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<document id="8A9B78DF24ACC921853775C1A2131B90" ID-DOI="10.11646/zootaxa.5326.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8243811" ID-ZooBank="2EC25BF0-D36F-4029-AD1C-A9B62A668FEE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1691944079938" checkinUser="plazi" docAuthor="Werner, Maria Johanna, Hausmann, Axel, Kostjuk, Igor, Wanke, Dominic &amp; Rajaei, Hossein" docDate="2023" docId="A420BA7FF05EFFE2FF1EFB9D7B12F8CA" docLanguage="en" docName="zootaxa.5326.1.1.pdf" docOrigin="Zootaxa 5326 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5326.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Phaselia deliciosaria" docType="treatment" docVersion="6" lastPageNumber="24" masterDocId="5819C207F049FFF5FF89FFA27F76FFE3" masterDocTitle="Integrative taxonomic revision of the genus Phaselia Guenée, [1858] (Geometridae: Ennominae) in the Middle East and Central Asia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="24" updateTime="1699557214275" updateUser="plazi">
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<mods:title id="2F96CB5F88DACE0478BF3DB464D058AB">Integrative taxonomic revision of the genus Phaselia Guenée, [1858] (Geometridae: Ennominae) in the Middle East and Central Asia</mods:title>
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<mods:namePart id="734685FEA0D3247309EE169781F7C571">Werner, Maria Johanna</mods:namePart>
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<mods:namePart id="340F11C4C1810D4B44D0B0D89DC06304">Hausmann, Axel</mods:namePart>
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<mods:namePart id="F2E53F70486DBF51CF9A62256BEC50C6">Wanke, Dominic</mods:namePart>
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<mods:namePart id="5C949EA8D0095746D6E24F46C0805CE4">Rajaei, Hossein</mods:namePart>
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<treatment id="A420BA7FF05EFFE2FF1EFB9D7B12F8CA" ID-DOI="http://doi.org/10.5281/zenodo.8247699" ID-Zenodo-Dep="8247699" LSID="urn:lsid:plazi:treatment:A420BA7FF05EFFE2FF1EFB9D7B12F8CA" httpUri="http://treatment.plazi.org/id/A420BA7FF05EFFE2FF1EFB9D7B12F8CA" lastPageNumber="24" pageId="23" pageNumber="24">
<subSubSection id="649358E2F05EFFE2FF1EFB9D7C98FB42" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="2C360B69F05EFFE2FF1EFB9D7D67FBBA" blockId="23.[151,1437,1087,1401]" box="[151,529,1087,1113]" pageId="23" pageNumber="24">
<heading id="777EBC05F05EFFE2FF1EFB9D7D67FBBA" box="[151,529,1087,1113]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="5">
<taxonomicName id="EB8970EAF05EFFE2FF1EFB9D7E4AFBBA" ID-CoL="4FVZ2" ID-ENA="1007372" baseAuthorityName="Lederer" baseAuthorityYear="1855" box="[151,316,1087,1113]" class="Insecta" family="Geometridae" genus="Phaselia" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="deliciosaria">
<emphasis id="1EFDD77BF05EFFE2FF1EFB9D7FDEFBBB" box="[151,168,1087,1112]" italics="true" pageId="23" pageNumber="24">P</emphasis>
.
<emphasis id="1EFDD77BF05EFFE2FF3FFB9D7E4AFBBA" box="[182,316,1087,1113]" italics="true" pageId="23" pageNumber="24">deliciosaria</emphasis>
</taxonomicName>
vs. (
<taxonomicName id="EB8970EAF05EFFE2FEFAFB9D7D7EFBBA" authorityName="Hausmann" authorityYear="1996" box="[371,520,1087,1113]" class="Insecta" family="Geometridae" genus="Phaselia" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="joestleinae">
<emphasis id="1EFDD77BF05EFFE2FEFAFB9D7D7EFBBA" box="[371,520,1087,1113]" italics="true" pageId="23" pageNumber="24">P. joestleinae</emphasis>
</taxonomicName>
)
</heading>
</paragraph>
<paragraph id="2C360B69F05EFFE2FF4EFBC17C98FB42" blockId="23.[151,1437,1087,1401]" pageId="23" pageNumber="24">
<emphasis id="1EFDD77BF05EFFE2FF4EFBC17ED0FB9E" box="[199,422,1123,1149]" italics="true" pageId="23" pageNumber="24">External characters</emphasis>
(figs 5053; 5658). On forewing antemedial line with three acute tips, middle one longer (antemedial line with rounded or acute tips, costal tooth reduced to a curve).
</paragraph>
</subSubSection>
<subSubSection id="649358E2F05EFFE2FF4EFB097CF7FA06" pageId="23" pageNumber="24" type="description">
<paragraph id="2C360B69F05EFFE2FF4EFB097FAEFAD2" blockId="23.[151,1437,1087,1401]" pageId="23" pageNumber="24">
<emphasis id="1EFDD77BF05EFFE2FF4EFB097E18FB26" box="[199,366,1195,1221]" italics="true" pageId="23" pageNumber="24">Male genitalia</emphasis>
(figs 103105; 107108). Sacculus triangular; spines gradually enlarging towards the ventral tip of sacculus; aedeagus with a long, straight, unbent cornutus (sacculus slightly oval, smoothly transitioning to valvula; without or with very small and fine, evenly distributed spines on the sacculus; claw-like cornutus strongly bent).
</paragraph>
<paragraph id="2C360B69F05EFFE2FF4EFA997E90FA9A" blockId="23.[151,1437,1087,1401]" pageId="23" pageNumber="24">
<emphasis id="1EFDD77BF05EFFE2FF4EFA997EF1FAB6" box="[199,391,1339,1365]" italics="true" pageId="23" pageNumber="24">Female genitalia</emphasis>
(figs 140; 142143). Antrum roundish to triangular, tapering towards corpus bursae (antrum small, roundish to triangular).
</paragraph>
<paragraph id="2C360B69F05EFFE2FF4EFA057CF7FA06" blockId="23.[151,1437,1447,1833]" pageId="23" pageNumber="24">
<emphasis id="1EFDD77BF05EFFE2FF4EFA057E3FFA22" bold="true" box="[199,329,1447,1473]" pageId="23" pageNumber="24">Phenology.</emphasis>
Investigated specimens have been collected from late April to late July. Bi- or trivoltine, early March to late July, August, and early September to late November.
</paragraph>
</subSubSection>
<subSubSection id="649358E2F05EFFE2FF4EFA4C7E97F996" pageId="23" pageNumber="24" type="biology_ecology">
<paragraph id="2C360B69F05EFFE2FF4EFA4C7A18F9EA" blockId="23.[151,1437,1447,1833]" box="[199,1390,1518,1545]" pageId="23" pageNumber="24">
<emphasis id="1EFDD77BF05EFFE2FF4EFA4C7E51F9EB" bold="true" box="[199,295,1518,1544]" pageId="23" pageNumber="24">Biology.</emphasis>
Larval stages possibly on
<taxonomicName id="EB8970EAF05EFFE2FDD8FA4D7C2FF9EA" box="[593,857,1519,1545]" class="Gnetopsida" family="Ephedraceae" genus="Ephedra" kingdom="Plantae" order="Ephedrales" pageId="23" pageNumber="57" phylum="Tracheophyta" rank="subGenus" subGenus="Ephedraceae">
<emphasis id="1EFDD77BF05EFFE2FDD8FA4D7DC7F9EA" box="[593,689,1519,1545]" italics="true" pageId="23" pageNumber="24">Ephedra</emphasis>
(Ephedraceae)
</taxonomicName>
(
<bibRefCitation id="48187698F05EFFE2FCE1FA4D7B12F9EA" author="Hausmann, A. &amp; Stadie, D. &amp; Fiebig, R." box="[872,1124,1519,1545]" pageId="23" pageNumber="24" pagination="7 - 742" refId="ref30591" refString="Hausmann, A., Stadie, D. &amp; Fiebig, R. (2016) Family Geometridae Leach [1815]. In: Hacker, H. H. (Ed.), Systematic and Illustrated Catalogue of the Macroheterocera and Superfamilies Cossoidea Leach, [1815], Zygaenoidea Latreille, 1809, Thyridoidea Herrich-Schaffer, 1846 and Hyblaeoidea Hampson, 1903 of the Arabian Peninsula, with a survey of their distribution (Lepidoptera). Esperiana, 20 (1), pp. 7 - 742." type="journal article" year="2016">
Hausmann
<emphasis id="1EFDD77BF05EFFE2FC60FA4D7B57F9EA" box="[1001,1057,1519,1545]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2016
</bibRefCitation>
;
<bibRefCitation id="48187698F05EFFE2FBE6FA4D7A10F9EA" author="Hausmann, A. &amp; Kravchenko, V. &amp; Muller, G." box="[1135,1382,1519,1545]" pageId="23" pageNumber="24" pagination="1 - 256" refId="ref30458" refString="Hausmann, A., Kravchenko, V. &amp; Muller, G. (2020) The Lepidoptera of Israel. Vol. 3. Geometridae. Proceedings of the Museum Witt Munich, 2020, pp. 1 - 256." type="journal article" year="2020">
Hausmann
<emphasis id="1EFDD77BF05EFFE2FB64FA4D7A50F9EA" box="[1261,1318,1519,1545]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2020
</bibRefCitation>
)
</paragraph>
<paragraph id="2C360B69F05EFFE2FF4EF9B07E97F996" blockId="23.[151,1437,1447,1833]" pageId="23" pageNumber="24">
<emphasis id="1EFDD77BF05EFFE2FF4EF9B07E5CF9CF" bold="true" box="[199,298,1554,1580]" pageId="23" pageNumber="24">Habitat.</emphasis>
Investigated specimens have been collected at altitudes from sea-level (
<collectingCountry id="549E4BF9F05EFFE2FBD6F9B17BD6F9CE" box="[1119,1184,1555,1581]" name="Israel" pageId="23" pageNumber="24">Israel</collectingCountry>
) up to
<quantity id="EB71A68CF05EFFE2FB7AF9B17A3FF9CE" box="[1267,1353,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="24" unit="m" value="2000.0">2000 m</quantity>
(North
<collectingCountry id="549E4BF9F05EFFE2FF1EF9957F96F9B2" box="[151,224,1591,1617]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
). Xerothermophilous, psammophilous. Deserts and semi-deserts, Mediterranean vegetation, and coasts (see also:
<bibRefCitation id="48187698F05EFFE2FF5AF9F97EA3F996" author="Hausmann, A. &amp; Kravchenko, V. &amp; Muller, G." box="[211,469,1627,1653]" pageId="23" pageNumber="24" pagination="1 - 256" refId="ref30458" refString="Hausmann, A., Kravchenko, V. &amp; Muller, G. (2020) The Lepidoptera of Israel. Vol. 3. Geometridae. Proceedings of the Museum Witt Munich, 2020, pp. 1 - 256." type="journal article" year="2020">
Hausmann
<emphasis id="1EFDD77BF05EFFE2FEDBF9F97EFCF996" box="[338,394,1627,1653]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
, 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="649358E2F05EFFE2FF4EF9DC7B12F8CA" pageId="23" pageNumber="24" type="distribution">
<paragraph id="2C360B69F05EFFE2FF4EF9DC7B74F95E" blockId="23.[151,1437,1447,1833]" pageId="23" pageNumber="24">
<emphasis id="1EFDD77BF05EFFE2FF4EF9DC7E17F97B" bold="true" box="[199,353,1662,1688]" pageId="23" pageNumber="24">Distribution.</emphasis>
From South
<collectingCountry id="549E4BF9F05EFFE2FE74F9DD7D38F97A" box="[509,590,1663,1689]" name="Turkey" pageId="23" pageNumber="24">Turkey</collectingCountry>
,
<collectingCountry id="549E4BF9F05EFFE2FDD4F9DD7DEDF97A" box="[605,667,1663,1689]" name="Syria" pageId="23" pageNumber="24">Syria</collectingCountry>
,
<collectingCountry id="549E4BF9F05EFFE2FD20F9DD7C78F97A" box="[681,782,1663,1689]" name="Lebanon" pageId="23" pageNumber="24">Lebanon</collectingCountry>
,
<collectingCountry id="549E4BF9F05EFFE2FC94F9DD7C2DF97A" box="[797,859,1663,1689]" name="Israel" pageId="23" pageNumber="24">Israel</collectingCountry>
and
<collectingCountry id="549E4BF9F05EFFE2FC11F9DD7C95F97A" box="[920,995,1663,1689]" name="Jordan" pageId="23" pageNumber="24">Jordan</collectingCountry>
to Saudi-Arabia and Northeast
<collectingCountry id="549E4BF9F05EFFE2FAD1F9DD7AEAF97A" box="[1368,1436,1663,1689]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
(
<bibRefCitation id="48187698F05EFFE2FF16F9017ED7F95E" author="Hausmann, A. &amp; Stadie, D. &amp; Fiebig, R." box="[159,417,1699,1725]" pageId="23" pageNumber="24" pagination="7 - 742" refId="ref30591" refString="Hausmann, A., Stadie, D. &amp; Fiebig, R. (2016) Family Geometridae Leach [1815]. In: Hacker, H. H. (Ed.), Systematic and Illustrated Catalogue of the Macroheterocera and Superfamilies Cossoidea Leach, [1815], Zygaenoidea Latreille, 1809, Thyridoidea Herrich-Schaffer, 1846 and Hyblaeoidea Hampson, 1903 of the Arabian Peninsula, with a survey of their distribution (Lepidoptera). Esperiana, 20 (1), pp. 7 - 742." type="journal article" year="2016">
Hausmann
<emphasis id="1EFDD77BF05EFFE2FE96F9017E2EF95E" box="[287,344,1699,1725]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
, 2016
</bibRefCitation>
;
<bibRefCitation id="48187698F05EFFE2FE25F9017D09F95E" author="Laszlo, G. M. &amp; Lodl, M. &amp; Gaal-Haszler, S. &amp; Galsworthy, A. &amp; Ronkay, G. &amp; Ronkay, L. &amp; Varga, Z." box="[428,639,1699,1725]" pageId="23" pageNumber="24" refId="ref30962" refString="Laszlo, G. M., Lodl, M., Gaal-Haszler, S., Galsworthy, A., Ronkay, G., Ronkay, L., &amp; Varga, Z. (2018) The Vartian Collection Part IV. Geometridae. Fibigeriana. Vol. 4. Heterocera Press, Budapest, pp. 1 - 166." type="book" year="2018">
László
<emphasis id="1EFDD77BF05EFFE2FE77F9017D40F95E" box="[510,566,1699,1725]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
, 2018
</bibRefCitation>
;
<bibRefCitation id="48187698F05EFFE2FD03F9017CF4F95E" author="Hausmann, A. &amp; Kravchenko, V. &amp; Muller, G." box="[650,898,1699,1725]" pageId="23" pageNumber="24" pagination="1 - 256" refId="ref30458" refString="Hausmann, A., Kravchenko, V. &amp; Muller, G. (2020) The Lepidoptera of Israel. Vol. 3. Geometridae. Proceedings of the Museum Witt Munich, 2020, pp. 1 - 256." type="journal article" year="2020">
Hausmann
<emphasis id="1EFDD77BF05EFFE2FC80F9017C37F95E" box="[777,833,1699,1725]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2020
</bibRefCitation>
) (fig. 144).
</paragraph>
<paragraph id="2C360B69F05EFFE2FF4EF9647B12F8CA" blockId="23.[151,1437,1447,1833]" pageId="23" pageNumber="24">
<emphasis id="1EFDD77BF05EFFE2FF4EF9647EFCF903" bold="true" box="[199,394,1734,1760]" pageId="23" pageNumber="24">DNA barcoding.</emphasis>
Genetic distances from morphologically most similar species:
<taxonomicName id="EB8970EAF05EFFE2FBD9F9657A44F902" authorityName="Hausmann" authorityYear="1996" box="[1104,1330,1735,1761]" class="Insecta" family="Geometridae" genus="Phaselia" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="joestleinae">
<emphasis id="1EFDD77BF05EFFE2FBD9F9657A44F902" box="[1104,1330,1735,1761]" italics="true" pageId="23" pageNumber="24">Phaselia joestleinae</emphasis>
</taxonomicName>
(7.94%),
<taxonomicName id="EB8970EAF05EFFE2FF1EF9497E4CF8E6" authorityName="Werner &amp; Hausmann &amp; Kostjuk &amp; Wanke &amp; Rajaei" authorityYear="2023" box="[151,314,1771,1797]" class="Insecta" family="Geometridae" genus="Phaselia" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="erika" subSpecies="jonubi">
<emphasis id="1EFDD77BF05EFFE2FF1EF9497E4CF8E6" box="[151,314,1771,1797]" italics="true" pageId="23" pageNumber="24">P. erika jonubi</emphasis>
</taxonomicName>
<emphasis id="1EFDD77BF05EFFE2FEC9F9497ED2F8E6" bold="true" box="[320,420,1771,1797]" pageId="23" pageNumber="24">ssp. nov.</emphasis>
(7.58%),
<emphasis id="1EFDD77BF05EFFE2FD9DF9497D5FF8E7" box="[532,553,1771,1796]" italics="true" pageId="23" pageNumber="24">P.</emphasis>
sp.
<taxonomicName id="EB8970EAF05EFFE2FDDCF9497D88F8E6" baseAuthorityName="Lederer" baseAuthorityYear="1855" box="[597,766,1771,1797]" class="Insecta" family="Geometridae" genus="Phaselia" isUncertain="true" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="deliciosaria">
cf.
<emphasis id="1EFDD77BF05EFFE2FDF1F9497D88F8E6" box="[632,766,1771,1797]" italics="true" pageId="23" pageNumber="24">deliciosaria</emphasis>
</taxonomicName>
(3.09%). Genetically closest species in percentage terms:
<taxonomicName id="EB8970EAF05EFFE2FA01F9497E4FF8CA" authorityName="Werner &amp; Hausmann &amp; Kostjuk &amp; Wanke &amp; Rajaei" authorityYear="2023" baseAuthorityName="Lederer" baseAuthorityYear="1855" class="Insecta" family="Geometridae" genus="Phaselia" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="phaeoleucaria" status="stat. nov.">
<emphasis id="1EFDD77BF05EFFE2FA01F9497E4FF8CA" italics="true" pageId="23" pageNumber="24">P. phaeoleucaria</emphasis>
</taxonomicName>
<emphasis id="1EFDD77BF05EFFE2FEC9F8AD7ED3F8CA" bold="true" box="[320,421,1807,1833]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="05CE6A00F05EFFE2FEC9F8AD7ED3F8CA" box="[320,421,1807,1833]" pageId="23" pageNumber="24">stat. rev.</taxonomicNameLabel>
</emphasis>
(1.99 %) and
<emphasis id="1EFDD77BF05EFFE2FDCCF8AD7D2CF8CB" box="[581,602,1807,1832]" italics="true" pageId="23" pageNumber="24">P.</emphasis>
sp.
<taxonomicName id="EB8970EAF05EFFE2FD0EF8AD7C47F8CA" baseAuthorityName="Lederer" baseAuthorityYear="1855" box="[647,817,1807,1833]" class="Insecta" family="Geometridae" genus="Phaselia" isUncertain="true" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="deliciosaria">
cf.
<emphasis id="1EFDD77BF05EFFE2FD22F8AD7C47F8CA" box="[683,817,1807,1833]" italics="true" pageId="23" pageNumber="24">deliciosaria</emphasis>
</taxonomicName>
(3.09 %) (fig. 145,
<tableCitation id="610B3ED2F05EFFE2FB87F8AD7B25F8CA" box="[1038,1107,1807,1833]" captionStart="TABLE 1" captionStartId="50.[151,239,992,1016]" captionText="TABLE 1: Tabular view of maximum likelihood analysis of 70 species of the genus Phaselia, based on COI 5 sequences, percentage differences (built with MEGA X; Kimura 2-parameter model; bootstrap method, 1000 replications using MEGA X (Kumar et al., 2018))." pageId="23" pageNumber="24">Tab. 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>