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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1027.54888" ID-GBIF-Dataset="a108311a-65d1-431d-b108-40e585c2646b" ID-PMC="PMC8035127" ID-Pensoft-Pub="1313-2970-1027-1" ID-Pensoft-UUID="C5103AABD06058C595280B912CBE9B8B" ID-PubMed="33867800" ID-ZooBank="7D179ED7D7A540A2A972E07BC648F3B9" ModsDocID="1313-2970-1027-1" checkinTime="1617435236653" checkinUser="pensoft" docAuthor="Godwin, Rebecca L. &amp; Bond, Jason E." docDate="2021" docId="3AC7CFB2651C58B78265435A739CA573" docLanguage="en" docName="ZooKeys 1027: 1-165" docOrigin="ZooKeys 1027" docPubDate="2021-04-02" docSource="http://dx.doi.org/10.3897/zookeys.1027.54888" docTitle="Ummidia carlosviquezi Godwin &amp; Bond 2021, sp. nov." docType="treatment" docUuid="299C26B2-6D75-438E-B23D-03795ADF968E" docUuidSource="ZooBank" docVersion="4" id="C5103AABD06058C595280B912CBE9B8B" lastPageNumber="1" masterDocId="C5103AABD06058C595280B912CBE9B8B" masterDocTitle="Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae)" masterLastPageNumber="165" masterPageNumber="1" pageNumber="1" updateTime="1668151966284" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Godwin, Rebecca L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2439-2868</mods:nameIdentifier>
<mods:affiliation>Department of Entomology and Nematology, University of California, Davis 1 Shields Ave, Davis, CA, 95616</mods:affiliation>
<mods:nameIdentifier type="email">rlc0004@tigermail.auburn.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bond, Jason E.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of California, Davis 1 Shields Ave, Davis, CA, 95616</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1027</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>165</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1027.54888</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1027.54888</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1027-1</mods:identifier>
<mods:identifier type="ZooBank">7D179ED7D7A540A2A972E07BC648F3B9</mods:identifier>
<mods:identifier type="Pensoft-UUID">C5103AABD06058C595280B912CBE9B8B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="180850647" LSID="urn:lsid:zoobank.org:act:299C26B2-6D75-438E-B23D-03795ADF968E" httpUri="http://treatment.plazi.org/id/3AC7CFB2651C58B78265435A739CA573" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/299C26B2-6D75-438E-B23D-03795ADF968E" authority="Godwin &amp; Bond, 2021" authorityName="Godwin &amp; Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia carlosviquezi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="carlosviquezi" status="sp. nov.">Ummidia carlosviquezi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 57" captionStartId="F62" captionText="Figure 57. Ummidia carlosviquezi sp. nov. from Puntarenas, Costa Rica. A male habitus illustration UMM 239 B female habitus illustration UMM 747. Scale bars: 4.0 mm." figureDoi="10.3897/zookeys.1027.54888.figure57" httpUri="https://binary.pensoft.net/fig/527521" pageId="0" pageNumber="1">Figs 57</figureCitation>
<figureCitation captionStart="Figure 58" captionStartId="F63" captionText="Figure 58. Ummidia carlosviquezi sp. nov. from Puntarenas, Costa Rica A-D male holotype (UMM 239) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female paratype (UMM 747). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1027.54888.figure58" httpUri="https://binary.pensoft.net/fig/527522" pageId="0" pageNumber="1">, 58</figureCitation>
<figureCitation captionStart="Map 5" captionStartId="F46" captionText="Map 5. Locality records for southern Mexico and Central American species." figureDoi="10.3897/zookeys.1027.54888.map5" httpUri="https://binary.pensoft.net/fig/527503" pageId="0" pageNumber="1">, Map 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
HOLOTYPE: 1 ♂ (UMM239) from 6 kilometers south of San Vito, Puntarenas, Costa Rica,
<geoCoordinate degrees="8.7125" direction="north" orientation="latitude" precision="5" value="8.7125">8.7125</geoCoordinate>
<geoCoordinate degrees="83.0051" direction="west" orientation="longitude" precision="5" value="-83.0051">-83.0051</geoCoordinate>
1, 355 m a.s.l., 13-18.iii.1967, coll. C Valerio, AMNH. PARATYPE: 1 ♀ (UMM747) from Las Cruces, Puntarenas, Costa Rica,
<geoCoordinate degrees="8.7919" direction="north" orientation="latitude" precision="5" value="8.7919">8.7919</geoCoordinate>
<geoCoordinate degrees="82.9551" direction="west" orientation="longitude" precision="5" value="-82.9551">-82.9551</geoCoordinate>
1, 1227 m a.s.l. coll. C Viquez, P Jordan 7-8.xi.2001, BME.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The specific epithet is a patronym in honor of Carlos Viquez, a Costa Rican arachnologist who assisted the authors in the field and provided a great deal of the Costa Rican material used in this revision, to include one of the type specimens for this species.</paragraph>
<caption doi="10.3897/zookeys.1027.54888.figure57" httpUri="https://binary.pensoft.net/fig/527521" pageId="0" pageNumber="1" start="Figure 57" startId="F62">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 57.</emphasis>
<taxonomicName authorityName="Godwin &amp; Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia carlosviquezi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="carlosviquezi">
<emphasis italics="true" pageId="0" pageNumber="1">Ummidia carlosviquezi</emphasis>
</taxonomicName>
sp. nov. from Puntarenas, Costa Rica.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
male habitus illustration UMM239
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
female habitus illustration UMM747. Scale bars: 4.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Godwin &amp; Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia carlosviquezi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="carlosviquezi">
<emphasis italics="true" pageId="0" pageNumber="1">Ummidia carlosviquezi</emphasis>
</taxonomicName>
males can be distinguished from
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. hondurena" order="Araneae" pageId="0" pageNumber="1" rank="species" species="hondurena">
<emphasis italics="true" pageId="0" pageNumber="1">U. hondurena</emphasis>
</taxonomicName>
,
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. yojoa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="yojoa">
<emphasis italics="true" pageId="0" pageNumber="1">U. yojoa</emphasis>
</taxonomicName>
,
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. rugosa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="rugosa">
<emphasis italics="true" pageId="0" pageNumber="1">U. rugosa</emphasis>
</taxonomicName>
, and
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. erema" order="Araneae" pageId="0" pageNumber="1" rank="species" species="erema">
<emphasis italics="true" pageId="0" pageNumber="1">U. erema</emphasis>
</taxonomicName>
by having a relatively short embolus with a single even curve and from
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. yojoa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="yojoa">
<emphasis italics="true" pageId="0" pageNumber="1">U. yojoa</emphasis>
</taxonomicName>
and
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. varablanca" order="Araneae" pageId="0" pageNumber="1" rank="species" species="varablanca">
<emphasis italics="true" pageId="0" pageNumber="1">U. varablanca</emphasis>
</taxonomicName>
by the presence of many prolateral (22 vs 0) and retrolateral (35 vs 2-3) spines on tibia I. Males can be further differentiated from
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. rugosa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="rugosa">
<emphasis italics="true" pageId="0" pageNumber="1">U. rugosa</emphasis>
</taxonomicName>
and
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. riverai" order="Araneae" pageId="0" pageNumber="1" rank="species" species="riverai">
<emphasis italics="true" pageId="0" pageNumber="1">U. riverai</emphasis>
</taxonomicName>
by the lack of a pale dorsal heart patch on the abdomen and from
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. matagalpa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="matagalpa">
<emphasis italics="true" pageId="0" pageNumber="1">U. matagalpa</emphasis>
</taxonomicName>
and
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. quepoa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="quepoa">
<emphasis italics="true" pageId="0" pageNumber="1">U. quepoa</emphasis>
</taxonomicName>
by the presence of a comb on the retrolateral face of tarsus IV. Females can be distinguished from all geographically proximate species except
<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. cerrohoya" order="Araneae" pageId="0" pageNumber="1" rank="species" species="cerrohoya">
<emphasis italics="true" pageId="0" pageNumber="1">U. cerrohoya</emphasis>
</taxonomicName>
by having straight, simple spermathecae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description of male holotype.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Specimen preparation and condition</emphasis>
. Specimen preserved in 80% EtOH. Left palp and leg I, right leg IV removed, in vial with specimen.
<emphasis italics="true" pageId="0" pageNumber="1">General coloration</emphasis>
. Carapace, chelicerae, and legs reddish black 10R 2.5/1, tarsi yellowish red 5YR 4/6. Abdomen reddish black 2.5YR 2.5/1.
<emphasis italics="true" pageId="0" pageNumber="1">Cephalothorax</emphasis>
. Carapace 6.7 long, 6.18 wide. Pars cephalica 4.13 long. Foveal groove procurved, 0.4 long, 1.12 wide. All eyes on moderate tubercle. AER procurved. PER recurved. Eye group 0.85 long, 1.39 wide, AME 0.35, PME 0.25, ALE 0.48, PLE 0.36. Sternum sparsely setose around outer 1/3, thicker anteriorly, STRl 3.73, STRw 3.49. Chelicerae with anterior tooth row comprising six teeth, posterior margin with six teeth. Palpal endites with 21 cuspules spread over proximal half of endite face, and four cuspules distally, ENDw 1.43, ENDl 2.53. Labium with five cuspules, LBw 1.37, LBl 0.91. Rastellum with many spines on process. Abdomen setose.
<emphasis italics="true" pageId="0" pageNumber="1">Legs</emphasis>
. F1 5.96; F1w 1.58; P1 3.1; Ti1 4.34; Mt1 2.6; Tr1 1.42; F3 4.33; F3w 1.66; P3 2.16; Ti3 2.4; Sd3 1.29; Mt3 2.47; Tr3 1.85; F4 5.55; F4w 1.66; P4 2.46; Ti4 3.68; Mt4 3.8; Tr4 1.94. Retrolateral face of tarsus IV with defined comb over length of tarsus. Leg I spination pattern: TSp 10, TSpv 11, TSrd 0, TSr 1, TSrv 34, MtSp 11, MtSr 15, TrSp 6, TrSr 8.
<emphasis italics="true" pageId="0" pageNumber="1">Pedipalps</emphasis>
. PTl 3.45, PTw 1.19, Bl 2.02. Embolus relatively short with even curve.
</paragraph>
<caption doi="10.3897/zookeys.1027.54888.figure58" httpUri="https://binary.pensoft.net/fig/527522" pageId="0" pageNumber="1" start="Figure 58" startId="F63">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 58.</emphasis>
<taxonomicName authorityName="Godwin &amp; Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia carlosviquezi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="carlosviquezi">
<emphasis italics="true" pageId="0" pageNumber="1">Ummidia carlosviquezi</emphasis>
</taxonomicName>
sp. nov. from Puntarenas, Costa Rica
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
male holotype (UMM239)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
prolateral aspect, leg I
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
retrolateral aspect, leg I
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
line drawings, leg I prolateral and retrolateral aspects
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
retrolateral aspect, pedipalp
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
cleared spermathecae female paratype (UMM747). Scale bars: 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation, males</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">(n = 14).</emphasis>
CL 5.17-7.82, 6.51
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21; CW 4.69-7.41, 6.05
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21; Cap 3.4-5.24, 4.16
<normalizedToken originalValue="±">+/-</normalizedToken>
0.15; ENDl 0.66-0.93, 0.79
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02; ENDw 1.26-1.99, 1.45
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05; STRl 2.93-4.43, 3.63
<normalizedToken originalValue="±">+/-</normalizedToken>
0.12; STRw 2.65-4.19, 3.28
<normalizedToken originalValue="±">+/-</normalizedToken>
0.13; LBl 0.65-1.12, 0.89
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04; LBw 1.02-1.61, 1.29
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05; F1 4.07-7.19, 5.78
<normalizedToken originalValue="±">+/-</normalizedToken>
0.22; F1w 1.14-1.8, 1.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05; P1 2.15-3.39, 2.82
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1; Ti1 2.58-4.88, 3.85
<normalizedToken originalValue="±">+/-</normalizedToken>
0.17; Mt1 1.78-3.02, 2.45
<normalizedToken originalValue="±">+/-</normalizedToken>
0.09; Tr1 1-1.42, 1.24
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03; F3 3.15-5.06, 4.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.15; F3w 1.37-2.11, 1.63
<normalizedToken originalValue="±">+/-</normalizedToken>
0.06; P3 1.62-2.61, 2.16
<normalizedToken originalValue="±">+/-</normalizedToken>
0.08; Ti3 1.77-2.97, 2.42
<normalizedToken originalValue="±">+/-</normalizedToken>
0.09; Mt3 1.5-2381, 172.29
<normalizedToken originalValue="±">+/-</normalizedToken>
169.9; Tr3 1.24-2.02, 1.75
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05; F4 4.11-6.93, 5.63
<normalizedToken originalValue="±">+/-</normalizedToken>
0.22; F4w 1.27-1.99, 1.54
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05; P4 2-2.96, 2.44
<normalizedToken originalValue="±">+/-</normalizedToken>
0.08; Ti4 2.62-4.83, 3.67
<normalizedToken originalValue="±">+/-</normalizedToken>
0.16; Mt4 2.61-5.07, 3.95
<normalizedToken originalValue="±">+/-</normalizedToken>
0.18; Tr4 1.49-2.29, 1.86
<normalizedToken originalValue="±">+/-</normalizedToken>
0.06; TSp 0-13, 3.64
<normalizedToken originalValue="±">+/-</normalizedToken>
1.13; TSpv 0-26, 8.43
<normalizedToken originalValue="±">+/-</normalizedToken>
1.92; TSr 0-11, 1.36
<normalizedToken originalValue="±">+/-</normalizedToken>
0.75; TSrv 9-38, 25.86
<normalizedToken originalValue="±">+/-</normalizedToken>
2.13; PTl 2.2-4.07, 3.37
<normalizedToken originalValue="±">+/-</normalizedToken>
0.15; PTw 0.85-1.35, 1.17
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04; BL 1.59-2.34, 1.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description of female paratype.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Specimen preparation and condition</emphasis>
. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen.
<emphasis italics="true" pageId="0" pageNumber="1">General coloration</emphasis>
. Carapace, chelicerae, and legs dark reddish brown 5YR 3/4. Abdomen black 7.5YR 2.5/1, spinnerets dark yellowish brown 10YR 4/6.
<emphasis italics="true" pageId="0" pageNumber="1">Cephalothorax</emphasis>
. Carapace 8.71 long, 7.08 wide. Pars cephalica 5.46 long. Foveal groove procurved, 0.79 long, 1.63 wide. Eye tubercle moderate under ME. AER procurved. PER straight. Eye group 0.79 long, 1.63 wide, AME 0.44, PME 0.38, ALE 0.62, PLE 0.39. Sternum sparsely setose around edges, thicker anteriorly STRl 4.99, STRw 4.58. Chelicerae with anterior row comprising six teeth, posterior margin with seven teeth. Palpal endites with 30 cuspules spread across proximal half of endite and 56 smaller cuspules distally, ENDw 1.81, ENDl 3.09. Labium with eight cuspules, LBw 1.76, LBl 1.41. Rastellum with strong spines along margin and bottom half of process with smaller spines on upper half of process and up cheliceral face ~ 1.5
<normalizedToken originalValue="×">x</normalizedToken>
length of process.
<emphasis italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
. Evenly setose with pale speckles clustered at apodemes.
<emphasis italics="true" pageId="0" pageNumber="1">Legs</emphasis>
. F1 5.37; F1w 1.89; P1 3.25; Ti1 3.25; Mt1 2.1; Tr1 1.25; F3 4.78; F3w 2.14; P3 2.74; Ti3 2.47, Sd3 1.51; Mt3 2.04; Tr3 1.83; F4 5.68; F4w 2.45; P4 2.77; Ti4 3.49; Mt4 3.27; Tr4 1.96. Retrolateral face tarsus IV with distinct comb over central half of tarsus becoming loose distally.
<emphasis italics="true" pageId="0" pageNumber="1">Pedipalps</emphasis>
. PF 4.85, PP 2.84, PTi 3.11, PTr 2.7. Spermathecae simple and straight, bulbs facing anteriorly.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation, females</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">(n = 8).</emphasis>
CL 4.2-9.79, 7.03
<normalizedToken originalValue="±">+/-</normalizedToken>
0.72; CW 3.86-9.13, 6.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.63; Cap 2.87-6.75, 4.73
<normalizedToken originalValue="±">+/-</normalizedToken>
0.47; ENDl 0.58-1.21, 0.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.08; ENDw 1.09-2.05, 1.56
<normalizedToken originalValue="±">+/-</normalizedToken>
0.11; STRl 2.34-5.98, 4.19
<normalizedToken originalValue="±">+/-</normalizedToken>
0.43; STRw 2.45-5.69, 4.01
<normalizedToken originalValue="±">+/-</normalizedToken>
0.38; LBl 0.72-1.63, 1.12
<normalizedToken originalValue="±">+/-</normalizedToken>
0.11; LBw 0.91-2.11, 1.49
<normalizedToken originalValue="±">+/-</normalizedToken>
0.14; F1 2.61-6.32, 4.39
<normalizedToken originalValue="±">+/-</normalizedToken>
0.49; F1w 1.02-2.11, 1.55
<normalizedToken originalValue="±">+/-</normalizedToken>
0.15; P1 1.57-3.85, 2.72
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3; Ti1 1.43-3.9, 2.61
<normalizedToken originalValue="±">+/-</normalizedToken>
0.32; Mt1 1.01-2.69, 1.75
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21; Tr1 0.85-1.58, 1.16
<normalizedToken originalValue="±">+/-</normalizedToken>
0.09; F3 2.24-5.15, 3.79
<normalizedToken originalValue="±">+/-</normalizedToken>
0.38; F3w 1.21-2.65, 1.95
<normalizedToken originalValue="±">+/-</normalizedToken>
0.18; P3 1.32-3.38, 2.28
<normalizedToken originalValue="±">+/-</normalizedToken>
0.25; Ti3 1.21-3, 2.08
<normalizedToken originalValue="±">+/-</normalizedToken>
0.22; Mt3 0.98-2.28, 1.67
<normalizedToken originalValue="±">+/-</normalizedToken>
0.17; Tr3 0.84-2.37, 1.64
<normalizedToken originalValue="±">+/-</normalizedToken>
0.18; F4 2.94-6.58, 4.69
<normalizedToken originalValue="±">+/-</normalizedToken>
0.43; F4w 1.22-6.47, 2.46
<normalizedToken originalValue="±">+/-</normalizedToken>
0.59; P4 1.52-3.46, 2.34
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21; Ti4 1.63-3.78, 2.76
<normalizedToken originalValue="±">+/-</normalizedToken>
0.28; Mt4 1.63-3.97, 2.72
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3; Tr4 1.13-3.55, 1.78
<normalizedToken originalValue="±">+/-</normalizedToken>
0.28; PF 2.26-5.84, 4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.46; PP 1.35-3.27, 2.33
<normalizedToken originalValue="±">+/-</normalizedToken>
0.25; PTi 1.39-3.76, 2.51
<normalizedToken originalValue="±">+/-</normalizedToken>
0.32; PTr 1.16-3.39, 2.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.28.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Costa Rica: Cartago</emphasis>
:
<normalizedToken originalValue="Cantón-Central">Canton-Central</normalizedToken>
,
<geoCoordinate degrees="9.7156" direction="north" orientation="latitude" precision="5" value="9.7156">9.7156</geoCoordinate>
<geoCoordinate degrees="83.7503" direction="west" orientation="longitude" precision="5" value="-83.7503">-83.7503</geoCoordinate>
9, 1743 m a.s.l. (UMM0587, 20.iv.1973, 1♂, Araya, Rodolfo, AMNH);
<emphasis bold="true" pageId="0" pageNumber="1">Guanacaste</emphasis>
:
<normalizedToken originalValue="Estación">Estacion</normalizedToken>
Cacao,
<geoCoordinate degrees="10.9291" direction="north" orientation="latitude" precision="5" value="10.9291">10.9291</geoCoordinate>
<geoCoordinate degrees="85.4952" direction="west" orientation="longitude" precision="5" value="-85.4952">-85.4952</geoCoordinate>
1, 1100 m a.s.l. (UMM0759b, vii.1989-iii.1990, 1♂, C Moraga, BME); Guanacaste Conservation Area, La Cruz, Guanacaste National Park, Maritza Station,
<geoCoordinate degrees="10.9625" direction="north" orientation="latitude" precision="5" value="10.9625">10.9625</geoCoordinate>
<geoCoordinate degrees="85.4952" direction="west" orientation="longitude" precision="5" value="-85.4952">-85.4952</geoCoordinate>
1, 600 m a.s.l. (UMM0761, 1-31.v.1996, 1♂, M Pereira, BME); La Cruz, 9 km S Santa Cecilia, Pitilla Station,
<geoCoordinate degrees="10.9904" direction="north" orientation="latitude" precision="5" value="10.9904">10.9904</geoCoordinate>
<geoCoordinate degrees="85.4272" direction="west" orientation="longitude" precision="5" value="-85.4272">-85.4272</geoCoordinate>
1, 700 m a.s.l. (UMM0760, 1-31.x.1996, 1♂, C Moraga, BME); Tenorio National Park,
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
Tenorio,
<geoCoordinate degrees="10.6764" direction="north" orientation="latitude" precision="5" value="10.6764">10.6764</geoCoordinate>
<geoCoordinate degrees="85.0225" direction="west" orientation="longitude" precision="5" value="-85.0225">-85.0225</geoCoordinate>
1, 1661 m a.s.l. (UMM0756, 4.xi.2003-15.ii.2004, 1♂, A Azofeifa, BME);
<emphasis bold="true" pageId="0" pageNumber="1">Heredia</emphasis>
: Santa Rosa, Cacao, 9.9705 -84.1 1, 1114 m a.s.l. (UMM0671, 1♂, BME);
<emphasis bold="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Limón">Limon</normalizedToken>
</emphasis>
: Barbilla,
<geoCoordinate degrees="10.0695" direction="north" orientation="latitude" precision="5" value="10.0695">10.0695</geoCoordinate>
<geoCoordinate degrees="83.3675" direction="west" orientation="longitude" precision="5" value="-83.3675">-83.3675</geoCoordinate>
1, 22 m a.s.l. (UMM0753, 20-24.x.2000, 1♀, Mantillo, BME); Hitoy Cerere,
<geoCoordinate degrees="9.6436" direction="north" orientation="latitude" precision="5" value="9.6436">9.6436</geoCoordinate>
<geoCoordinate degrees="83.0711" direction="west" orientation="longitude" precision="5" value="-83.0711">-83.0711</geoCoordinate>
1, 364 m a.s.l. (UMM0679, 1♂, BME); R.B. Hitoy Cerere. Send. Espavel,
<geoCoordinate degrees="9.6614" direction="north" orientation="latitude" precision="5" value="9.6614">9.6614</geoCoordinate>
<geoCoordinate degrees="83.0307" direction="west" orientation="longitude" precision="5" value="-83.0307">-83.0307</geoCoordinate>
1, 560 m a.s.l. (UMM0763, 18.ix-3.x.2003, 1♂, W Arana, B Gamboa, E Rojas, BME);
<normalizedToken originalValue="Río">Rio</normalizedToken>
Yorkin and
<normalizedToken originalValue="Río">Rio</normalizedToken>
Telire,
<geoCoordinate degrees="9.5599" direction="north" orientation="latitude" precision="5" value="9.5599">9.5599</geoCoordinate>
<geoCoordinate degrees="82.924" direction="west" orientation="longitude" precision="55" value="-82.924">-82.924</geoCoordinate>
1, 1300 m a.s.l. (UMM0677, 24-26.x.2008, 1♂, A Solis, BME);
<emphasis bold="true" pageId="0" pageNumber="1">Puntarenas</emphasis>
: Platanares,
<geoCoordinate degrees="9.138" direction="north" orientation="latitude" precision="55" value="9.138">9.138</geoCoordinate>
<geoCoordinate degrees="83.2777" direction="west" orientation="longitude" precision="5" value="-83.2777">-83.2777</geoCoordinate>
1, 413 m a.s.l. (UMM0678, 16.vii-4.viii.2001, 1♂, R
<normalizedToken originalValue="Gonzálz">Gonzalz</normalizedToken>
, BME); 6km S San Vito,
<geoCoordinate degrees="8.7125" direction="north" orientation="latitude" precision="5" value="8.7125">8.7125</geoCoordinate>
<geoCoordinate degrees="83.0051" direction="west" orientation="longitude" precision="5" value="-83.0051">-83.0051</geoCoordinate>
1, 355 m a.s.l. (UMM0239, 13-18.iii.1967, 1♂, AMNH); Gulfito,
<geoCoordinate degrees="8.6391" direction="north" orientation="latitude" precision="5" value="8.6391">8.6391</geoCoordinate>
<geoCoordinate degrees="83.1625" direction="west" orientation="longitude" precision="5" value="-83.1625">-83.1625</geoCoordinate>
5, 112 m a.s.l. (UMM0120, iv.1972, 1♀, ICECU, AMNH); Las Cruces,
<geoCoordinate degrees="8.7919" direction="north" orientation="latitude" precision="5" value="8.7919">8.7919</geoCoordinate>
<geoCoordinate degrees="82.9551" direction="west" orientation="longitude" precision="5" value="-82.9551">-82.9551</geoCoordinate>
1, 1227 m a.s.l. (UMM0747, 1♀, BME);
<emphasis bold="true" pageId="0" pageNumber="1">
San
<normalizedToken originalValue="José">Jose</normalizedToken>
</emphasis>
: (UMM0624, 5.iv.1980, 1♀, AMNH); Grandilla,
<geoCoordinate degrees="9.9295" direction="north" orientation="latitude" precision="5" value="9.9295">9.9295</geoCoordinate>
<geoCoordinate degrees="84.0193" direction="west" orientation="longitude" precision="5" value="-84.0193">-84.0193</geoCoordinate>
5, 1290 m a.s.l. (UMM0545, 10.vi.1986, 1♀, CE Valerio, AMNH); BME building,
<geoCoordinate degrees="9.973" direction="north" orientation="latitude" precision="55" value="9.973">9.973</geoCoordinate>
<geoCoordinate degrees="84.0937" direction="west" orientation="longitude" precision="5" value="-84.0937">-84.0937</geoCoordinate>
1, 1127 m a.s.l. (UMM0672, 4.v.2001, 1♂, A Solis, BME); La Cangreja National Park, Ecotropica Station,
<geoCoordinate degrees="9.6883" direction="north" orientation="latitude" precision="5" value="9.6883">9.6883</geoCoordinate>
<geoCoordinate degrees="84.3692" direction="west" orientation="longitude" precision="5" value="-84.3692">-84.3692</geoCoordinate>
1, 300-400 m a.s.l. (UMM0758, 13-17.vii.2004, 1♂, Porras,
<normalizedToken originalValue="Cárdenas">Cardenas</normalizedToken>
, Gamboa, Briceno, Moraga, Mata, BME); La Congreja National Park,
<geoCoordinate degrees="9.7032" direction="north" orientation="latitude" precision="5" value="9.7032">9.7032</geoCoordinate>
<geoCoordinate degrees="84.3972" direction="west" orientation="longitude" precision="5" value="-84.3972">-84.3972</geoCoordinate>
1, 448 m a.s.l. (UMM0748, 1♀, BME); La Fonda,
<geoCoordinate degrees="10.0495" direction="north" orientation="latitude" precision="5" value="10.0495">10.0495</geoCoordinate>
<geoCoordinate degrees="84.0077" direction="west" orientation="longitude" precision="5" value="-84.0077">-84.0077</geoCoordinate>
1, 1606 m a.s.l. (UMM0745, 1♀, BME); San
<normalizedToken originalValue="José">Jose</normalizedToken>
,
<geoCoordinate degrees="9.933" direction="north" orientation="latitude" precision="55" value="9.933">9.933</geoCoordinate>
<geoCoordinate degrees="84.0831" direction="west" orientation="longitude" precision="5" value="-84.0831">-84.0831</geoCoordinate>
6, 1156 m a.s.l. (UMM0203, 1♀, MCZ); Zurqui de Moravia,
<geoCoordinate degrees="10.0495" direction="north" orientation="latitude" precision="5" value="10.0495">10.0495</geoCoordinate>
<geoCoordinate degrees="84.0159" direction="west" orientation="longitude" precision="5" value="-84.0159">-84.0159</geoCoordinate>
1, 1600 m a.s.l. (UMM0762, 22.vii-8.viii.2010, 1♂, W Porras, BME);
<emphasis bold="true" pageId="0" pageNumber="1">Nicaragua: Granada</emphasis>
:
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
Mombacho, non-organic coffee plantation,
<geoCoordinate degrees="12.4219" direction="north" orientation="latitude" precision="5" value="12.4219">12.4219</geoCoordinate>
<geoCoordinate degrees="86.5448" direction="west" orientation="longitude" precision="5" value="-86.5448">-86.5448</geoCoordinate>
1, 700-800 m a.s.l. (UMM0685, 2.vi.1998, 1♂, JM Maes, BME).
</paragraph>
</subSubSection>
</treatment>
</document>