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<document ID-DOI="http://dx.doi.org/10.3897/zse.99.103003" ID-Pensoft-Pub="1860-0743-2-423" ID-Pensoft-UUID="63B9DC36F52C5FEEA5ADD00CA079394D" ID-ZooBank="CFCE08BC6C7347EA807DEB9C3F2547E5" ModsDocID="1860-0743-99-2-423" checkinTime="1696476058948" checkinUser="pensoft" docAuthor="Segadilha, Juliana Lopes, Bird, Graham &amp; Tavares, Marcos" docDate="2023" docId="877E898DBF3D50E69551057C6EE22794" docLanguage="en" docName="ZoosystEvol 99(2): 423-437" docOrigin="Zoosystematics and Evolution 99 (2)" docPubDate="2023-10-04" docSource="http://dx.doi.org/10.3897/zse.99.103003" docTitle="Stenotanais Bird &amp; Holdich 1984" docType="treatment" docVersion="1" id="63B9DC36F52C5FEEA5ADD00CA079394D" lastPageNumber="423" masterDocId="63B9DC36F52C5FEEA5ADD00CA079394D" masterDocTitle="Stenotanais (Crustacea, Tanaidacea) from the Santos Basin: the first described species of the family Akanthophoreidae off the Brazilian coast" masterLastPageNumber="437" masterPageNumber="423" pageNumber="423" updateTime="1696476058948" updateUser="pensoft">
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<mods:title id="6CD90B4086C4BFC7E0CC5CAB9E071D82">Stenotanais (Crustacea, Tanaidacea) from the Santos Basin: the first described species of the family Akanthophoreidae off the Brazilian coast</mods:title>
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<mods:namePart id="B016F846EA3948FF155FBF0760C62BE4">Segadilha, Juliana Lopes</mods:namePart>
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<mods:affiliation id="5148669EF79E1E622C401CA21858EFF3">Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo-SP, 04263 - 000, Brazil</mods:affiliation>
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<mods:namePart id="C24D37A0E408389673FB54B9186A3F7B">Bird, Graham</mods:namePart>
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<mods:affiliation id="FABE67D4F9F76A9ACD8EE0E8BDD74A45">Waikanae, Kapiti, New Zealand</mods:affiliation>
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<mods:namePart id="D2E5E4801A562D8CED24564E5E39A758">Tavares, Marcos</mods:namePart>
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<subSubSection id="3B474016FA80617218DEBDB8128F8824" pageId="0" pageNumber="423" type="nomenclature">
<paragraph id="B47EF14B6487880E48FD72AE4773FCBB" pageId="0" pageNumber="423">
Genus
<taxonomicName id="C7CFE089C8A549943AA737C819BF48DF" ID-CoL="84TTT" LSID="877E898D-BF3D-50E6-9551-057C6EE22794" authority="Bird &amp; Holdich, 1984" authorityName="Bird &amp; Holdich" authorityYear="1984" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">Stenotanais Bird &amp; Holdich, 1984</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D25335D5BFAF12165DF2046A4ED4407E" pageId="0" pageNumber="423" type="diagnosis">
<paragraph id="E52FC4DB258D523B6FB23DF2CBDFFE3D" pageId="0" pageNumber="423">Diagnosis</paragraph>
<paragraph id="FCCD1EED227B9C96547A7DAC2BE10E2B" pageId="0" pageNumber="423">
(Modified from
<bibRefCitation id="7846FF23CC75843F5CAFD8CBC46894C8" DOI="https://doi.org/10.1163/9789047416883" author="Larsen, K" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="423" refId="B15" refString="Larsen, K, 2005. Deep-Sea Tanaidacea (Peracarida) from the Gulf of Mexico. Brill, Leiden, DOI: https://doi.org/10.1163/9789047416883" title="Deep-Sea Tanaidacea (Peracarida) from the Gulf of Mexico." url="https://doi.org/10.1163/9789047416883" year="2005">Larsen 2005</bibRefCitation>
,
<bibRefCitation id="98EC9E8625C771F55FB4143A9C1C6BE2" DOI="https://doi.org/10.11646/zootaxa.3088.1.4" author="Larsen, K" journalOrPublisher="Zootaxa" pageId="0" pageNumber="423" pagination="39 - 53" refId="B16" refString="Larsen, K, 2011. The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088 (1): 39 - 53, DOI: https://doi.org/10.11646/zootaxa.3088.1.4" title="The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea)." url="https://doi.org/10.11646/zootaxa.3088.1.4" volume="3088" year="2011">2011</bibRefCitation>
). Female. Body elongate (11.5-13
<normalizedToken id="DAC2617E126532BBD8AEB625DAB1B95C" originalValue="×">x</normalizedToken>
L:W). Pereonites longer than wide, with straight lateral edges. Pleon short (15-20% of TBL). Antennule shorter than carapace, four-articled. Antenna six-articled. Mandibular molar tapering, with terminal spines. Maxillule with eight to ten terminal spines. Maxilliped endite with rounded cusps, spiniform lateral corners and distal seta. Cheliped carpus robust with large distoventral shield; chela robust, fixed finger with two robust ventral spines. Pereopods 1-3 basis wider than on pereopods 4-6; merus and carpus with spine. Pereopods 4-6 without coxae. Pereopods 4-6 dactylus and unguis not fused; dactylus with double-row of small spines. Pleopods short and broad, with plumose or simple setae. Uropods prominent, often as long as antennae: endopod two-articled; exopod one- or two-articled; specialised setae with a wide and flat basis present on either exopod or endopod.
</paragraph>
<paragraph id="E1105A9F35B4DA7F1A4FBC7B39769E84" pageId="0" pageNumber="423">
Male. Immature (preparatory) male with antennule thicker than in female, functional mouthparts present (see remarks on genus). Sexually mature
<normalizedToken id="0DAE951904197484C95DBAD4A9BB132D" originalValue="swimming">'swimming'</normalizedToken>
type: cephalothorax as long as first three pereonites; pereon shorter than in female; pleon well developed, as long as pereon, caudally pointed; multi-articulate antennule with multiple aesthetascs, distal article longer than preceding article; no functional mouthparts; uropod biramous, endopod and exopod two-articled.
</paragraph>
</subSubSection>
<subSubSection id="A7E4E2236B4DF9E8DB611DACF4932E5D" pageId="0" pageNumber="423" type="type species">
<paragraph id="C081084CBA6D2996DD3B89F2093F4F40" pageId="0" pageNumber="423">Type species.</paragraph>
<paragraph id="F48534812F06DE93700D458C768970AF" pageId="0" pageNumber="423">
<taxonomicName id="95C5432E40D2711F1941847F90304DA5" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais crassiseta" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="crassiseta">
<emphasis id="FCE3A15A4736FC83682D412D5161C096" italics="true" pageId="0" pageNumber="423">Stenotanais crassiseta</emphasis>
</taxonomicName>
Bird &amp; Holdich, 1984, by original designation. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="42D4DDEF92524CC164AECCF81B461166" pageId="0" pageNumber="423" type="species included">
<paragraph id="8D39FFD6A348F853624ED7BC5D8B79FF" pageId="0" pageNumber="423">Species included.</paragraph>
<paragraph id="A5566038AFCB4A6D0BC591F18FCC7DD9" pageId="0" pageNumber="423">
<taxonomicName id="575B064DE84818434F5E42FB0270699D" authorityName="Larsen" authorityYear="2011" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais arenasi" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="arenasi">
<emphasis id="0774A65CD1695A5161A20592A34396A9" italics="true" pageId="0" pageNumber="423">Stenotanais arenasi</emphasis>
</taxonomicName>
Larsen, 2011;
<taxonomicName id="94163FB08E076E6F42F0506069A76A01" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. crassiseta" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="crassiseta">
<emphasis id="CAF46EF359EA657BC4B3E8143041CDE8" italics="true" pageId="0" pageNumber="423">S. crassiseta</emphasis>
</taxonomicName>
Bird &amp; Holdich, 1984;
<taxonomicName id="3BB201D557E1D7D1854049358FFA94E1" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. hamicauda" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="hamicauda">
<emphasis id="060D7533022F8790E489C5E7D7E4AA07" italics="true" pageId="0" pageNumber="423">S. hamicauda</emphasis>
</taxonomicName>
Bird &amp; Holdich, 1984;
<taxonomicName id="46C95F35863D39E300597A474698647E" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. macrodactylus" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="377C418AFB0ABBEF20BCE607D60B2D4C" italics="true" pageId="0" pageNumber="423">S. macrodactylus</emphasis>
</taxonomicName>
Larsen, 2005;
<taxonomicName id="49C6F243E34867DB64726FFCF79CDE79" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. leonardoi" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="leonardoi">
<emphasis id="DEFCA69B5A92D445F7BA0B58E734D1C7" italics="true" pageId="0" pageNumber="423">S. leonardoi</emphasis>
</taxonomicName>
sp. nov.;
<taxonomicName id="F3C4EFA2A1A9E07072F3B22D7DE62435" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. uropedon" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="uropedon">
<emphasis id="C9E8EE30DE8FAF173B8A374380145E7A" italics="true" pageId="0" pageNumber="423">S. uropedon</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="58E20B3ACFF27B4FCF8AC15036249989" pageId="0" pageNumber="423" type="remarks">
<paragraph id="A675DD617C3138A8F27874FE4DE58049" pageId="0" pageNumber="423">Remarks.</paragraph>
<paragraph id="8AFA2F73E3BCFBB3709494D52F24A1A2" pageId="0" pageNumber="423">
The rather infrequently-recorded
<taxonomicName id="3E7050C8874288C36F3843BED938575C" authorityName="Bird &amp; Holdich" authorityYear="1984" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="6F01C22D0828542AFAD429E6878CDBD6" italics="true" pageId="0" pageNumber="423">Stenotanais</emphasis>
</taxonomicName>
is similar to akanthophoreid genera such as
<taxonomicName id="F8601864CE66160709F946A59E7ACDA6" authorityName="Lubinevsky, Tom &amp; Bird" authorityYear="2022" class="Malacostraca" family="Akanthophoreidae" genus="Pseudakanthophoreus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudakanthophoreus" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="5C228095D9CBEC18F5608DA3A4516E5F" italics="true" pageId="0" pageNumber="423">Pseudakanthophoreus</emphasis>
</taxonomicName>
Lubinevsky, Tom &amp; Bird, 2022 and
<taxonomicName id="3C7FA71D3442DEE2B7B6A38883AA25D1" authorityName="Larsen &amp; Araujo-Silva" authorityYear="2014" class="Malacostraca" family="Akanthophoreidae" genus="Parakanthophoreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parakanthophoreus" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="67E307AE9EFE8754D00082B0C8932F40" italics="true" pageId="0" pageNumber="423">Parakanthophoreus</emphasis>
</taxonomicName>
Larsen &amp;
<normalizedToken id="1D371E22061154F0FCDC6ADC006FB1F3" originalValue="Araújo-Silva">Araujo-Silva</normalizedToken>
, 2014, by having chelipeds without extensive surface ornamentation and the absence of spurs on pleonite-5, pleotelson and the uropod basal article or endopod article-1. However,
<taxonomicName id="6BD75E7CB0FC4E71A7463102396433FE" authorityName="Bird &amp; Holdich" authorityYear="1984" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="37A66F2279710994DFA346F0B9DBDA60" italics="true" pageId="0" pageNumber="423">Stenotanais</emphasis>
</taxonomicName>
can be distinguished mainly by having the basis of pereopods 1-3 thicker than the basis of pereopods 4-6 and the presence of specialised setae with a wide and flat basal attachment on the uropod (
<bibRefCitation id="8DEB097E8C2B8BD9279F0F1A9322C6C5" DOI="https://doi.org/10.1111/j.1463-6409.1984.tb00044.x" author="Bird, GJ" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="423" pagination="285 - 315" refId="B4" refString="Bird, GJ, Holdich, DM, 1984. New Deep-sea Leptognathiid Tanaids (Crustacea, Tanaidacea) from the North-east Atlantic. Zoologica Scripta 13 (4): 285 - 315, DOI: https://doi.org/10.1111/j.1463-6409.1984.tb00044.x" title="New Deep-sea Leptognathiid Tanaids (Crustacea, Tanaidacea) from the North-east Atlantic." url="https://doi.org/10.1111/j.1463-6409.1984.tb00044.x" volume="13" year="1984">Bird and Holdich 1984</bibRefCitation>
;
<bibRefCitation id="9B8A8371295BCC18DAB541344EEE2E47" DOI="https://doi.org/10.11646/zootaxa.3088.1.4" author="Larsen, K" journalOrPublisher="Zootaxa" pageId="0" pageNumber="423" pagination="39 - 53" refId="B16" refString="Larsen, K, 2011. The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088 (1): 39 - 53, DOI: https://doi.org/10.11646/zootaxa.3088.1.4" title="The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea)." url="https://doi.org/10.11646/zootaxa.3088.1.4" volume="3088" year="2011">Larsen 2011</bibRefCitation>
).
</paragraph>
<paragraph id="41B2CB9D78FA2714691A3B8E08D5389A" pageId="0" pageNumber="423">
<bibRefCitation id="2233D30F5B87B01FB12B0EC4370EFBAB" DOI="https://doi.org/10.11646/zootaxa.3088.1.4" author="Larsen, K" journalOrPublisher="Zootaxa" pageId="0" pageNumber="423" pagination="39 - 53" refId="B16" refString="Larsen, K, 2011. The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088 (1): 39 - 53, DOI: https://doi.org/10.11646/zootaxa.3088.1.4" title="The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea)." url="https://doi.org/10.11646/zootaxa.3088.1.4" volume="3088" year="2011">Larsen (2011)</bibRefCitation>
described
<taxonomicName id="C2B9483CF92DEA3902A3D3108A2CCBB6" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. arenasi" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="arenasi">
<emphasis id="1AF8566FC3C1E4FEF35222FCDC3136AA" italics="true" pageId="0" pageNumber="423">S. arenasi</emphasis>
</taxonomicName>
, based on a single specimen and expressed uncertainty as to whether it really belongs to
<taxonomicName id="AFB5D8D45517D7842BD2E06111B268F6" authorityName="Bird &amp; Holdich" authorityYear="1984" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="13E5A36D065000ED1A40431EB304D974" italics="true" pageId="0" pageNumber="423">Stenotanais</emphasis>
</taxonomicName>
, so that its generic affiliation should be considered tentative. We agree that this species may belong to a different genus as its cheliped lacks a strong carpal shield, the shape of the antennule articles 3-4 does not conform to other
<taxonomicName id="CADA14361C64BB6FFB0C09A7FA57B440" authorityName="Bird &amp; Holdich" authorityYear="1984" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="2A8045A968CA8375EC35887334500525" italics="true" pageId="0" pageNumber="423">Stenotanais</emphasis>
</taxonomicName>
species (article-3 not rectangular and article-4 minute, as long as wide; which can be a sexually dimorphic character) and the uropods are not ventrally deflexed (i.e. not folded back on themselves). Although not an impediment for supporting the exclusion of
<taxonomicName id="5ECF5504EBF0DBDF5C961F2C45B4A459" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. arenasi" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="arenasi">
<emphasis id="5E2DA59759266DD29139C29144571AEE" italics="true" pageId="0" pageNumber="423">S. arenasi</emphasis>
</taxonomicName>
from
<taxonomicName id="5FDDB72C51FF9932A28460F49C258D28" authorityName="Bird &amp; Holdich" authorityYear="1984" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="EA0EC37621D10C9358D2947CF57FF853" italics="true" pageId="0" pageNumber="423">Stenotanais</emphasis>
</taxonomicName>
, it is of note that it is so far the only species of the genus reported from the Pacific Ocean (Fig.
<figureCitation id="B1ABA00CA5EA4D88C33D544476F4F26F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic distribution of Stenotanais species based on the present study and literature data (Bird and Holdich 1984; Schmidt and Brandt 2001; Larsen 2005, 2011)." figureDoi="10.3897/zse.99.103003.figure8" httpUri="https://binary.pensoft.net/fig/916462" pageId="0" pageNumber="423">8</figureCitation>
). Despite these considerations, we adhere to the current classification until more evidence is available.
</paragraph>
<paragraph id="0D8D7B78243D8AF703043D4B7585A137" pageId="0" pageNumber="423">
The single available specimen of
<taxonomicName id="EB658EFD71A33CE02B035F4D578E88BB" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. arenasi" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="arenasi">
<emphasis id="E6EC3D73D5D73AF68A0E2A7F6F1D36B3" italics="true" pageId="0" pageNumber="423">S. arenasi</emphasis>
</taxonomicName>
may be a preparatory male, based on characters of the antennule articles 1-2 (as suggested by
<bibRefCitation id="9D0844F2620CCF2070767A53D73A8CCF" DOI="https://doi.org/10.11646/zootaxa.3088.1.4" author="Larsen, K" journalOrPublisher="Zootaxa" pageId="0" pageNumber="423" pagination="39 - 53" refId="B16" refString="Larsen, K, 2011. The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088 (1): 39 - 53, DOI: https://doi.org/10.11646/zootaxa.3088.1.4" title="The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea)." url="https://doi.org/10.11646/zootaxa.3088.1.4" volume="3088" year="2011">Larsen 2011</bibRefCitation>
) and has functional mouthparts. Here, we record the first sexually mature
<normalizedToken id="E7C52393CD26516C0E142FE0E0E5F9AB" originalValue="swimming">'swimming'</normalizedToken>
male of
<taxonomicName id="6B27C3E04A24251E30C03CCF26EE1844" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. leonardoi" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="leonardoi">
<emphasis id="2348C81F2F3C90BF7DE086882E05BAB2" italics="true" pageId="0" pageNumber="423">S. leonardoi</emphasis>
</taxonomicName>
sp. nov. (see below), the first to have been noticed in
<taxonomicName id="589C31FD3E9BC8A7D58C768F034B66CF" authorityName="Bird &amp; Holdich" authorityYear="1984" class="Malacostraca" family="Akanthophoreidae" genus="Stenotanais" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenotanais" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="71442232944E2AD228E701E65C114C05" italics="true" pageId="0" pageNumber="423">Stenotanais</emphasis>
</taxonomicName>
, although it would have been desirable to confirm the link between male and female forms of the same species using molecular analysis.
</paragraph>
<paragraph id="814BF0B7E70F51CB4D587B8FE87630AB" pageId="0" pageNumber="423">
Another character in the original generic diagnosis of
<bibRefCitation id="64B2D0CD6ADBFFCBDB22BA33BDD7E58A" DOI="https://doi.org/10.1111/j.1463-6409.1984.tb00044.x" author="Bird, GJ" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="423" pagination="285 - 315" refId="B4" refString="Bird, GJ, Holdich, DM, 1984. New Deep-sea Leptognathiid Tanaids (Crustacea, Tanaidacea) from the North-east Atlantic. Zoologica Scripta 13 (4): 285 - 315, DOI: https://doi.org/10.1111/j.1463-6409.1984.tb00044.x" title="New Deep-sea Leptognathiid Tanaids (Crustacea, Tanaidacea) from the North-east Atlantic." url="https://doi.org/10.1111/j.1463-6409.1984.tb00044.x" volume="13" year="1984">Bird and Holdich (1984)</bibRefCitation>
is the pereopods 1-3 propodus having a distinct convex inferior margin armed with small strong spinules. This needs further investigation as it is true for
<taxonomicName id="35E97F1A576D1C3FCC2CD9F44F3337AE" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. crassiseta" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="crassiseta">
<emphasis id="2A3C4B30A6DDC8DEC12B5BB6721DF7D1" italics="true" pageId="0" pageNumber="423">S. crassiseta</emphasis>
</taxonomicName>
,
<taxonomicName id="D8F870135C4A7DF80832F2E15356CC1C" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. hamicauda" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="hamicauda">
<emphasis id="8DCEB1F8E1B4DDB04E7A979BE79D084B" italics="true" pageId="0" pageNumber="423">S. hamicauda</emphasis>
</taxonomicName>
and both new species described here, but not for
<taxonomicName id="37D44CA7686B7B82013CA58D1285EA11" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. arenasi" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="arenasi">
<emphasis id="90D8F79CA4BD1C84400D65B81157DCD6" italics="true" pageId="0" pageNumber="423">S. arenasi</emphasis>
</taxonomicName>
and
<taxonomicName id="3D80F868BFDE3A0C221CCEB579F0FF7D" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. macrodactylus" order="Tanaidacea" pageId="0" pageNumber="423" phylum="Arthropoda" rank="species" species="macrodactylus">
<emphasis id="76461E2CEFDD0BC00DF3F9464EBE43C5" italics="true" pageId="0" pageNumber="423">S. macrodactylus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>