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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.981.53583" ID-GBIF-Dataset="8e14a5b5-aa20-4050-b2b4-f393764fc6c7" ID-PMC="PMC7644702" ID-Pensoft-Pub="1313-2970-981-1" ID-Pensoft-UUID="7C44C797C4125A71BAE032A55E6FA5DC" ID-PubMed="33199958" ID-ZooBank="5194AAC86B8A473F8A41470A60182A0B" ModsDocID="1313-2970-981-1" checkinTime="1604011718171" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Sajan, Sheikh, Tripathy, Basudev, Meng, Kaibaryer, Asami, Takahiro &amp; Ablett, Jonathan D." docDate="2020" docId="5A497BF8D0A55EF5AD06461C6BA4E177" docLanguage="en" docName="ZooKeys 981: 1-220" docOrigin="ZooKeys 981" docSource="http://dx.doi.org/10.3897/zookeys.981.53583" docTitle="Metalycaeus rathouisianus" docType="treatment" docVersion="6" id="7C44C797C4125A71BAE032A55E6FA5DC" lastPageNumber="1" masterDocId="7C44C797C4125A71BAE032A55E6FA5DC" masterDocTitle="Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue" masterLastPageNumber="220" masterPageNumber="1" pageNumber="1" updateTime="1668169683351" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sajan, Sheikh</mods:namePart>
<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India &amp; Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2785-6824</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tripathy, Basudev</mods:namePart>
<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Meng, Kaibaryer</mods:namePart>
<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Asami, Takahiro</mods:namePart>
<mods:affiliation>Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5706-0272</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>981</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>220</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.981.53583</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.981.53583</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-981-1</mods:identifier>
<mods:identifier type="ZooBank">5194AAC86B8A473F8A41470A60182A0B</mods:identifier>
<mods:identifier type="Pensoft-UUID">7C44C797C4125A71BAE032A55E6FA5DC</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="168715125" LSID="urn:lsid:plazi:treatment:5A497BF8D0A55EF5AD06461C6BA4E177" httpUri="http://treatment.plazi.org/id/5A497BF8D0A55EF5AD06461C6BA4E177" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="5A497BF8-D0A5-5EF5-AD06-461C6BA4E177" authority="(Heude, 1882)" baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus rathouisianus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">Metalycaeus rathouisianus (Heude, 1882)</taxonomicName>
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Metalycaeus rathouisianus (Heude, 1882), syntype: labelled as lectotype of Alycaeus dolomiticus (HMT- 221 a). Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.981.53583.figure34" httpUri="https://binary.pensoft.net/fig/468679" pageId="0" pageNumber="1">Figs 34</figureCitation>
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Alycaeus neglectus Heude, 1885, syntype (HMT- 220 a; synonym of Metalycaeus rathouisianus (Heude, 1882 )). Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.981.53583.figure35" httpUri="https://binary.pensoft.net/fig/468680" pageId="0" pageNumber="1">, 35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1882" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus rathouisianus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">Alycaeus rathouisianus</taxonomicName>
Heude, 1882: 7, pl. 12, figs 12, 12a.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1882" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus rathouisianus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">Alycaeus rathouisianus</taxonomicName>
-
<bibRefCitation author="Moellendorff, OF von" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/16360044" refId="B132" refString="Moellendorff, OF von, 1882. Materialen zur Fauna for China. Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 9: 251-278, 337-356. https://biodiversitylibrary.org/page/16360044" title="Materialen zur Fauna for China. Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 9: 251 - 278, 337 - 356." url="https://biodiversitylibrary.org/page/16360044" year="1882">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1882
</bibRefCitation>
: 345.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycoeus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Alycoeus</taxonomicName>
[sic]
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="neglectus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="neglectus">neglectus</taxonomicName>
Heude, 1885: 96, pl. 24, fig. 4. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1882" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Orthalycaeus) rathouisianus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus" subGenus="Orthalycaeus">Alycaeus (Orthalycaeus) rathouisianus</taxonomicName>
-
<bibRefCitation author="Moellendorff, OF von" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="156 - 210" publicationUrl="https://www.biodiversitylibrary.org/page/16355855" refId="B134" refString="Moellendorff, OF von, 1886. Materialen zur Fauna for China. Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 13: 156 - 210, https://www.biodiversitylibrary.org/page/16355855" title="Materialen zur Fauna for China." url="https://www.biodiversitylibrary.org/page/16355855" volume="13" year="1886">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1886
</bibRefCitation>
: 169.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Orthalycaeus) neglectus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="neglectus" subGenus="Orthalycaeus">Alycaeus (Orthalycaeus) neglectus</taxonomicName>
-
<bibRefCitation author="Moellendorff, OF von" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="156 - 210" publicationUrl="https://www.biodiversitylibrary.org/page/16355855" refId="B134" refString="Moellendorff, OF von, 1886. Materialen zur Fauna for China. Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 13: 156 - 210, https://www.biodiversitylibrary.org/page/16355855" title="Materialen zur Fauna for China." url="https://www.biodiversitylibrary.org/page/16355855" volume="13" year="1886">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1886
</bibRefCitation>
: 170.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) neglectus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="neglectus" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) neglectus</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1897
</bibRefCitation>
: 149;
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1000211" refId="B97" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin, https://biodiversitylibrary.org/page/1000211" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://biodiversitylibrary.org/page/1000211" year="1902">Kobelt 1902</bibRefCitation>
: 360.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1882" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) rathouisianus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) rathouisianus</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1897
</bibRefCitation>
: 149;
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1000211" refId="B97" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin, https://biodiversitylibrary.org/page/1000211" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://biodiversitylibrary.org/page/1000211" year="1902">Kobelt 1902</bibRefCitation>
: 362.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1890" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus dolomiticus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dolomiticus">Alycaeus dolomiticus</taxonomicName>
Heude, 1890: 130, pl. 38, fig. 1.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus rathouisianus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">Chamalycaeus rathouisianus</taxonomicName>
-
<bibRefCitation author="Yen, T-C" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" pageId="0" pageNumber="1" refId="B229" refString="Yen, T-C, 1939. Die Chinesischen Land-und Suesswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" title="Die Chinesischen Land-und Suesswasser-Gastropoden des Natur-Museums Senckenberg." year="1939">Yen 1939</bibRefCitation>
: 30, pl. 2, fig. 36.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus (Chamalycaeus) rathouisianus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus" subGenus="Chamalycaeus">Chamalycaeus (Chamalycaeus) rathouisianus</taxonomicName>
-
<bibRefCitation author="Zilch, A" journalOrPublisher="Ltd., Beijing" pageId="0" pageNumber="1" refId="B232" refString="Zilch, A, 1957. Die Typen und Typoide des Natur-Museums Senckenberg, 20: Mollusca, Cyclophoridae, Alycaeinae. Archiv fuer Molluskenkunde 86(4/6): 141-150." title="Die Typen und Typoide des Natur-Museums Senckenberg, 20: Mollusca, Cyclophoridae, Alycaeinae. Archiv fuer Molluskenkunde 86 (4 / 6): 141 - 150." year="1957">Zilch 1957</bibRefCitation>
: 145.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus rathouisianus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">Metalycaeus rathouisianus</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: 98-103, figs 55A-C, 63A, B, 65A, B.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicharax neglectus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="neglectus">Dicharax (?) neglectus</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: 105-107, fig. 69K.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;E collibus juxta civitatem Song-kiang (松江),
<normalizedToken originalValue="provinciæ">provinciae</normalizedToken>
Kiang-sou, ad montes districtus Tong-lieou, sed non ubique&quot; (
<taxonomicName authorityName="Heude" authorityYear="1882" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus rathouisianus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus rathouisianus</emphasis>
</taxonomicName>
); &quot;
<emphasis italics="true" pageId="0" pageNumber="1">
ad rupestres colles civitatis
<normalizedToken originalValue="Kien-té">Kien-te</normalizedToken>
,
<normalizedToken originalValue="provinciæ">provinciae</normalizedToken>
</emphasis>
<normalizedToken originalValue="Ngan-houé">Ngan-houe</normalizedToken>
<emphasis italics="true" pageId="0" pageNumber="1">legeram</emphasis>
&quot; (
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus neglectus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus neglectus</emphasis>
</taxonomicName>
); &quot;Ad calcarios lapides
<normalizedToken originalValue="juxtà">juxta</normalizedToken>
civitatem
<emphasis italics="true" pageId="0" pageNumber="1">Tchen-kiang</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">Kiang-sou</emphasis>
)&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
China,
<normalizedToken originalValue="Tsing-p’on">Tsing-p'on</normalizedToken>
, MCZ 167143 (23 syntypes of
<taxonomicName lsidName="A. rathouisianus" pageId="0" pageNumber="1" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">A. rathouisianus</emphasis>
</taxonomicName>
); Sung-kiang: China, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Heude, SMF 39243 (5 syntypes of
<taxonomicName lsidName="A. rathouisianus" pageId="0" pageNumber="1" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">A. rathouisianus</emphasis>
</taxonomicName>
); same data, 39303 (3 syntypes of
<taxonomicName lsidName="A. rathouisianus" pageId="0" pageNumber="1" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">A. rathouisianus</emphasis>
</taxonomicName>
); China, Tchein-kiang (Kiang-sou), MCZ 167209 (29 syntypes of
<taxonomicName lsidName="A. dolomiticus" pageId="0" pageNumber="1" rank="species" species="dolomiticus">
<emphasis italics="true" pageId="0" pageNumber="1">A. dolomiticus</emphasis>
</taxonomicName>
); HMT-221a (labelled as lectotype of
<taxonomicName authorityName="Heude" authorityYear="1890" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus dolomiticus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dolomiticus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus dolomiticus</emphasis>
</taxonomicName>
, Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Metalycaeus rathouisianus (Heude, 1882), syntype: labelled as lectotype of Alycaeus dolomiticus (HMT- 221 a). Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.981.53583.figure34" httpUri="https://binary.pensoft.net/fig/468679" pageId="0" pageNumber="1">34</figureCitation>
) deposited in IZCAS: locality information were deleted; HMT-221 (labelled as paralectotype of
<taxonomicName authorityName="Heude" authorityYear="1890" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus dolomiticus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dolomiticus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus dolomiticus</emphasis>
</taxonomicName>
), same data with HMT-221; HMT-220a (1 syntype, labelled as lectotype of
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus neglectus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus neglectus</emphasis>
</taxonomicName>
, Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Alycaeus neglectus Heude, 1885, syntype (HMT- 220 a; synonym of Metalycaeus rathouisianus (Heude, 1882 )). Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.981.53583.figure35" httpUri="https://binary.pensoft.net/fig/468680" pageId="0" pageNumber="1">35</figureCitation>
), deposited in IZCAS: Tong-lieu (Dong-Liu town, Dong-Zhi County, Chi-Zhou City, An-Hui Province, China); Same data, HMT-220 (1 syntype, labelled as paralectotype of
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus neglectus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus neglectus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1890" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus dolomiticus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dolomiticus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus dolomiticus</emphasis>
</taxonomicName>
Heude, 1890 is a synonym of
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus rathouisianus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus rathouisianus</emphasis>
</taxonomicName>
(see
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
According to
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.141074" author="Johnson, RJ" journalOrPublisher="Cambridge, Massachusetts (Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University)" pageId="0" pageNumber="1" refId="B92" refString="Johnson, RJ, 1973. Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude. Cambridge, Massachusetts (Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University), DOI: https://doi.org/10.5962/bhl.title.141074" title="Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude." url="https://doi.org/10.5962/bhl.title.141074" year="1973">Johnson (1973)</bibRefCitation>
, paratypes of
<taxonomicName lsidName="A. neglectus" pageId="0" pageNumber="1" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">A. neglectus</emphasis>
</taxonomicName>
are present in the MCZ (inv. number: 167222) and the USNM (inventory number: 472336). In the revision of Chinese
<taxonomicName authorityName="Blanford" authorityYear="1864" family="Alycaeidae" lsidName="" pageId="0" pageNumber="1" rank="family">Alycaeidae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al 2017
</bibRefCitation>
) we examined the former sample, but the locality (&quot;China, Tong-lieou&quot; in Anhui Province) did not match the type locality given in the original description (
<normalizedToken originalValue="“Kien-té”">&quot;Kien-te&quot;</normalizedToken>
in Zhejiang Province). Consequently, we did not consider them as type specimens; moreover, on a hand-written label of the MCZ 167222 sample the following text was written: &quot;not types, only holotype in
<normalizedToken originalValue="Heude’s">Heude's</normalizedToken>
coll.&quot;. We have now been able to examine the specimens in the Beijing museum, and found that the specimens labelled as types also have the same type locality (
<normalizedToken originalValue="“Tong-lieu”">&quot;Tong-lieu&quot;</normalizedToken>
). The Beijing specimens were also labelled as lectotype (HMT-221a), and paralectotypes (HMT-221). It appears that although the locality on the type specimen labels and that in the original description do not match, the shells labelled as types are indeed types of
<taxonomicName lsidName="A. neglectus" pageId="0" pageNumber="1" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">A. neglectus</emphasis>
</taxonomicName>
. Since we did not find traces of the lectotype designation, we consider all type specimens of
<taxonomicName lsidName="A. neglectus" pageId="0" pageNumber="1" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">A. neglectus</emphasis>
</taxonomicName>
syntypes.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Since we did not consider the specimens in the MCZ types in
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. (2017)
</bibRefCitation>
, we had to rely on the original description regarding the identity of
<taxonomicName lsidName="A. neglectus" pageId="0" pageNumber="1" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">A. neglectus</emphasis>
</taxonomicName>
, and based on the low protoconch in the figure of the original description, we tentatively classified it in
<taxonomicName authorityName="Kobelt &amp; Moellendorff" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicharax" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Dicharax</emphasis>
</taxonomicName>
. However, this opinion must now be revised: the shells labelled as types in the Beijing Museum and in the MCZ are identical, and they are also identical to typical
<taxonomicName authorityName="Heude" authorityYear="1882" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus rathouisianus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus rathouisianus</emphasis>
</taxonomicName>
shells. Thus,
<taxonomicName lsidName="A. neglectus" pageId="0" pageNumber="1" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">A. neglectus</emphasis>
</taxonomicName>
is considered a synonym of
<taxonomicName lsidName="A. rathouisianus" pageId="0" pageNumber="1" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">A. rathouisianus</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/zookeys.981.53583.figure34" httpUri="https://binary.pensoft.net/fig/468679" pageId="0" pageNumber="1" start="Figure 34" startId="F34">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 34.</emphasis>
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus rathouisianus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus rathouisianus</emphasis>
</taxonomicName>
(Heude, 1882), syntype: labelled as lectotype of
<taxonomicName authorityName="Heude" authorityYear="1890" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus dolomiticus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dolomiticus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus dolomiticus</emphasis>
</taxonomicName>
(HMT-221a). Photographs: Kaibaryer Meng.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.981.53583.figure35" httpUri="https://binary.pensoft.net/fig/468680" pageId="0" pageNumber="1" start="Figure 35" startId="F35">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 35.</emphasis>
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus neglectus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="neglectus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus neglectus</emphasis>
</taxonomicName>
Heude, 1885, syntype (HMT-220a; synonym of
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus rathouisianus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rathouisianus">
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus rathouisianus</emphasis>
</taxonomicName>
(Heude, 1882)). Photographs: Kaibaryer Meng.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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