233 lines
33 KiB
XML
233 lines
33 KiB
XML
<document id="AEBA8E469170C5162F5105640F79E8AA" ID-DOI="10.5281/zenodo.184416" ID-GBIF-Dataset="bf3414c0-6326-4ff1-9670-55460d7b85b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="184416" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459981288142" checkinUser="plazi" docAuthor="Escalona, Hermes E. & Ślipiński, Adam" docDate="2008" docId="7149B174F67E4A5B2485FA59D7E00B85" docLanguage="en" docName="zt01895p065.pdf" docOrigin="Zootaxa 1895" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ranomafana pollocki Escalona & Ślipiński, 2008, sp. n." docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="8D70C90CF67C4A5D2412FF99D3330F1F" masterDocTitle="A new species of Ranomafana from Madagascar (Coleoptera: Pyrochroidae: Pilipalpinae)" masterLastPageNumber="65" masterPageNumber="59" pageNumber="61" updateTime="1698231831888" updateUser="plazi">
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<mods:titleInfo id="3E93CFD5FCE919A44EDDB332A48812F6">
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<mods:title id="A5CA7C742DA28486493A77AA1FEEC009">A new species of Ranomafana from Madagascar (Coleoptera: Pyrochroidae: Pilipalpinae)</mods:title>
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<mods:name id="03943254EB9880D12F5F636ED3FAA938" type="personal">
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<mods:namePart id="8329BBC2C5EE9E6E51E87BD8894599B9">Escalona, Hermes E.</mods:namePart>
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<mods:name id="7F2BFF7AC096BF9EAE316CF3E9035E60" type="personal">
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<mods:namePart id="F843AC599593D69C27B761D111D6B893">Ślipiński, Adam</mods:namePart>
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<mods:typeOfResource id="19DAE63701A86218F6B318D1359C0E60">text</mods:typeOfResource>
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<mods:title id="5EB8B0A4D5FB1C541442EC8AA710BDDB">Zootaxa</mods:title>
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<mods:part id="199EBFDD611F952CCF1E6105E26F7C46">
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<mods:date id="E5FF05AA8632F3869DEDE7F2C753AD62">2008</mods:date>
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<mods:identifier id="A72941C5F3E7E6C581951CF755580F15" type="DOI">10.5281/zenodo.184416</mods:identifier>
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<treatment id="7149B174F67E4A5B2485FA59D7E00B85" ID-DOI="http://doi.org/10.5281/zenodo.6231182" ID-GBIF-Taxon="119378605" ID-Zenodo-Dep="6231182" LSID="urn:lsid:plazi:treatment:7149B174F67E4A5B2485FA59D7E00B85" httpUri="http://treatment.plazi.org/id/7149B174F67E4A5B2485FA59D7E00B85" lastPageId="6" lastPageNumber="65" pageId="2" pageNumber="61">
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<subSubSection id="B1FA53E9F67E4A5F2485FA59D2150AE3" pageId="2" pageNumber="61" type="nomenclature">
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<paragraph id="F95F0062F67E4A5F2485FA59D2D60AC5" blockId="2.[151,485,1472,1532]" box="[151,485,1472,1498]" pageId="2" pageNumber="61">
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<heading id="A217B70EF67E4A5F2485FA59D2D60AC5" bold="true" box="[151,485,1472,1498]" fontSize="11" level="1" pageId="2" pageNumber="61" reason="1">
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<emphasis id="CB94DC70F67E4A5F2485FA59D2D60AC5" bold="true" box="[151,485,1472,1498]" pageId="2" pageNumber="61">
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<taxonomicName id="3EE07BE1F67E4A5F2485FA59D2AF0AC5" box="[151,412,1472,1498]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pollocki" status="sp. nov.">
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<emphasis id="CB94DC70F67E4A5F2485FA59D2AF0AC5" bold="true" box="[151,412,1472,1498]" italics="true" pageId="2" pageNumber="61">Ranomafana pollocki</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="D0A7610BF67E4A5F25B1FA59D2D60AC5" box="[419,485,1472,1498]" pageId="2" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="F95F0062F67E4A5F2485FA7BD2150AE3" blockId="2.[151,485,1472,1532]" box="[151,294,1506,1532]" pageId="2" pageNumber="61">(Figs. 3–20)</paragraph>
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</subSubSection>
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||
<subSubSection id="B1FA53E9F67E4A5F2485F9ABD1DA09DB" pageId="2" pageNumber="61" type="materials_examined">
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||
<paragraph id="F95F0062F67E4A5F2485F9ABD17C096B" blockId="2.[151,1436,1586,2012]" pageId="2" pageNumber="61">
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<emphasis id="CB94DC70F67E4A5F2485F9ABD2210953" bold="true" box="[151,274,1586,1612]" pageId="2" pageNumber="61">
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<typeStatus id="265BBEC0F67E4A5F2485F9ABD23D0953" box="[151,270,1586,1612]" pageId="2" pageNumber="61" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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%,
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<collectingCountry id="81F740F2F67E4A5F2552F9ABD2E30953" box="[320,464,1586,1612]" name="Madagascar" pageId="2" pageNumber="61">Madagascar</collectingCountry>
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, Tamatave prov., Andasibe (Maromizaha),
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<date id="8D5E26A2F67E4A5F27C6F9ABD7BC0953" box="[980,1167,1586,1612]" pageId="2" pageNumber="61" value="1995-11-21" valueMax="1995-11-24" valueMin="1995-11-21">21–24–xi–1995</date>
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, Ivo Jenis leg. /
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<typeStatus id="265BBEC0F67E4A5F2145F9ABD3FF096B" pageId="2" pageNumber="61" type="holotype">Holotype</typeStatus>
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,
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<taxonomicName id="3EE07BE1F67E4A5F24CAF9C2D2E7096B" box="[216,468,1627,1652]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pollocki">
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<emphasis id="CB94DC70F67E4A5F24CAF9C2D2E7096B" box="[216,468,1627,1652]" italics="true" pageId="2" pageNumber="61">Ranomafana pollocki</emphasis>
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</taxonomicName>
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(
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<collectionCode id="9FF198A7F67E4A5F25F1F9C3D177096B" box="[483,580,1626,1652]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="61">MHNG</collectionCode>
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).
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</paragraph>
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<paragraph id="F95F0062F67E4A5F24D4F91BD1DA09DB" blockId="2.[151,1436,1586,2012]" pageId="2" pageNumber="61">
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<typeStatus id="265BBEC0F67E4A5F24D4F91BD20E0983" box="[198,317,1666,1692]" pageId="2" pageNumber="61" type="paratype">Paratypes</typeStatus>
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: 15 (9 %%, 6 ΨΨ): same data as
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<typeStatus id="265BBEC0F67E4A5F26A7F91BD02D0983" box="[693,798,1666,1692]" pageId="2" pageNumber="61" type="holotype">holotype</typeStatus>
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, 2 %%, 1 Ψ (1 % disarticulated on slide),
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<collectionCode id="9FF198A7F67E4A5F20EFF91BD67B0983" LSID="urn:lsid:biocol.org:col:32981" box="[1277,1352,1666,1692]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="2" pageNumber="61">ANIC</collectionCode>
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; 1 %, 1 Ψ,
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<collectionCode id="9FF198A7F67E4A5F24ABF933D23809DB" box="[185,267,1706,1732]" pageId="2" pageNumber="61">DAPC</collectionCode>
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; 5 %%, 3 ΨΨ,
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<collectionCode id="9FF198A7F67E4A5F25BCF933D13809DB" box="[430,523,1706,1732]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="61">MHNG</collectionCode>
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; 1 %, 1 Ψ
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<collectionCode id="9FF198A7F67E4A5F2695F933D1D609DB" box="[647,741,1706,1732]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="61">NMNH</collectionCode>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B1FA53E9F67E4A5F24D4F94BD199080B" pageId="2" pageNumber="61" type="etymology">
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<paragraph id="F95F0062F67E4A5F24D4F94BD199080B" blockId="2.[151,1436,1586,2012]" pageId="2" pageNumber="61">
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<emphasis id="CB94DC70F67E4A5F24D4F94BD26409F3" bold="true" box="[198,343,1746,1772]" pageId="2" pageNumber="61">Etymology:</emphasis>
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Dedicated to Darren A. Pollock, renowned coleopterist and friend, for his substantial contributions to the systematics of Tenebrionoidea.
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</paragraph>
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</subSubSection>
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<subSubSection id="B1FA53E9F67E4A5F24D4F8BBD6B608AB" pageId="2" pageNumber="61" type="diagnosis">
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<paragraph id="F95F0062F67E4A5F24D4F8BBD6B608AB" blockId="2.[151,1436,1586,2012]" pageId="2" pageNumber="61">
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<emphasis id="CB94DC70F67E4A5F24D4F8BBD27B0823" bold="true" box="[198,328,1826,1852]" pageId="2" pageNumber="61">Diagnosis:</emphasis>
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<taxonomicName id="3EE07BE1F67E4A5F2541F8BAD1630823" box="[339,592,1827,1852]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pollocki">
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<emphasis id="CB94DC70F67E4A5F2541F8BAD1630823" box="[339,592,1827,1852]" italics="true" pageId="2" pageNumber="61">Ranomafana pollocki</emphasis>
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</taxonomicName>
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(Fig. 3a) readily differs from the uniformly rufotestaceous
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<taxonomicName id="3EE07BE1F67E4A5F2133F8BAD6AF0823" box="[1313,1436,1827,1852]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="steineri">
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<emphasis id="CB94DC70F67E4A5F2133F8BAD6AF0823" box="[1313,1436,1827,1852]" italics="true" pageId="2" pageNumber="61">R. steineri</emphasis>
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</taxonomicName>
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(Fig. 1) in having a bicoloured body, with antennomeres 5–11, most of head, scutellum and central part of pronotum black. Additionally,
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<taxonomicName id="3EE07BE1F67E4A5F25EDF8EAD14D0893" box="[511,638,1907,1932]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pollocki">
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<emphasis id="CB94DC70F67E4A5F25EDF8EAD14D0893" box="[511,638,1907,1932]" italics="true" pageId="2" pageNumber="61">R. pollocki</emphasis>
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</taxonomicName>
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possesses a distinctly transverse pronotum with six lateral tubercles (Fig. 2 vs.
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<figureCitation id="61DB1CE7F67E4A5F2507F803D26908AB" box="[277,346,1946,1972]" captionStart="FIGURES 9 – 11" captionStartId="3.[151,269,1571,1595]" captionTargetBox="[186,1402,208,1538]" captionTargetId="figure@3.[170,1417,194,1547]" captionTargetPageId="3" captionText="FIGURES 9 – 11. Ranomafana pollocki sp. n.: Fig. 9, pronotum; Fig. 10, scutellum; Fig. 11, metendosternite." httpUri="https://zenodo.org/record/184418/files/figure.png" pageId="2" pageNumber="61">Fig. 9</figureCitation>
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) and male genitalia with subcylindrical tegminal accessory lobes (Figs. 19–20 vs. Fig. 21).
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</paragraph>
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</subSubSection>
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<subSubSection id="B1FA53E9F67E4A5B24D4F85BD7E00B85" lastPageId="6" lastPageNumber="65" pageId="2" pageNumber="61" type="description">
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<paragraph id="F95F0062F67E4A5F24D4F85BD16408C3" blockId="2.[151,1436,1586,2012]" box="[198,599,1986,2012]" pageId="2" pageNumber="61">
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<emphasis id="CB94DC70F67E4A5F24D4F85BD25208C3" bold="true" box="[198,353,1986,2012]" pageId="2" pageNumber="61">Description:</emphasis>
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Length
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<quantity id="3E18AD87F67E4A5F25D0F85BD16008C3" box="[450,595,1986,2012]" metricMagnitude="-3" metricUnit="m" metricValue="5.699999999999999" metricValueMax="6.6" metricValueMin="4.8" pageId="2" pageNumber="61" unit="mm" value="5.699999999999999" valueMax="6.6" valueMin="4.8">4.8–6.6 mm</quantity>
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.
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</paragraph>
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<caption id="AD9F50EAF67F4A5E2485F9BAD6170925" box="[151,1316,1571,1595]" httpUri="https://zenodo.org/record/184418/files/figure.png" pageId="3" pageNumber="62" targetBox="[186,1402,208,1538]" targetPageId="3">
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<paragraph id="F95F0062F67F4A5E2485F9BAD6170925" blockId="3.[151,1316,1571,1595]" box="[151,1316,1571,1595]" pageId="3" pageNumber="62">
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<emphasis id="CB94DC70F67F4A5E2485F9BAD27C0924" bold="true" box="[151,335,1571,1595]" pageId="3" pageNumber="62">FIGURES 9–11.</emphasis>
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<taxonomicName id="3EE07BE1F67F4A5E2547F9BAD1090925" box="[341,570,1571,1594]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pollocki" status="sp. nov.">
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<emphasis id="CB94DC70F67F4A5E2547F9BAD1090925" box="[341,570,1571,1594]" italics="true" pageId="3" pageNumber="62">Ranomafana pollocki</emphasis>
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</taxonomicName>
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<emphasis id="CB94DC70F67F4A5E2652F9BAD14F0924" bold="true" box="[576,636,1571,1595]" pageId="3" pageNumber="62">
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<taxonomicNameLabel id="D0A7610BF67F4A5E2652F9BAD14F0924" box="[576,636,1571,1595]" pageId="3" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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: Fig. 9, pronotum; Fig. 10, scutellum; Fig. 11, metendosternite.
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</paragraph>
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</caption>
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<paragraph id="F95F0062F67F4A5E24D4F9F0D2CC083C" blockId="3.[151,1436,1641,2027]" pageId="3" pageNumber="62">Colour (Figs. 3–4) rufotestaceous with pronotum along the middle, antennomeres 5–11, most of the head behind the anterior border of eyes (excluding the gular area), mesothorax (including scutellum but excluding part of the mesoventral disc), and metaventrite dark piceous to black. Body (Fig. 3a) slightly convex, elongate, 2.3–2.6 times as long as wide; dorsal surfaces covered with homogeneous, generally dense adpressed and few erect, yellowish setae.</paragraph>
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<paragraph id="F95F0062F67F4A5924D4F8A8D7A20D05" blockId="3.[151,1436,1641,2027]" lastBlockId="4.[151,1437,152,538]" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
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Head (
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<figureCitation id="61DB1CE7F67F4A5E2507F8A8D2680854" box="[277,347,1841,1867]" captionStart="FIGURES 5 – 8" captionStartId="2.[151,269,1357,1381]" captionTargetBox="[166,1372,314,1310]" captionTargetId="figure@2.[151,1436,314,1333]" captionTargetPageId="2" captionText="FIGURES 5 – 8. Ranomafana pollocki sp. n.: Fig. 5, mandible; Fig. 6, maxilla; Fig. 7, labium; Fig. 8, head." httpUri="https://zenodo.org/record/184417/files/figure.png" pageId="3" pageNumber="62">Fig. 8</figureCitation>
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) about 0.8 times as long as wide, moderately constricted behind the eyes; covered (Figs. 3b, 8) with coarse, subcontiguous, umbilicated punctures (see
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<bibRefCitation id="9D717D93F67F4A5E2751F8C0D0D1086C" author="Pollock" box="[835,994,1881,1907]" pageId="3" pageNumber="65" refString="Pollock, D. A. (1995) Classification, reconstructed phylogeny, and geographical history of genera of Pilipalpinae (Coleoptera: Tenebrionoidea: Pyrochroidae). Invertebrate Taxonomy, 9, 563 - 708." type="journal article" year="1995">Pollock 1995</bibRefCitation>
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||
), the punctures sparser ventrally, near posterior margin, and much less coarse in the gular area. Eyes entire, about 0.4 times as long as head, strongly protuberant, coarsely facetted with fine interfacetal setae. Frons broad, in some specimens slightly depressed at the centre behind the frontoclypeal suture, lateral ridges slightly elevated. Clypeus wider than long; frontoclypeal suture curved and very weakly indicated; apex bordered by a membrane. Gular sutures well developed (Fig. 4) and separated; gula longer than wide. Antennae (Figs. 3a, 4) extending at least to the base of the pronotum; antennomeres 1–5 filiform, subequal in length, 6–10 submoniliform; 11 conical. Labrum wider than long, broadly rounded, slightly convex; tormae converging posteriorly. Mandible (
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<figureCitation id="61DB1CE7F6784A592081FE89D7EB0E35" box="[1171,1240,272,298]" captionStart="FIGURES 5 – 8" captionStartId="2.[151,269,1357,1381]" captionTargetBox="[166,1372,314,1310]" captionTargetId="figure@2.[151,1436,314,1333]" captionTargetPageId="2" captionText="FIGURES 5 – 8. Ranomafana pollocki sp. n.: Fig. 5, mandible; Fig. 6, maxilla; Fig. 7, labium; Fig. 8, head." httpUri="https://zenodo.org/record/184417/files/figure.png" pageId="4" pageNumber="63">Fig. 5</figureCitation>
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) about 1.4 times as long as wide, apex bidentate; ventral surface fringed with an oblique row of fine microtrichia; mola 0.30 times as long as the mandible, with small, basal membranous lobe; prostheca hyaline, slightly shorter than the mola. Maxilla (
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<figureCitation id="61DB1CE7F6784A592559FE11D2A30EBD" box="[331,400,392,418]" captionStart="FIGURES 5 – 8" captionStartId="2.[151,269,1357,1381]" captionTargetBox="[166,1372,314,1310]" captionTargetId="figure@2.[151,1436,314,1333]" captionTargetPageId="2" captionText="FIGURES 5 – 8. Ranomafana pollocki sp. n.: Fig. 5, mandible; Fig. 6, maxilla; Fig. 7, labium; Fig. 8, head." httpUri="https://zenodo.org/record/184417/files/figure.png" pageId="4" pageNumber="63">Fig. 6</figureCitation>
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) with galea robust and longer than lacinia, densely setose in apical half; lacinia with fine dense setae, apex narrow and blunt; apical maxillary palpomere securiform. Labium (
|
||
<figureCitation id="61DB1CE7F6784A5920B3FE29D7D90ED5" box="[1185,1258,432,458]" captionStart="FIGURES 5 – 8" captionStartId="2.[151,269,1357,1381]" captionTargetBox="[166,1372,314,1310]" captionTargetId="figure@2.[151,1436,314,1333]" captionTargetPageId="2" captionText="FIGURES 5 – 8. Ranomafana pollocki sp. n.: Fig. 5, mandible; Fig. 6, maxilla; Fig. 7, labium; Fig. 8, head." httpUri="https://zenodo.org/record/184417/files/figure.png" pageId="4" pageNumber="63">Fig. 7</figureCitation>
|
||
) with mentum transverse, subtruncated apically, with sparse fine punctures; prementum broadly emarginated and covered with fine dense setae; apical palpomere cylindrical; hypopharynx mostly membranous.
|
||
</paragraph>
|
||
<caption id="AD9F50EAF6784A592485F90AD01609D2" httpUri="https://zenodo.org/record/184419/files/figure.png" pageId="4" pageNumber="63" targetBox="[183,1402,601,1653]" targetPageId="4">
|
||
<paragraph id="F95F0062F6784A592485F90AD01609D2" blockId="4.[151,1436,1683,1741]" pageId="4" pageNumber="63">
|
||
<emphasis id="CB94DC70F6784A592485F90AD26D09B4" bold="true" box="[151,350,1683,1707]" pageId="4" pageNumber="63">FIGURES 12–16.</emphasis>
|
||
<taxonomicName id="3EE07BE1F6784A592577F90AD17909B5" box="[357,586,1683,1706]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="pollocki" status="sp. nov.">
|
||
<emphasis id="CB94DC70F6784A592577F90AD17909B5" box="[357,586,1683,1706]" italics="true" pageId="4" pageNumber="63">Ranomafana pollocki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0A7610BF6784A592643F90AD1BD09B4" box="[593,654,1683,1707]" pageId="4" pageNumber="63" rank="species">
|
||
<emphasis id="CB94DC70F6784A592643F90AD1BD09B4" bold="true" box="[593,654,1683,1707]" pageId="4" pageNumber="63">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: Fig. 12, anterior leg; Fig. 13, middle leg; Fig. 14, posterior leg; Fig. 15, middle leg, apex of tibia and tarsus; Fig. 16, mesotarsal claw.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F95F0062F6784A5824D4F898D0E50E1D" blockId="4.[151,1436,1793,2019]" lastBlockId="5.[151,1436,152,258]" lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63">
|
||
Pronotum (
|
||
<figureCitation id="61DB1CE7F6784A592559F898D2A20804" box="[331,401,1793,1819]" captionStart="FIGURES 9 – 11" captionStartId="3.[151,269,1571,1595]" captionTargetBox="[186,1402,208,1538]" captionTargetId="figure@3.[170,1417,194,1547]" captionTargetPageId="3" captionText="FIGURES 9 – 11. Ranomafana pollocki sp. n.: Fig. 9, pronotum; Fig. 10, scutellum; Fig. 11, metendosternite." httpUri="https://zenodo.org/record/184418/files/figure.png" pageId="4" pageNumber="63">Fig. 9</figureCitation>
|
||
) about 0.70 times as long as wide, widest at anterior third; sides sinuate; base narrower than combined elytral bases; lateral sides (
|
||
<figureCitation id="61DB1CE7F6784A592694F8B0D1FF085C" box="[646,716,1833,1859]" captionStart="FIGURES 9 – 11" captionStartId="3.[151,269,1571,1595]" captionTargetBox="[186,1402,208,1538]" captionTargetId="figure@3.[170,1417,194,1547]" captionTargetPageId="3" captionText="FIGURES 9 – 11. Ranomafana pollocki sp. n.: Fig. 9, pronotum; Fig. 10, scutellum; Fig. 11, metendosternite." httpUri="https://zenodo.org/record/184418/files/figure.png" pageId="4" pageNumber="63">Fig. 9</figureCitation>
|
||
), each with 6 setiferous tubercles, the two anterior considerably smaller; anterior angles obtuse; posterior angles tuberculate basally; anterior edge slightly and broadly emarginated; posterior edge slightly emarginated, with narrow or inconspicuous bead at centre; disc convex with multiple somewhat irregular depressions medially and laterally; surface with umbilicated punctures similar in size and separation to those on head. Prosternum (Fig. 4) in front of coxae 0.9 times longer than the transverse diameter of the coxal cavity, moderately convex; prosternal process acute; notosternal suture complete; procoxae conical, projecting well below prosternum, trochantin clearly visible on slide mounted specimens; coxal cavities transverse, broadly open internally and externally.
|
||
</paragraph>
|
||
<paragraph id="F95F0062F6794A582485F91BD60409A2" blockId="5.[151,1436,1664,1725]" pageId="5" pageNumber="64">
|
||
<emphasis id="CB94DC70F6794A582485F91BD26E0985" bold="true" box="[151,349,1666,1690]" pageId="5" pageNumber="64">FIGURES 17–21.</emphasis>
|
||
Figs. 17–20,
|
||
<taxonomicName id="3EE07BE1F6794A5825FFF91BD1E20986" box="[493,721,1666,1689]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="pollocki" status="sp. nov.">
|
||
<emphasis id="CB94DC70F6794A5825FFF91BD1E20986" box="[493,721,1666,1689]" italics="true" pageId="5" pageNumber="64">Ranomafana pollocki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CB94DC70F6794A5826C5F91BD0210985" bold="true" box="[727,786,1666,1690]" pageId="5" pageNumber="64">
|
||
<taxonomicNameLabel id="D0A7610BF6794A5826C5F91BD0210985" box="[727,786,1666,1690]" pageId="5" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, %: Fig. 17, ventrites 4-5, Fig. 18, spiculum gastrale; Fig. 19, aedeagus, tegmen; Fig. 20, aedeagus, accessory lobes. Fig. 21,
|
||
<taxonomicName id="3EE07BE1F6794A582725F93FD0CC09A2" authority="Pollock" authorityName="Pollock" box="[823,1023,1702,1725]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="steineri">
|
||
<emphasis id="CB94DC70F6794A582725F93FD09709A2" box="[823,932,1702,1725]" italics="true" pageId="5" pageNumber="64">R. steineri</emphasis>
|
||
Pollock
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="265BBEC0F6794A582018F93FD75909A2" box="[1034,1130,1702,1725]" pageId="5" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
, aedeagus, apicale.
|
||
</paragraph>
|
||
<paragraph id="F95F0062F6794A5824D4F968D20608B4" blockId="5.[151,1436,1777,2003]" pageId="5" pageNumber="64">
|
||
Elytra about 1.80–1.90 times as long as wide and about 3.50–3.70 times as long as pronotum; sides subparallel, narrowed at posterior fourth and broadly rounded at apex, covered with dense punctures (Fig. 3d); epipleuron gradually narrowed apically, extending to ventrite 5. Scutellum (
|
||
<figureCitation id="61DB1CE7F6794A582039F8D8D7B00844" box="[1067,1155,1857,1883]" captionStart="FIGURES 9 – 11" captionStartId="3.[151,269,1571,1595]" captionTargetBox="[186,1402,208,1538]" captionTargetId="figure@3.[170,1417,194,1547]" captionTargetPageId="3" captionText="FIGURES 9 – 11. Ranomafana pollocki sp. n.: Fig. 9, pronotum; Fig. 10, scutellum; Fig. 11, metendosternite." httpUri="https://zenodo.org/record/184418/files/figure.png" pageId="5" pageNumber="64">Fig. 10</figureCitation>
|
||
) not abruptly elevated, broad, rounded laterally, covered with punctures similar to those of pronotum. Secondary setae (see
|
||
<bibRefCitation id="9D717D93F6794A582150F8F0D3E008B4" author="Pollock" pageId="5" pageNumber="65" refString="Pollock, D. A. (1995) Classification, reconstructed phylogeny, and geographical history of genera of Pilipalpinae (Coleoptera: Tenebrionoidea: Pyrochroidae). Invertebrate Taxonomy, 9, 563 - 708." type="journal article" year="1995">Pollock 1995</bibRefCitation>
|
||
) absent.
|
||
</paragraph>
|
||
<paragraph id="F95F0062F6794A5B24D4F820D2770ED5" blockId="5.[151,1436,1777,2003]" lastBlockId="6.[151,1437,152,1178]" lastPageId="6" lastPageNumber="65" pageId="5" pageNumber="64">
|
||
Mesoventrite subtriangular, separated by complete sutures from mesepisterna; mesepisterna narrowly separated anteriorly, disc with oblique carina, procoxal rest well developed; mesoventral process moderately long, extending well beyond middle of mesocoxal cavities, joined to mesoventral projection with single knob; mesocoxa projecting, narrowly separated; mesocoxal cavities open laterally, mesotrochantin exposed. Metaventrite about 0.60 times as long as wide, convex; discrimen 0.3 times as long as the metaventrite; exposed portion of metepisternum about 4.60 times as long as wide; metacoxae subcontiguous and transverse. Metendosternite (
|
||
<figureCitation id="61DB1CE7F67A4A5B2575FEF9D28F0E65" box="[359,444,352,378]" captionStart="FIGURES 9 – 11" captionStartId="3.[151,269,1571,1595]" captionTargetBox="[186,1402,208,1538]" captionTargetId="figure@3.[170,1417,194,1547]" captionTargetPageId="3" captionText="FIGURES 9 – 11. Ranomafana pollocki sp. n.: Fig. 9, pronotum; Fig. 10, scutellum; Fig. 11, metendosternite." httpUri="https://zenodo.org/record/184418/files/figure.png" pageId="6" pageNumber="65">Fig. 11</figureCitation>
|
||
) with long stalk and lateral arms; laminae well developed; anterior tendons widely separated on lateral arms. Metathoracic wings well developed, veins similar to those in
|
||
<taxonomicName id="3EE07BE1F67A4A5B20BCFE11D61B0EBE" box="[1198,1320,392,418]" class="Insecta" family="Pyrochroidae" genus="Ranomafana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="steineri">
|
||
<emphasis id="CB94DC70F67A4A5B20BCFE11D7F30EBE" box="[1198,1216,392,417]" italics="true" pageId="6" pageNumber="65">R</emphasis>
|
||
.
|
||
<emphasis id="CB94DC70F67A4A5B20C2FE11D61B0EBE" box="[1232,1320,392,417]" italics="true" pageId="6" pageNumber="65">steineri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="9D717D93F67A4A5B212BFE11D2040ED5" author="Pollock" pageId="6" pageNumber="65" refString="Pollock, D. A. (1995) Classification, reconstructed phylogeny, and geographical history of genera of Pilipalpinae (Coleoptera: Tenebrionoidea: Pyrochroidae). Invertebrate Taxonomy, 9, 563 - 708." type="journal article" year="1995">Pollock, 1995, Fig. 28</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F95F0062F67A4A5B24D4FE41D6530D75" blockId="6.[151,1437,152,1178]" pageId="6" pageNumber="65">
|
||
Tibiae (
|
||
<figureCitation id="61DB1CE7F67A4A5B2532FE41D29F0EED" box="[288,428,472,498]" captionStart="FIGURES 12 – 16" captionStartId="4.[151,269,1683,1707]" captionTargetBox="[183,1402,601,1653]" captionTargetId="figure@4.[167,1420,594,1659]" captionTargetPageId="4" captionText="FIGURES 12 – 16. Ranomafana pollocki sp. n.: Fig. 12, anterior leg; Fig. 13, middle leg; Fig. 14, posterior leg; Fig. 15, middle leg, apex of tibia and tarsus; Fig. 16, mesotarsal claw." httpUri="https://zenodo.org/record/184419/files/figure.png" pageId="6" pageNumber="65">Figs. 12–14</figureCitation>
|
||
) covered with dense fine setae; protibia (
|
||
<figureCitation id="61DB1CE7F67A4A5B279FFE41D0D20EED" box="[909,993,472,498]" captionStart="FIGURES 12 – 16" captionStartId="4.[151,269,1683,1707]" captionTargetBox="[183,1402,601,1653]" captionTargetId="figure@4.[167,1420,594,1659]" captionTargetPageId="4" captionText="FIGURES 12 – 16. Ranomafana pollocki sp. n.: Fig. 12, anterior leg; Fig. 13, middle leg; Fig. 14, posterior leg; Fig. 15, middle leg, apex of tibia and tarsus; Fig. 16, mesotarsal claw." httpUri="https://zenodo.org/record/184419/files/figure.png" pageId="6" pageNumber="65">Fig. 12</figureCitation>
|
||
) slightly expanded at apex; mesotibia (
|
||
<figureCitation id="61DB1CE7F67A4A5B248DFD99D3C40D05" box="[159,247,512,538]" captionStart="FIGURES 12 – 16" captionStartId="4.[151,269,1683,1707]" captionTargetBox="[183,1402,601,1653]" captionTargetId="figure@4.[167,1420,594,1659]" captionTargetPageId="4" captionText="FIGURES 12 – 16. Ranomafana pollocki sp. n.: Fig. 12, anterior leg; Fig. 13, middle leg; Fig. 14, posterior leg; Fig. 15, middle leg, apex of tibia and tarsus; Fig. 16, mesotarsal claw." httpUri="https://zenodo.org/record/184419/files/figure.png" pageId="6" pageNumber="65">Fig. 13</figureCitation>
|
||
) cylindrical and uniform; metatibia (
|
||
<figureCitation id="61DB1CE7F67A4A5B26A7FD99D03F0D05" box="[693,780,512,538]" captionStart="FIGURES 12 – 16" captionStartId="4.[151,269,1683,1707]" captionTargetBox="[183,1402,601,1653]" captionTargetId="figure@4.[167,1420,594,1659]" captionTargetPageId="4" captionText="FIGURES 12 – 16. Ranomafana pollocki sp. n.: Fig. 12, anterior leg; Fig. 13, middle leg; Fig. 14, posterior leg; Fig. 15, middle leg, apex of tibia and tarsus; Fig. 16, mesotarsal claw." httpUri="https://zenodo.org/record/184419/files/figure.png" pageId="6" pageNumber="65">Fig. 14</figureCitation>
|
||
) longer than the others and slightly expanded at apex. Tibial spurs paired, well developed. Tarsal formula 5–5–4; mesotarsomere 1 (
|
||
<figureCitation id="61DB1CE7F67A4A5B202BFDB1D7A30D5D" box="[1081,1168,552,578]" captionStart="FIGURES 12 – 16" captionStartId="4.[151,269,1683,1707]" captionTargetBox="[183,1402,601,1653]" captionTargetId="figure@4.[167,1420,594,1659]" captionTargetPageId="4" captionText="FIGURES 12 – 16. Ranomafana pollocki sp. n.: Fig. 12, anterior leg; Fig. 13, middle leg; Fig. 14, posterior leg; Fig. 15, middle leg, apex of tibia and tarsus; Fig. 16, mesotarsal claw." httpUri="https://zenodo.org/record/184419/files/figure.png" pageId="6" pageNumber="65">Fig. 15</figureCitation>
|
||
) subequal to 2+3, tarsomeres 2 and 3 similar in shape, 4 lobed below, 5 as long as 1+2. Claws (
|
||
<figureCitation id="61DB1CE7F67A4A5B27EFFDC9D7610D75" box="[1021,1106,592,618]" captionStart="FIGURES 12 – 16" captionStartId="4.[151,269,1683,1707]" captionTargetBox="[183,1402,601,1653]" captionTargetId="figure@4.[167,1420,594,1659]" captionTargetPageId="4" captionText="FIGURES 12 – 16. Ranomafana pollocki sp. n.: Fig. 12, anterior leg; Fig. 13, middle leg; Fig. 14, posterior leg; Fig. 15, middle leg, apex of tibia and tarsus; Fig. 16, mesotarsal claw." httpUri="https://zenodo.org/record/184419/files/figure.png" pageId="6" pageNumber="65">Fig. 16</figureCitation>
|
||
) with long basal tooth.
|
||
</paragraph>
|
||
<paragraph id="F95F0062F67A4A5B24D4FDE1D0BA0DFD" blockId="6.[151,1437,152,1178]" pageId="6" pageNumber="65">Abdomen about 1.5 times as long as wide; intercoxal process narrowly acute; ventrites 1–4 subequal; ventrites pubescent and free; pygidium, broadly rounded apically, covered with fine setae, with binding patches in posterior part; sexual dimorphism present in ventrite 5 (Fig. 17).</paragraph>
|
||
<paragraph id="F95F0062F67A4A5B24D4FD69D1C90CCD" blockId="6.[151,1437,152,1178]" pageId="6" pageNumber="65">Male genitalia: spiculum gastrale with separated branches (Fig. 18). Aedeagus (Figs. 19–20), as long as ventrites 4–5 combined; tegmen gradually narrowed to apex; basale glabrous; apicale 0.85 times as long as basale with few apical setae; accessory lobes apical (Fig. 20), subcylindrical, 0.5 times as long as apicale, with dense conspicuous setae from basal third; penis well sclerotised, as long as tegmen and accessory lobes, somewhat dorsoventrally compressed, broader at base, narrow from apical third, apex acute, dorsal apical third with two membranous areas, basal struts short.</paragraph>
|
||
<paragraph id="F95F0062F67A4A5B24D4FC79D2C40B55" blockId="6.[151,1437,152,1178]" pageId="6" pageNumber="65">Female genitalia: spiculum ventrale, as long as ventrites 1–5; ovipositor as long as ventrites 2–5; coxites pubescent, slightly sclerotised, each coxite divided into 3 parts; styli apical, well developed. Spermatheca 3- branched, bursa membranous.</paragraph>
|
||
<paragraph id="F95F0062F67A4A5B24D4FBC1D7E00B85" blockId="6.[151,1437,152,1178]" pageId="6" pageNumber="65">
|
||
Sexual dimorphism: apex of ventrite 5 slightly emarginated in male (Fig. 17) and rounded in female; this form of sexual dimorphism had not been previously recorded in
|
||
<taxonomicName id="3EE07BE1F67A4A5B279BFB19D7FC0B85" authority="(Pollock 1995)" baseAuthorityName="Pollock" baseAuthorityYear="1995" box="[905,1231,1152,1178]" class="Insecta" family="Pyrochroidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="subFamily" subFamily="Pilipalpinae">
|
||
Pilipalpinae (
|
||
<bibRefCitation id="9D717D93F67A4A5B2036FB19D7F50B85" author="Pollock" box="[1060,1222,1152,1178]" pageId="6" pageNumber="65" refString="Pollock, D. A. (1995) Classification, reconstructed phylogeny, and geographical history of genera of Pilipalpinae (Coleoptera: Tenebrionoidea: Pyrochroidae). Invertebrate Taxonomy, 9, 563 - 708." type="journal article" year="1995">Pollock 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |