151 lines
20 KiB
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151 lines
20 KiB
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<document id="7A73DE4331EBC4A0DC665ACD54FA372E" ID-DOI="10.37520/fi.2023.004" ID-ISSN="2533-4069" ID-Zenodo-Dep="10481035" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704924026152" checkinUser="felipe" docAuthor="Smith, MacKenzie A., Greenwalt, Dale E. & Manchester, Steven R." docDate="2023" docId="4D2487A3EF598265FEDBFB006AA2FDB6" docLanguage="en" docName="FossilImprint.79.1.37-88.pdf" docOrigin="Fossil Imprint 79 (1)" docSource="http://dx.doi.org/10.37520/fi.2023.004" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Abies MILL." docType="treatment" docVersion="2" lastPageNumber="43" masterDocId="B11DFFDBEF5F8263FF8CFFB16F30FFE9" masterDocTitle="Diverse Fruits And Seeds Of The Mid-Eocene Kishenehn Formation, Northwestern Montana, Usa, And Their Implications For Biogeography" masterLastPageNumber="88" masterPageNumber="37" pageNumber="43" updateTime="1704925034431" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="6530E1DA506D8E272EED644E98810107">Diverse Fruits And Seeds Of The Mid-Eocene Kishenehn Formation, Northwestern Montana, Usa, And Their Implications For Biogeography</mods:title>
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<mods:namePart id="121C36C5C19D572B580F64E9BCB93D76">Smith, MacKenzie A.</mods:namePart>
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<mods:namePart id="C5D7C061CEAF138E1FF26A09ADD2F5A9">Greenwalt, Dale E.</mods:namePart>
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<mods:namePart id="FBCB099B8CFEFBF99CF748FB55149370">Manchester, Steven R.</mods:namePart>
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<mods:typeOfResource id="9989207663AA073049697399E3B2CBAE">text</mods:typeOfResource>
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<mods:title id="7210AF6DA7D0AA1C07EE812AA3AFD864">Fossil Imprint</mods:title>
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<mods:part id="097B2F497FE6F25A6BF150B0BE1D32D5">
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<mods:date id="7C4257E2F47FABBD789CBB4CA4505CB3">2023</mods:date>
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<mods:number id="2DFC7F09F4361BEE37CBD2E442BF9AC3">2023-11-07</mods:number>
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<mods:number id="B574DE68011CFDD3FBA4F85F0C1537E1">79</mods:number>
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<mods:identifier id="21A56519A61122B84FEDFC725097B056" type="ISSN">2533-4069</mods:identifier>
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<treatment id="4D2487A3EF598265FEDBFB006AA2FDB6" ID-DOI="http://doi.org/10.5281/zenodo.10481045" ID-Zenodo-Dep="10481045" LSID="urn:lsid:plazi:treatment:4D2487A3EF598265FEDBFB006AA2FDB6" httpUri="http://treatment.plazi.org/id/4D2487A3EF598265FEDBFB006AA2FDB6" lastPageNumber="43" pageId="6" pageNumber="43">
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<subSubSection id="8D97653EEF598265FEDBFB006ECDFB03" pageId="6" pageNumber="43" type="nomenclature">
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<paragraph id="C53236B5EF598265FEDBFB006D1AFB23" blockId="6.[343,554,1201,1258]" box="[343,554,1201,1226]" pageId="6" pageNumber="43">
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<heading id="9E7A81D9EF598265FEDBFB006D1AFB23" box="[343,554,1201,1226]" centered="true" fontSize="8" level="2" pageId="6" pageNumber="43" reason="2">
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<emphasis id="F7F9EAA7EF598265FEDBFB006E90FB20" bold="true" box="[343,416,1201,1225]" pageId="6" pageNumber="43">Genus</emphasis>
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<taxonomicName id="028D4D36EF598265FE2AFB006D1AFB23" ID-CoL="627WF" ID-ENA="3319" authority="MILL." authorityName="MILL." box="[422,554,1201,1226]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="43" phylum="Tracheophyta" rank="genus">
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<emphasis id="F7F9EAA7EF598265FE2AFB006D1AFB23" bold="true" box="[422,554,1201,1226]" pageId="6" pageNumber="43">
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<emphasis id="F7F9EAA7EF598265FE2AFB006ED3FB20" bold="true" box="[422,483,1201,1225]" italics="true" pageId="6" pageNumber="43">Abies</emphasis>
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<smallCapsWord id="C3D4A069EF598265FE65FB036D12FB2E" baselines="1220,1219" box="[489,546,1202,1226]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Mill" pageId="6" pageNumber="43">MILL</smallCapsWord>
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.
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="C53236B5EF598265FE0FFB636ECDFB03" blockId="6.[343,554,1201,1258]" box="[387,509,1234,1258]" pageId="6" pageNumber="43">
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<figureCitation id="5DB62A30EF598265FE0FFB636ECDFB03" box="[387,509,1234,1258]" captionStart="Text-fig" captionStartId="7.[125,206,1821,1842]" captionTargetBox="[125,1440,151,1805]" captionTargetId="figure-6@7.[699,1100,583,1220]" captionTargetPageId="7" captionText="Text-fig. 3. Pinaceae from the Kishenehn Formation. a: USNM PAL 621424 Abies sp. seed. Scale bar 5mm. b: USNM PAL 621991 Larix sp. winged seed. Scale bar 2mm. c: USNM PAL 622093cf. Larix sp. ovuliferous cone. Scale bar 3mm. d: USNM PAL 776558 Picea sp. 1 winged seed. Scale bar 3mm. e: USNM PAL 776565 Picea sp. 2 seed. Scale bar 5mm. f: USNM PAL 619964 Pinaceae Cone 1. Scale bar 1cm. g: USNM PAL 776557 Pinaceae Cone 2. Scale bar 1cm. h: USNM PAL 620025 Pinaceae Cone 3. Scale bar 1cm. i: USNM PAL 776556 Pinus sp. C1. Scale bar 2cm. j: DMNH EPI. 48178 Pinus sp. L1 3-leaf fascicle. Scale bar 2cm. k: USNM PAL 776570 Pinus sp. C2 Pinus subg. Strobus female cone. Scale bar 1cm. l: USNM PAL 621980 Pinus subg. Strobus, Pinus sp. S1 winged seed. Scale bar 3mm. m: USNM PAL 622300 Pinus subg. Strobus, Pinus sp. S1 winged seed. Scale bar 5mm. n: USNM PAL 776567 Pinus subg. Strobus, Pinus sp. S2 seed. Scale bar 2mm. o: USNM PAL 620785 Pinus subg. Strobus, Pinus sp. S3 winged seed, with seed body disarticulated. Scale bar 2mm. p: USNM PAL 626064 Pinus subg. Pinus female cone scale Pinus sp. C3. Scale" figureDoi="http://doi.org/10.5281/zenodo.10481041" httpUri="https://zenodo.org/record/10481041/files/figure.png" pageId="6" pageNumber="43">Text-fig. 3a</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8D97653EEF598265FF2AFAB36AA2FDB6" pageId="6" pageNumber="43" type="description">
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<paragraph id="C53236B5EF598265FF2AFAB36D29FAF3" blockId="6.[166,537,1282,1306]" box="[166,537,1282,1306]" pageId="6" pageNumber="43">M a t e r i a l. USNM PAL 621424.</paragraph>
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<paragraph id="C53236B5EF598265FF2AFA836E5EFAA0" blockId="6.[166,366,1329,1354]" box="[166,366,1329,1354]" pageId="6" pageNumber="43">L o c a l i t y. Park.</paragraph>
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<paragraph id="C53236B5EF598265FF2AFAD36E8BF9B5" blockId="6.[128,768,1377,1629]" pageId="6" pageNumber="43">
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D e s c r i p t i o n. Winged seed linear,
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<quantity id="02759B50EF598265FDD6FAD06D8CFA90" box="[602,700,1377,1401]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="6" pageNumber="43" unit="mm" value="11.7">11.7 mm</quantity>
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long,
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<quantity id="02759B50EF598265FF0DFA306FE4FA70" box="[129,212,1409,1433]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="43" unit="mm" value="5.3">5.3 mm</quantity>
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wide at widest; seed body elliptical
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<quantity id="02759B50EF598265FDE0FA336D8FFA70" box="[620,703,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="43" unit="mm" value="3.8">3.8 mm</quantity>
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long,
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<quantity id="02759B50EF598265FF0CFA136FE0FA53" box="[128,208,1442,1466]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="43" unit="mm" value="2.3">2.3 mm</quantity>
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wide oriented with long axis parallel to medial edge; seed body on proximal side of seed; seed body contains numerous, round resin vesicles; abmedial margin begins approximately one quarter of the way up seed body extending so widest area of wing is distal; distal edge straight; medial edge convex in distal quarter.
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</paragraph>
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<paragraph id="C53236B5EF598265FF2AF9C46AA2FDB6" blockId="6.[128,768,1652,2100]" lastBlockId="6.[843,1482,518,607]" pageId="6" pageNumber="43">
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R e m a r k s. Winged seeds with resin vesicles and with the wings widest in the distal half belong to
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<taxonomicName id="028D4D36EF598265FDFDF9246E7BF927" authority="(Wolfe and Schorn 1990)" baseAuthorityName="Wolfe and Schorn" baseAuthorityYear="1990" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="43" phylum="Tracheophyta" rank="genus">
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<emphasis id="F7F9EAA7EF598265FDFDF9246D9CF944" box="[625,684,1685,1709]" italics="true" pageId="6" pageNumber="43">Abies</emphasis>
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(
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<bibRefCitation id="A11C4B44EF598265FD31F9246E73F927" author="Wolfe, J. A. & Schorn, H. E." pageId="6" pageNumber="43" pagination="1 - 69" refId="ref44290" refString="Wolfe, J. A., Schorn, H. E. (1990): Taxonomic revision of the Spermatopsida of the Oligocene Creede flora, Southern Colorado. - U. S. Geological Survey Bulletin, 1923: 1 - 69. https: // doi. org / 10.3133 / b 1923" type="journal article" year="1990">Wolfe and Schorn 1990</bibRefCitation>
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)
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</taxonomicName>
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.
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<taxonomicName id="028D4D36EF598265FED5F9076EA4F927" baseAuthorityName="Wolfe and Schorn" baseAuthorityYear="1990" box="[345,404,1718,1742]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="43" phylum="Tracheophyta" rank="genus">
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<emphasis id="F7F9EAA7EF598265FED5F9076EA4F927" box="[345,404,1718,1742]" italics="true" pageId="6" pageNumber="43">Abies</emphasis>
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</taxonomicName>
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pollen is known from the early Eocene McAbee and Quilchena floras of
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<collectingRegion id="0749F857EF598265FDC9F9676DCFF907" box="[581,767,1750,1774]" country="Canada" name="British Columbia" pageId="6" pageNumber="43">British Columbia</collectingRegion>
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(
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<bibRefCitation id="A11C4B44EF598265FF0BF9466E63F8E7" author="Dillhoff, R. M. & Leopold, E. B. & Manchester, S. R." box="[135,339,1782,1807]" pageId="6" pageNumber="43" pagination="151 - 166" refId="ref35034" refString="Dillhoff, R. M., Leopold, E. B., Manchester, S. R. (2005): The McAbee flora of British Columbia and its relation to the early - middle Eocene Okanagan Highlands flora of the Pacific Northwest. - Canadian Journal of Earth Sciences, 42: 151 - 166. https: // doi. org / 10.1139 / e 04 - 084" type="journal article" year="2005">Dillhoff et al. 2005</bibRefCitation>
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,
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<bibRefCitation id="A11C4B44EF598265FED0F9466D3EF8E6" author="Lowe, A. J. & Greenwood, D. R. & West, C. K. & Galloway, J. M. & Sudermann, M. & Reichgelt, T." box="[348,526,1783,1807]" pageId="6" pageNumber="43" refId="ref39070" refString="Lowe, A. J., Greenwood, D. R., West, C. K., Galloway, J. M., Sudermann, M., Reichgelt, T. (2018): Plant community ecology and climate on an upland volcanic landscape during the early Eocene Climatic Optimum: McAbee Fossil Beds, British Columbia, Canada. - Palaeogeography, Palaeoclimatology, Palaeoecology, 511: 443 - 338. https: // doi. org / 10.1016 / j. palaeo. 2018.09.010" type="book" year="2018">Lowe et al. 2018</bibRefCitation>
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,
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<bibRefCitation id="A11C4B44EF598265FD9BF9466DC8F8E6" author="Mathewes, R. W. & Greenwood, D. R. & Archibald, S. B." box="[535,760,1783,1807]" pageId="6" pageNumber="43" pagination="1 - 17" refId="ref40238" refString="Mathewes, R. W., Greenwood, D. R., Archibald, S. B. (2016): Paleoenvironment of the Quilchena flora, British Columbia, during the early Eocene Climatic Optimum. - Canadian Journal of Earth Sciences, 53: 1 - 17. https: // doi. org / 10.1139 / cjes- 2015 - 0163" type="journal article" year="2016">Mathewes et al. 2016</bibRefCitation>
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) and Yellowstone,
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<collectingRegion id="0749F857EF598265FEDEF8A66E8BF8C6" box="[338,443,1815,1839]" country="United States of America" name="Wyoming" pageId="6" pageNumber="43">Wyoming</collectingRegion>
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(
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<bibRefCitation id="A11C4B44EF598265FE5FF8A66D69F8C6" author="Wing, S. L." box="[467,601,1815,1839]" pageId="6" pageNumber="43" pagination="748 - 784" refId="ref44013" refString="Wing, S. L. (1987): Eocene and Oligocene floras of the Rocky Mountains. - Annals of the Missouri Botanical Garden, 74 (4): 748 - 784. https: // doi. org / 10.2307 / 2399449" type="journal article" year="1987">Wing 1987</bibRefCitation>
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). Macrofossils are known from the early Eocene McAbee and Falkland floras of
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<collectingRegion id="0749F857EF598265FF64F8E96E95F899" box="[232,421,1880,1904]" country="Canada" name="British Columbia" pageId="6" pageNumber="43">British Columbia</collectingRegion>
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(
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<bibRefCitation id="A11C4B44EF598265FE3BF8E96DA5F899" author="Dillhoff, R. M. & Leopold, E. B. & Manchester, S. R." box="[439,661,1880,1904]" pageId="6" pageNumber="43" pagination="151 - 166" refId="ref35034" refString="Dillhoff, R. M., Leopold, E. B., Manchester, S. R. (2005): The McAbee flora of British Columbia and its relation to the early - middle Eocene Okanagan Highlands flora of the Pacific Northwest. - Canadian Journal of Earth Sciences, 42: 151 - 166. https: // doi. org / 10.1139 / e 04 - 084" type="journal article" year="2005">Dillhoff et al. 2005</bibRefCitation>
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,
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<bibRefCitation id="A11C4B44EF598265FD28F8E86FEBF878" author="Lowe, A. J. & Greenwood, D. R. & West, C. K. & Galloway, J. M. & Sudermann, M. & Reichgelt, T." pageId="6" pageNumber="43" refId="ref39070" refString="Lowe, A. J., Greenwood, D. R., West, C. K., Galloway, J. M., Sudermann, M., Reichgelt, T. (2018): Plant community ecology and climate on an upland volcanic landscape during the early Eocene Climatic Optimum: McAbee Fossil Beds, British Columbia, Canada. - Palaeogeography, Palaeoclimatology, Palaeoecology, 511: 443 - 338. https: // doi. org / 10.1016 / j. palaeo. 2018.09.010" type="book" year="2018">Lowe et al. 2018</bibRefCitation>
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,
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<bibRefCitation id="A11C4B44EF598265FF65F8C86E9AF878" author="Smith, R. Y. & Basinger, J. F. & Greenwood, D. R." box="[233,426,1913,1937]" pageId="6" pageNumber="43" pagination="309 - 328" refId="ref42801" refString="Smith, R. Y., Basinger, J. F., Greenwood, D. R. (2012): Early Eocene plant diversity and dynamics in the Falkland flora, British Columbia, Canada. - Palaeobiodiversity and Palaeoenvironments, 92: 309 - 328. https: // doi. org / 10.1007 / s 12549 - 011 - 0061 - 5" type="journal article" year="2012">Smith et al. 2012</bibRefCitation>
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), Republic flora of
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<collectingRegion id="0749F857EF598265FD0DF8C86C30F878" box="[641,768,1913,1937]" country="United States of America" name="Washington" pageId="6" pageNumber="43">Washington</collectingRegion>
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(
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<bibRefCitation id="A11C4B44EF598265FF04F8286E71F858" author="Pigg, K. B. & DeVore, M. L. & Volkman, K. E." box="[136,321,1945,1969]" pageId="6" pageNumber="43" refId="ref41920" refString="Pigg, K. B., DeVore, M. L., Volkman, K. E. (2011): Fossil Plants from Republic: A Guidebook. - Stonerose Interpretive Center, Republic, Washington, 86 pp." type="book" year="2011">Pigg et al. 2011</bibRefCitation>
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), Green River Formation (
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<bibRefCitation id="A11C4B44EF598265FDE0F8286DC8F858" author="Brown, R. W." box="[620,760,1945,1969]" pageId="6" pageNumber="43" pagination="279 - 292" refId="ref33528" refString="Brown, R. W. (1929): Additions to the flora of the Green River Formation. - United States Geological Service Professional Paper, 154 - J: 279 - 292. https: // doi. org / 10.3133 / pp 154 j" type="book chapter" year="1929">Brown 1929</bibRefCitation>
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) and Oligocene Beaverhead flora of
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<taxonomicName id="028D4D36EF598265FD8BF80B6DCCF83B" authority="(Wing 1987)" baseAuthorityName="Wing" baseAuthorityYear="1987" box="[519,764,1978,2002]" class="Magnoliopsida" family="Eucommiaceae" genus="Eucommia" kingdom="Plantae" order="Garryales" pageId="6" pageNumber="43" phylum="Tracheophyta" rank="species" species="montana">
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Montana (
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<bibRefCitation id="A11C4B44EF598265FDF9F80B6DC3F83B" author="Wing, S. L." box="[629,755,1978,2002]" pageId="6" pageNumber="43" pagination="748 - 784" refId="ref44013" refString="Wing, S. L. (1987): Eocene and Oligocene floras of the Rocky Mountains. - Annals of the Missouri Botanical Garden, 74 (4): 748 - 784. https: // doi. org / 10.2307 / 2399449" type="journal article" year="1987">Wing 1987</bibRefCitation>
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)
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</taxonomicName>
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.
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<taxonomicName id="028D4D36EF598265FF0CF86B6F8BF81B" baseAuthorityName="Wolfe and Schorn" baseAuthorityYear="1990" box="[128,187,2010,2034]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="43" phylum="Tracheophyta" rank="genus">
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<emphasis id="F7F9EAA7EF598265FF0CF86B6F8BF81B" box="[128,187,2010,2034]" italics="true" pageId="6" pageNumber="43">Abies</emphasis>
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</taxonomicName>
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is also recognized based on seeds and foliage from the mid-Eocene Thunder Mountain flora of
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<collectingRegion id="0749F857EF598265FDD8F84A6DA1F7FA" box="[596,657,2043,2067]" country="United States of America" name="Idaho" pageId="6" pageNumber="43">Idaho</collectingRegion>
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(
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<bibRefCitation id="A11C4B44EF598265FD2AF84A6F89F7DD" author="Axelrod, D. I." pageId="6" pageNumber="43" pagination="1 - 193" refId="ref32761" refString="Axelrod, D. I. (1998): The Eocene thunder mountain flora of Central Idaho. - University of California Publications Geological Sciences, 142: 1 - 193." type="journal article" year="1998">Axelrod 1998</bibRefCitation>
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). Although the identity of some of the specimens that were attributed to
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<taxonomicName id="028D4D36EF598265FB94FDB76A0EFDF4" authority="AXELROD" authorityName="AXELROD" box="[1048,1342,518,542]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="43" phylum="Tracheophyta" rank="species" species="deweyensis">
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<emphasis id="F7F9EAA7EF598265FB94FDB76BE6FDF7" box="[1048,1238,518,542]" italics="true" pageId="6" pageNumber="43">Abies deweyensis</emphasis>
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<smallCapsWord id="C3D4A069EF598265FB53FDB76A0EFDF4" baselines="536,537" box="[1247,1342,518,542]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Axelrod" pageId="6" pageNumber="43">AXELROD</smallCapsWord>
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</taxonomicName>
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, by
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<bibRefCitation id="A11C4B44EF598265FAFEFDB76CA2FDD6" author="Axelrod, D. I." pageId="6" pageNumber="43" pagination="1 - 193" refId="ref32761" refString="Axelrod, D. I. (1998): The Eocene thunder mountain flora of Central Idaho. - University of California Publications Geological Sciences, 142: 1 - 193." type="journal article" year="1998">Axelrod (1998)</bibRefCitation>
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has been questioned due to disarticulated parts and a lack of diagnostic features (
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<bibRefCitation id="A11C4B44EF598265FB08FDF66AB7FDB6" author="Erwin, D. M. & Schorn, H. E." box="[1156,1415,583,607]" pageId="6" pageNumber="43" pagination="125 - 145" refId="ref35244" refString="Erwin, D. M., Schorn, H. E. (2005): Revisions of the conifers from the Eocene Thunder Mountain flora, Idaho, U. S. A. - Review of Palaeobotany and Palynology, 137: 125 - 145. https: // doi. org / 10.1016 / j. revpalbo. 2005.09.002" type="journal article" year="2005">Erwin and Schorn 2005</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |