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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.912.47719" ID-GBIF-Dataset="26396469-500e-42b8-acba-f6a22f652923" ID-PMC="PMC7040049" ID-Pensoft-Pub="1313-2970-912-65" ID-Pensoft-UUID="D1FDF644344C5B5B83E6024904E40A05" ID-PubMed="32123500" ID-ZooBank="4F361E38B2684B70944DD6B6FCC66542" ModsDocID="1313-2970-912-65" checkinTime="1581958640118" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2020" docId="804F0C62E90D5A52857B2D8019323DBC" docLanguage="en" docName="ZooKeys 912: 65-124" docOrigin="ZooKeys 912" docSource="http://dx.doi.org/10.3897/zookeys.912.47719" docTitle="Theopea bicolor Kimoto 1989" docType="treatment" docVersion="4" id="D1FDF644344C5B5B83E6024904E40A05" lastPageNumber="65" masterDocId="D1FDF644344C5B5B83E6024904E40A05" masterDocTitle="Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species" masterLastPageNumber="124" masterPageNumber="65" pageNumber="65" updateTime="1668168482312" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>912</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>65</mods:start>
<mods:end>124</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.912.47719</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.912.47719</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-912-65</mods:identifier>
<mods:identifier type="ZooBank">4F361E38B2684B70944DD6B6FCC66542</mods:identifier>
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<treatment ID-GBIF-Taxon="162147241" LSID="urn:lsid:plazi:treatment:804F0C62E90D5A52857B2D8019323DBC" httpUri="http://treatment.plazi.org/id/804F0C62E90D5A52857B2D8019323DBC" lastPageNumber="65" pageId="0" pageNumber="65">
<subSubSection pageId="0" pageNumber="65" type="nomenclature">
<paragraph pageId="0" pageNumber="65">
<taxonomicName LSID="804F0C62-E90D-5A52-857B-2D8019323DBC" authority="Kimoto, 1989" authorityName="Kimoto" authorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicolor" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicolor">Theopea bicolor Kimoto, 1989</taxonomicName>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Theopea bicolor, T. bicoloroides sp. nov., and T. mouhoti. A T. bicolor, male, dorsal view B Same, ventral view C T. bicolor, female, dorsal view D T. bicoloroides sp. nov., male, dorsal view E Same, ventral view F T. bicoloroides sp. nov., female, dorsal view G T. mouhoti, male H Same, ventral view I T. mouhoti, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65">Figs 1A-C</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="reference_group">
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Kimoto" authorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicolor" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicolor">Theopea bicolor</taxonomicName>
Kimoto, 1989: 199 (Vietnam); Mohamedsaid &amp; Costant, 2007 (Thailand).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="materials_examined">
<paragraph pageId="0" pageNumber="65">Type.</paragraph>
<paragraph pageId="0" pageNumber="65">Holotype ♂ (BPBM): &quot;VIET NAM. 20 km / N. of Pleiku / 650m. 9.V.1960 [p, w] // L. W. Quate / Collector [p, w] // Theopea / bicolor / n. sp. [h, w] // HOLOTYPE [p, r]&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="materials_examined">
<paragraph pageId="0" pageNumber="65">Other material.</paragraph>
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">THAILAND</emphasis>
. Chiang Mai: 3♂♂, 2♀♀ (SEHU), Chiang Dao Valley, 2.V.1980, leg. Y. Komiya; 5♀♀ (SEHU), same locality, 24.V.1983, leg. Y. Komiya; 2♀♀ (SEHU), same but with &quot;leg. H. Akiyama; 3♀♀ (SEHU), same locality, 30.V.1983, leg. Y. Komiya; 1♀ (SEHU), same but with &quot;leg. K. Ikeda&quot;; 1♀ (SEHU), same but with &quot;H. Akiyama&quot;; 4♀♀ (NHMB), same locality, 10-16.V.1991, leg. V.
<normalizedToken originalValue="Kubáň">Kuban</normalizedToken>
; 1♂, 7♀♀ (1♂, 6♀♀: NHMB; 1♀: MSNG), same locality, 17-24.V.1991, leg. V.
<normalizedToken originalValue="Kubáň">Kuban</normalizedToken>
; 4♂♂ (SEHU), Doi Pui, 28.IV.-1.V.1980, leg. Y. Komiya; 1♀ (NMPC), Doi Suthep, 19-22.-IV.1991, leg. S.
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
; 1♂ (JBCB), Doi Suthep to Doi Pui,
<geoCoordinate degrees="18" direction="north" minutes="49" orientation="latitude" precision="925" value="18.816668">18°49N</geoCoordinate>
<geoCoordinate degrees="99" direction="east" minutes="00" orientation="longitude" precision="925" value="99.0">99°00E</geoCoordinate>
, 19.-23.IV.1991, leg. L.
<normalizedToken originalValue="Dembický">Dembicky</normalizedToken>
; 1♂ (MSNG), Palong,
<geoCoordinate degrees="19" direction="north" minutes="55" orientation="latitude" precision="925" value="19.916666">19°55'N</geoCoordinate>
<geoCoordinate degrees="99" direction="east" minutes="06" orientation="longitude" precision="925" value="99.1">99°06'E</geoCoordinate>
, 750 m, 26-28.V.1991, leg. V.
<normalizedToken originalValue="Kubáň">Kuban</normalizedToken>
; Kanchanaburi: 1♂, 4♀♀ (SEHU), Ban Nong Bang, 15.V.1985, leg. Y. Komiya; Mae Hong Son: 6♂♂ (JBCB), Ban Huai Po,
<geoCoordinate degrees="19" direction="north" minutes="19" orientation="latitude" precision="925" value="19.316668">19°19N</geoCoordinate>
<geoCoordinate degrees="97" direction="east" minutes="59" orientation="longitude" precision="925" value="97.98333">97°59E</geoCoordinate>
, 1600-2000 m, 9.-16.V.1991, leg. L.
<normalizedToken originalValue="Dembický">Dembicky</normalizedToken>
; 1♀ (NHMUK), same locality, 9-16.V.1992, leg. J.
<normalizedToken originalValue="Horák">Horak</normalizedToken>
; 33♂♂, 2♀♀ (24♂♂, 2♀♀: NHMUK; 9♂♂: JBCB), Ban Si Lang, 1200 m, 1-8.V.1992, leg. J.
<normalizedToken originalValue="Horák">Horak</normalizedToken>
; 1♀ (JBCB), Kiwlom-pass near Soppong,
<geoCoordinate degrees="19" direction="north" minutes="26" orientation="latitude" precision="925" value="19.433332">19°26N</geoCoordinate>
<geoCoordinate degrees="98" direction="east" minutes="19" orientation="longitude" precision="925" value="98.316666">98°19E</geoCoordinate>
, 1400 m, 23.VI.-2.VII.2002, leg. R. and H.
<normalizedToken originalValue="Fouqué">Fouque</normalizedToken>
; 1♀ (JBCB), SE of Soppong,
<geoCoordinate degrees="19" direction="north" minutes="27" orientation="latitude" precision="925" value="19.45">19°27N</geoCoordinate>
<geoCoordinate degrees="98" direction="east" minutes="20" orientation="longitude" precision="925" value="98.333336">98°20E</geoCoordinate>
, 1500 m, 23.-27.V.1999, leg. M.
<normalizedToken originalValue="Řiha">Řiha</normalizedToken>
; 3♂♂, 1♀ (NHMB), Soppong-Pai, 1800 m, leg.
<normalizedToken originalValue="Pacholátko">Pacholatko</normalizedToken>
; Nan: 1♂ (JBCB), Ban Huay Kon env., 27.V.-10.VI.2002, leg. P.
<normalizedToken originalValue="Průdek">Prudek</normalizedToken>
, M.
<normalizedToken originalValue="Obořil">Oboril</normalizedToken>
; 1♂ (NHMUK), Doi Phuka N.P., V.2000, leg. local collector; 1♀ (SEHU), Mae Kamme Forest, 17.V.1985, leg. Y. Komiya; 1♂ (SEHU), Nan Watershed Res. Station, 17.V.1985, leg. Y. Komiya; 1♂ (SEHU), Wiang Sa, 15.V.1993, leg. S. Ohmomo; 2♀♀ (SEHU), Wieng Ko Sai N.P., 18.V.1985, leg. Y. Komiya; Prachinburi: 1♀ (HNHM), Sakaerat Ecol. Research Institute, 4.VI.2001, leg. E.
<normalizedToken originalValue="Harváth">Harvath</normalizedToken>
and G.
<normalizedToken originalValue="Szirákl">Szirakl</normalizedToken>
; VIETNAM. Daklak: 1♂ (MSNG), 12 km SW of Buon Ma Thout, Lake Eakao, 400 m, 26-27.IV.1986, leg. L. Medvedev.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="redescription">
<paragraph pageId="0" pageNumber="65">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="65">
Length 5.8-6.2 mm, width 1.9-2.2 mm. Body color (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Theopea bicolor, T. bicoloroides sp. nov., and T. mouhoti. A T. bicolor, male, dorsal view B Same, ventral view C T. bicolor, female, dorsal view D T. bicoloroides sp. nov., male, dorsal view E Same, ventral view F T. bicoloroides sp. nov., female, dorsal view G T. mouhoti, male H Same, ventral view I T. mouhoti, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65">1A-C</figureCitation>
) dark brown or blackish brown except elytra reddish brown. Antennae filiform in males, but antennomeres VI-VIII slightly swollen (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2A</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.3: 0.8: 1.1: 1.2: 1.2: 1.2: 1.1: 1.1: 1.0: 1.2, length to width ratios of antennomeres I-XI 2.8: 1.2: 2.5: 3.3: 3.6: 3.4: 3.4: 3.2: 3.5: 3.4: 4.1; filiform in females (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2B</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.3: 0.8: 1.0: 1.0: 0.9: 0.9: 0.9: 0.9: 0.8: 0.9, length to width ratios of antennomeres I-XI 3.0: 1.4: 3.1: 3.5: 3.5: 3.2: 3.4: 3.4: 3.6: 3.4: 3.7. Elytra elongate, parallel-sided, 2.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one weak longitudinal ridge between two longitudinal rows of punctures, basally abbreviated. Tarsomeres I of front legs slightly swollen in males; subparallel in females. Aedeagus (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2C-E</figureCitation>
) slender, 6.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; sides widest at middle, gradually narrowed towards basal 1/4, gradually and apically narrowed towards apical 1/5, parallel between apical 1/5 and 1/12, apex with shallow notch; tectum well sclerotized, basally broadened, as broad as aedeagus, with hollow area behind base of tectum; moderately curved in lateral view; ventral surface with deep notch from near apex, apically extending into basal opening, more approximate at apical 1/5; triangular sclerites small; internal sac with one median, elongate sclerite, 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, apically tapering from basal 1/3, apex acute, connected with short broad sclerite at base, disc with dense transverse rows of hair-like setae and with one pair of elongate, longitudinal rows of stout setae at sides. Gonocoxae (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2G</figureCitation>
) elongate, widest at apical 1/6, both gonocoxae joined from basal 1/8 to apical 1/7; apices narrowly rounded, each gonocoxa with eight setae along lateral margin from apex to apical 1/6; with one pair of short lateral processes at basal 2/5. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2F</figureCitation>
) elongate and well sclerotized; disc with several long setae at sides and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2H</figureCitation>
) strongly swollen; pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short.
</paragraph>
<caption doi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 1.</emphasis>
Habitus of
<taxonomicName authorityName="Kimoto" authorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicolor" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea bicolor</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName lsidName="T. mouhoti" pageId="0" pageNumber="65" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">T. mouhoti</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
<taxonomicName lsidName="T. bicolor" pageId="0" pageNumber="65" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicolor</emphasis>
</taxonomicName>
, male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
<taxonomicName lsidName="T. bicolor" pageId="0" pageNumber="65" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicolor</emphasis>
</taxonomicName>
, female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov., male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov., female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
<taxonomicName lsidName="T. mouhoti" pageId="0" pageNumber="65" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">T. mouhoti</emphasis>
</taxonomicName>
, male
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">I</emphasis>
<taxonomicName lsidName="T. mouhoti" pageId="0" pageNumber="65" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">T. mouhoti</emphasis>
</taxonomicName>
, female, dorsal view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 2.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Kimoto" authorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicolor" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea bicolor</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
Antenna, male
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Antenna, female
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
Aedeagus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Aedeagus, lateral view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
Aedeagus, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
Abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
Gonocoxae
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
Spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="remarks">
<paragraph pageId="0" pageNumber="65">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="65">
Populations from Laos and southwest China were misidentified. They represent
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicoloroides" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea bicoloroides</emphasis>
</taxonomicName>
sp. nov. (see below).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="diagnosis">
<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Kimoto" authorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicolor" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea bicolor</emphasis>
</taxonomicName>
Kimoto,
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName lsidName="T. mouhoti" pageId="0" pageNumber="65" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">T. mouhoti</emphasis>
</taxonomicName>
Baly are characterized by their reddish brown elytra.
<taxonomicName authorityName="Kimoto" authorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicolor" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea bicolor</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Theopea bicolor, T. bicoloroides sp. nov., and T. mouhoti. A T. bicolor, male, dorsal view B Same, ventral view C T. bicolor, female, dorsal view D T. bicoloroides sp. nov., male, dorsal view E Same, ventral view F T. bicoloroides sp. nov., female, dorsal view G T. mouhoti, male H Same, ventral view I T. mouhoti, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65">1A-F</figureCitation>
) can be easily separately from
<taxonomicName lsidName="T. mouhoti" pageId="0" pageNumber="65" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">T. mouhoti</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Theopea bicolor, T. bicoloroides sp. nov., and T. mouhoti. A T. bicolor, male, dorsal view B Same, ventral view C T. bicolor, female, dorsal view D T. bicoloroides sp. nov., male, dorsal view E Same, ventral view F T. bicoloroides sp. nov., female, dorsal view G T. mouhoti, male H Same, ventral view I T. mouhoti, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure1" httpUri="https://binary.pensoft.net/fig/380656" pageId="0" pageNumber="65">1G-I</figureCitation>
) by the dark brown or blackish brown head, prothorax, and scutellum, and indistinct ridges in the elytra. Besides, males of
<taxonomicName lsidName="T. bicolor" pageId="0" pageNumber="65" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicolor</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov. possess a median elongate sclerite internally in the aedeagus that is covered with transverse rows of hair-like setae (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2C, D</figureCitation>
;
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic characters of Theopea bicoloroides sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure3" httpUri="https://binary.pensoft.net/fig/380658" pageId="0" pageNumber="65">3C, D</figureCitation>
). This differs from those of
<taxonomicName lsidName="T. mouhoti" pageId="0" pageNumber="65" rank="species" species="mouhoti">
<emphasis italics="true" pageId="0" pageNumber="65">T. mouhoti</emphasis>
</taxonomicName>
, which lacks such hair-like setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Theopea mouhoti. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Same, lateral view E Same, ventral view F Aedeagus, variation, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure4" httpUri="https://binary.pensoft.net/fig/380659" pageId="0" pageNumber="65">4C, D</figureCitation>
).
<taxonomicName authorityName="Kimoto" authorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea bicolor" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea bicolor</emphasis>
</taxonomicName>
differs from
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov. by the relatively slender antennae (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2A</figureCitation>
) in males (length to width ratios of antennomeres V-X more than 3.0 in
<taxonomicName lsidName="T. bicolor" pageId="0" pageNumber="65" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicolor</emphasis>
</taxonomicName>
, relatively broader antenna (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic characters of Theopea bicoloroides sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure3" httpUri="https://binary.pensoft.net/fig/380658" pageId="0" pageNumber="65">3A</figureCitation>
), less than 3.0 in
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov.), the narrowly rounded apex of the ventral surface of the aedeagus (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2E</figureCitation>
) (broadly rounded apex of aedeagus in
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic characters of Theopea bicoloroides sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure3" httpUri="https://binary.pensoft.net/fig/380658" pageId="0" pageNumber="65">3E</figureCitation>
)), endophallic sclerite broad and without longitudinal groove in lateral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2D</figureCitation>
) (dorso-ventrally flattened and with longitudinal groove in lateral view in
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic characters of Theopea bicoloroides sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure3" httpUri="https://binary.pensoft.net/fig/380658" pageId="0" pageNumber="65">3D</figureCitation>
)), and slender notch at apex of gonocoxae (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Theopea bicolor. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure2" httpUri="https://binary.pensoft.net/fig/380657" pageId="0" pageNumber="65">2G</figureCitation>
) (broad notch at apex of gonocoxae in
<taxonomicName lsidName="T. bicoloroides" pageId="0" pageNumber="65" rank="species" species="bicoloroides">
<emphasis italics="true" pageId="0" pageNumber="65">T. bicoloroides</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic characters of Theopea bicoloroides sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure3" httpUri="https://binary.pensoft.net/fig/380658" pageId="0" pageNumber="65">3G</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="distribution">
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="65">Thailand, Vietnam.</paragraph>
</subSubSection>
</treatment>
</document>