a row of dark spots (at least eight black spots) along with Sc and R1 that are evidently absent in
|
||
<taxonomicName id="B1F948E39746FFE5FC2FFDDB5C14D94B" authorityName="Ren & Engel" authorityYear="2007" box="[898,1030,536,559]" class="Insecta" family="Osmylidae" genus="Nilionympha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="149" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="448DEF729746FFE5FC2FFDDB5C14D94B" box="[898,1030,536,559]" italics="true" pageId="3" pageNumber="149">N. pulchella</emphasis>
|
||
</taxonomicName>
|
||
. Additionally the new species is also differentiated from
|
||
<taxonomicName id="B1F948E39746FFE5FEC5FDFB59F9D92B" authorityName="Ren & Engel" authorityYear="2007" box="[360,491,568,591]" class="Insecta" family="Osmylidae" genus="Nilionympha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="149" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="448DEF729746FFE5FEC5FDFB59F9D92B" box="[360,491,568,591]" italics="true" pageId="3" pageNumber="149">N. pulchella</emphasis>
|
||
</taxonomicName>
|
||
according to the position of fusion of Sc and R
|
||
<quantity id="B1019E859746FFE5FC43FDFB5C06D92B" box="[1006,1044,568,591]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="149" unit="in" value="1.0">1 in</quantity>
|
||
forewing and simply branched CuA in hind wing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |