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<document ID-DOI="http://dx.doi.org/10.5733/afin.049.0105" ID-GBIF-Dataset="01968b77-015c-45df-aa06-4a7b1090904e" ID-ISSN="2305-2562" ID-Zenodo-Dep="7667189" checkinTime="1676639426290" checkinUser="felipe" docAuthor="Morita, Shelah" docDate="2008" docId="B17087D25930FFFE31DFFB88FB88FBCC" docLanguage="en" docName="AfrInvertebr.49.1.129-158.pdf" docOrigin="African Invertebrates 49 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.2:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="2" docTitle="Philoliche (Philoliche) aethiopica" docType="key" docVersion="4" lastPageNumber="153" masterDocId="4D49FFAA5928FFE73120FF81FFD8FFD3" masterDocTitle="A revision of the Philoliche aethiopica species complex (Diptera: Tabanidae)" masterLastPageNumber="158" masterPageNumber="129" pageNumber="152" updateTime="1677095106358" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>A revision of the Philoliche aethiopica species complex (Diptera: Tabanidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Morita, Shelah</mods:namePart>
<mods:affiliation>Population Biology Graduate Group, University of California at Davis, 1 Shields Ave., Davis, CA 95616, USA; Current address: Department of Botany &amp; Zoology, Stellenbosch University, P. Bag X 1, Matieland, 7602 South Africa</mods:affiliation>
<mods:nameIdentifier type="email">simorita@ncsu.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>49</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>158</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.5733/afin.049.0105</mods:identifier>
<mods:identifier type="GBIF-Dataset">01968b77-015c-45df-aa06-4a7b1090904e</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7667189</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7649421" ID-Zenodo-Dep="7649421" LSID="urn:lsid:plazi:treatment:B17087D25930FFFE31DFFB88FB88FBCC" httpUri="http://treatment.plazi.org/id/B17087D25930FFFE31DFFB88FB88FBCC" lastPageId="25" lastPageNumber="153" pageId="24" pageNumber="152">
<subSubSection box="[255,1067,1033,1058]" pageId="24" pageNumber="152" type="nomenclature">
<paragraph blockId="24.[255,1067,1033,1058]" box="[255,1067,1033,1058]" pageId="24" pageNumber="152">
<heading box="[255,1067,1033,1058]" centered="true" fontSize="36" level="2" pageId="24" pageNumber="152" reason="5">
Key to species in the
<taxonomicName ID-CoL="8D9WF" ID-ENA="415279" baseAuthorityName="Thunberg" baseAuthorityYear="1789" box="[496,876,1033,1058]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="152" phylum="Arthropoda" rank="species" species="aethiopica" subGenus="Philoliche">
<emphasis box="[496,611,1033,1057]" italics="true" pageId="24" pageNumber="152">Philoliche</emphasis>
(
<emphasis box="[627,742,1033,1057]" italics="true" pageId="24" pageNumber="152">Philoliche</emphasis>
)
<emphasis box="[758,876,1033,1057]" italics="true" pageId="24" pageNumber="152">aethiopica</emphasis>
</taxonomicName>
species complex
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="153" pageId="24" pageNumber="152" type="key">
<keyStep pageId="24" pageNumber="152">
<paragraph blockId="24.[181,1142,1075,1668]" box="[181,1139,1075,1100]" pageId="24" pageNumber="152">
<keyLead box="[181,1139,1075,1100]" pageId="24" pageNumber="152">1 Holoptic: eyes meeting or nearly meeting in front (males) .................................. 2</keyLead>
</paragraph>
<paragraph blockId="24.[181,1142,1075,1668]" pageId="24" pageNumber="152">
<keyLead pageId="24" pageNumber="152"> Dichoptic: eyes clearly separated, not touching or close to touching at any point (females) ............................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="24" pageNumber="152">
<paragraph blockId="24.[181,1142,1075,1668]" pageId="24" pageNumber="152">
<keyLead pageId="24" pageNumber="152">
2 Fore tarsi with long process on first and second segments (lappets); eye facets all of similar size or only gradually increasing in size from ventral to dorsal; eyes meeting until slightly above antennae leaving a small triangle of a frons (
<emphasis bold="true" box="[1020,1142,1243,1268]" pageId="24" pageNumber="152">aethiopica</emphasis>
group).................................................................................................................... 4
</keyLead>
</paragraph>
<paragraph blockId="24.[181,1142,1075,1668]" pageId="24" pageNumber="152">
<keyLead pageId="24" pageNumber="152">
Fore tarsi without long process on first and second segments; if process exists, it does not extend beyond the distal margin of the third tarsomere; eye facets abruptly increasing in size from ventral to dorsal forming a demarcating, horizontal line; eyes contiguous to near the base of antennae, no vestige of frons remaining (
<emphasis bold="true" box="[1048,1141,1403,1428]" pageId="24" pageNumber="152">rondani</emphasis>
group).................................................................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="24" pageNumber="152">
<paragraph blockId="24.[181,1142,1075,1668]" pageId="24" pageNumber="152">
<keyLead pageId="24" pageNumber="152">
3 Ratio of frons width at base to frons width at vertex greater than or equal to 2.1 (
<figureCitation box="[238,319,1507,1532]" captionStart="Fig" captionStartId="6.[182,215,1575,1595]" captionTargetBox="[184,1134,1216,1556]" captionTargetId="figure-422@6.[178,1138,1210,1556]" captionTargetPageId="6" captionText="Fig. 3. Frons divergence in females of (a) P. rondani and (b) P. aethiopica. The index of frons divergence is the ratio of the width of the frons at the clypeal suture (cs) to the width of the frons at the vertex (v). Note also that most P. aethiopica also have an obtuse angle in the inner eye margin (ea), though this is variable." figureDoi="http://doi.org/10.5281/zenodo.7667199" httpUri="https://zenodo.org/record/7667199/files/figure.png" pageId="24" pageNumber="152">Fig. 3b</figureCitation>
) (
<emphasis bold="true" box="[344,467,1507,1532]" pageId="24" pageNumber="152">aethiopica</emphasis>
group) ................................................................................. 4
</keyLead>
</paragraph>
<paragraph blockId="24.[181,1142,1075,1668]" pageId="24" pageNumber="152">
<keyLead pageId="24" pageNumber="152">
Ratio of frons width at base to frons width at vertex less than 2.1 (
<figureCitation box="[946,1024,1539,1564]" captionStart="Fig" captionStartId="6.[182,215,1575,1595]" captionTargetBox="[184,1134,1216,1556]" captionTargetId="figure-422@6.[178,1138,1210,1556]" captionTargetPageId="6" captionText="Fig. 3. Frons divergence in females of (a) P. rondani and (b) P. aethiopica. The index of frons divergence is the ratio of the width of the frons at the clypeal suture (cs) to the width of the frons at the vertex (v). Note also that most P. aethiopica also have an obtuse angle in the inner eye margin (ea), though this is variable." figureDoi="http://doi.org/10.5281/zenodo.7667199" httpUri="https://zenodo.org/record/7667199/files/figure.png" pageId="24" pageNumber="152">Fig. 3a</figureCitation>
) (
<emphasis bold="true" box="[1047,1141,1539,1564]" pageId="24" pageNumber="152">rondani</emphasis>
group).................................................................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep lastPageId="25" lastPageNumber="153" pageId="24" pageNumber="152">
<paragraph blockId="24.[181,1142,1075,1668]" lastBlockId="25.[144,1104,206,1055]" lastPageId="25" lastPageNumber="153" pageId="24" pageNumber="152">
<keyLead lastPageId="25" lastPageNumber="153" pageId="24" pageNumber="152">
4 Dorsum of mesothorax with distinct, narrow, grey sublateral stripes. Females: females are not yet known for this species; this is their provisional outcome in this key as predicted by the distribution of other characters in the group ..................... ...................................................................................................
<taxonomicName authority="(Austen)" baseAuthorityName="Austen" box="[898,1101,238,263]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="153" phylum="Arthropoda" rank="species" species="formosa">
<emphasis bold="true" box="[898,996,238,263]" pageId="25" pageNumber="153">formosa</emphasis>
(Austen)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="25.[144,1104,206,1055]" pageId="25" pageNumber="153">
<keyLead pageId="25" pageNumber="153">
Dorsum of mesothorax without stripes, or with indistinct, broad, dull reddish or greyish ones .......................................................................
<taxonomicName authority="(Thunberg)" baseAuthorityName="Thunberg" baseAuthorityYear="1789" box="[844,1101,302,327]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="153" phylum="Arthropoda" rank="species" species="aethiopica">
<emphasis bold="true" box="[844,967,302,327]" pageId="25" pageNumber="153">aethiopica</emphasis>
(Thunberg)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="25" pageNumber="153">
<paragraph blockId="25.[144,1104,206,1055]" pageId="25" pageNumber="153">
<keyLead pageId="25" pageNumber="153">
5 Dorsum of mesothorax with grey band on lateral margins and with grey on posterior margin; scutellum greyish pink (
<figureCitation box="[556,620,374,399]" captionStart="Fig" captionStartId="2.[182,215,1599,1619]" captionTargetBox="[246,1079,905,1580]" captionTargetId="figure-318@2.[246,1079,905,1580]" captionTargetPageId="2" captionText="Fig. 1. Dorsal view of (a) Philoliche rubiginosa and (b) P. elegans showing differences in patterning of the mesoscutum and frons. White arrows point to the transition between colouring of the grey hind margin of the mesoscutum and the main part of the mesothorax." figureDoi="http://doi.org/10.5281/zenodo.7667191" httpUri="https://zenodo.org/record/7667191/files/figure.png" pageId="25" pageNumber="153">Fig. 1</figureCitation>
) ............................................................... 6
</keyLead>
</paragraph>
<paragraph blockId="25.[144,1104,206,1055]" pageId="25" pageNumber="153">
<keyLead pageId="25" pageNumber="153"> Dorsum of mesothorax with grey band on lateral margins, but not on posterior margin; scutellum concolourous with centre of mesothorax, not greyish pink .... 7</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="25" pageNumber="153">
<paragraph blockId="25.[144,1104,206,1055]" pageId="25" pageNumber="153">
<keyLead pageId="25" pageNumber="153">
6 Grey posterior margin of mesothorax sharply contrasting with brown at centre; frons generally light coloured and pollinose, not wholly shiny (
<figureCitation box="[808,885,510,535]" captionStart="Fig" captionStartId="2.[182,215,1599,1619]" captionTargetBox="[246,1079,905,1580]" captionTargetId="figure-318@2.[246,1079,905,1580]" captionTargetPageId="2" captionText="Fig. 1. Dorsal view of (a) Philoliche rubiginosa and (b) P. elegans showing differences in patterning of the mesoscutum and frons. White arrows point to the transition between colouring of the grey hind margin of the mesoscutum and the main part of the mesothorax." figureDoi="http://doi.org/10.5281/zenodo.7667191" httpUri="https://zenodo.org/record/7667191/files/figure.png" pageId="25" pageNumber="153">Fig. 1b</figureCitation>
) ....
<taxonomicName authority="(Bigot)" baseAuthorityName="Bigot" box="[932,1101,510,535]" class="Insecta" family="Tabanidae" genus="Mycteromyia" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="153" phylum="Arthropoda" rank="species" species="elegans">
<emphasis bold="true" box="[932,1016,510,535]" pageId="25" pageNumber="153">elegans</emphasis>
(Bigot)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="25.[144,1104,206,1055]" pageId="25" pageNumber="153">
<keyLead pageId="25" pageNumber="153">
Grey posterior margin of mesothorax gradually blending into brown anteriorly; frons generally dark and shiny. If frons pollinose, then only along eye margin (
<figureCitation box="[200,278,606,631]" captionStart="Fig" captionStartId="2.[182,215,1599,1619]" captionTargetBox="[246,1079,905,1580]" captionTargetId="figure-318@2.[246,1079,905,1580]" captionTargetPageId="2" captionText="Fig. 1. Dorsal view of (a) Philoliche rubiginosa and (b) P. elegans showing differences in patterning of the mesoscutum and frons. White arrows point to the transition between colouring of the grey hind margin of the mesoscutum and the main part of the mesothorax." figureDoi="http://doi.org/10.5281/zenodo.7667191" httpUri="https://zenodo.org/record/7667191/files/figure.png" pageId="25" pageNumber="153">Fig. 1a</figureCitation>
) .......................................................................................
<taxonomicName authority="Dias" authorityName="Dias" box="[915,1101,606,631]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="153" phylum="Arthropoda" rank="species" species="rubiginosa">
<emphasis bold="true" box="[915,1043,606,631]" pageId="25" pageNumber="153">rubiginosa</emphasis>
Dias
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="25" pageNumber="153">
<paragraph blockId="25.[144,1104,206,1055]" pageId="25" pageNumber="153">
<keyLead pageId="25" pageNumber="153">
7 All legs similarly coloured on proximal segments: coxae grey, paler dorsally and with white hairs; trochanters black; femora orange. Posterolateral margin of segments 4 and 5 with tufts of black or white hairs as follows: 4 white only, 5 black and white .......................................................................
<taxonomicName authority="(Bertoloni)" baseAuthorityName="Bertoloni" box="[875,1101,742,767]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="153" phylum="Arthropoda" rank="species" species="rondani">
<emphasis bold="true" box="[875,969,742,767]" pageId="25" pageNumber="153">rondani</emphasis>
(Bertoloni)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="25.[144,1104,206,1055]" pageId="25" pageNumber="153">
<keyLead pageId="25" pageNumber="153">
Fore and mid legs similarly coloured, hind legs different: fore and mid coxae grey to pinkish grey, paler dorsally and with white and a few black hairs, trochanters black; hind coxa yellow, trochanters orange; fore and mid femora orange, hind femur yellow, same as hind coxa. Tibia of foreleg and midleg paler than femora. Hind tibia darker than hind femur, with black hairs. Posterolateral margin of segments 4 and 5 with tufts of black or white hairs as follows: 4 black and white, 5 black only. Third abdominal segment relatively paler dorsally than the other abdominal segments. Proboscis usually 2× greater in length than
<taxonomicName box="[897,1010,998,1022]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="153" phylum="Arthropoda" rank="species" species="rondani">
<emphasis box="[897,1010,998,1022]" italics="true" pageId="25" pageNumber="153">P. rondani</emphasis>
</taxonomicName>
. Known from the Albany
<collectingRegion box="[377,452,1030,1055]" country="France" name="Centre" pageId="25" pageNumber="153">Centre</collectingRegion>
of endemism only.........
<taxonomicName authority="(Ricardo)" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[717,1006,1030,1055]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="153" phylum="Arthropoda" rank="species" species="umbratipennis" status="stat. rev.">
<emphasis bold="true" box="[717,893,1030,1055]" pageId="25" pageNumber="153">umbratipennis</emphasis>
(Ricardo)
</taxonomicName>
<taxonomicNameLabel box="[1011,1104,1030,1055]" pageId="25" pageNumber="153">stat. rev.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>