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<document id="6E9E2A235BAB519402C9629DD3F5CC12" ID-CLB-Dataset="189256" ID-DOI="10.11646/phytotaxa.573.1.1" ID-GBIF-Dataset="55e0a16e-c362-41d5-8bdc-855a7ffb0d63" ID-ISSN="1179-3163" ID-Zenodo-Dep="7329401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668667493236" checkinUser="plazi" docAuthor="Nilanthi, Rajapakse Mudiyanselage Renuka, Samarakoon, Hiruna, Jayawardana, Nuwan, Hathurusinghe, Bhagya, Wijesundara, Siril &amp; Bandaranayake, Pradeepa Chandani Gunathilake" docDate="2022" docId="226337481075433FFF6A84F5EA292E58" docLanguage="en" docName="phytotaxa.573.1.1.pdf" docOrigin="Phytotaxa 573 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.573.1.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Strobilanthes glandulata Nilanthi 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="DE5A4F3010734334FFE28401E8382F7C" masterDocTitle="Strobilanthes glandulata (Acanthaceae), a new species from Sri Lanka based on the morphological and molecular evidences" masterLastPageNumber="14" masterPageNumber="1" pageNumber="7" updateTime="1699411605016" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6E601D0AB98802FCC567FC6E24D1598C">Strobilanthes glandulata (Acanthaceae), a new species from Sri Lanka based on the morphological and molecular evidences</mods:title>
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<mods:namePart id="BB8520F3B4E734861B9D5A2A90411991">Nilanthi, Rajapakse Mudiyanselage Renuka</mods:namePart>
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<mods:affiliation id="CEEEBAF8838945F07F6827664D453716">Department of Wildlife Conservation, Battaramulla 10120, Sri Lanka &amp; Post Graduate Institute of Science, University of Peradeniya, Peradeniya 20400, Sri Lanka &amp; nilanthi. dwc @ gmail. com; https: // orcid. org / 0000 - 0002 - 8398 - 541 X</mods:affiliation>
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<mods:namePart id="D0C4E90FCF34EE4EC5FA9C6A0AD5ED2D">Samarakoon, Hiruna</mods:namePart>
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<mods:affiliation id="9CB6CB0074A6D09731B84DCC1F4BA6B4">Agriculture Biotechnology Centre, Faculty of Agriculture, University of Peradeniya, Peradeniya 20400, Sri Lanka &amp; hirunas @ eng. pdn. ac. lk; https: // orcid. org / 0000 - 0002 - 5812 - 1046</mods:affiliation>
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<mods:namePart id="4F6DA4C5DE235F8E6A917261C21BC611">Jayawardana, Nuwan</mods:namePart>
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<mods:affiliation id="C3F5F24477CCE0E847AA4609359E70F2">Department of Wildlife Conservation, Battaramulla 10120, Sri Lanka &amp; chathuranga. nuwan. ka @ gmail. com; https: // orcid. org / 0000 - 0002 - 8273 - 3050</mods:affiliation>
<mods:nameIdentifier id="27F83326BDC4CACFB57D1D0616E9BEF7" type="email">chathuranga.nuwan.ka@gmail.com</mods:nameIdentifier>
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<mods:namePart id="40267FA23EA2B9DEA423E26FC17C04EB">Hathurusinghe, Bhagya</mods:namePart>
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<mods:affiliation id="8A96DBB503E2F62943B59DD7AE4FB1DD">Agriculture Biotechnology Centre, Faculty of Agriculture, University of Peradeniya, Peradeniya 20400, Sri Lanka &amp; bhagya. hathurusinghe 93 @ gmail. com; https: // orcid. org / 0000 - 0002 - 0816 - 2501</mods:affiliation>
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<mods:namePart id="FDDCDA3DD6A358875DB33146701BF719">Wijesundara, Siril</mods:namePart>
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<mods:affiliation id="1E113D514B758EB247BB43E29102C997">National Institute of Fundamental Studies, Hantane Road, Kandy 20000, Sri Lanka &amp; siril. wi @ nifs. ac. lk; https: // orcid. org / 0000 - 0002 - 6754 - 8201</mods:affiliation>
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<mods:namePart id="3DD33D7B88378733A52CC710EF4947B7">Bandaranayake, Pradeepa Chandani Gunathilake</mods:namePart>
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<mods:affiliation id="00D031F0C46826AA7CCCDB5642E83ECD">Agriculture Biotechnology Centre, Faculty of Agriculture, University of Peradeniya, Peradeniya 20400, Sri Lanka &amp; pradeepag @ agri. pdn. ac. lk, pgunathilake @ ucdavis. edu; https: // orcid. org / 0000 - 0001 - 8241 - 1775</mods:affiliation>
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<mods:title id="A7D15FE056A0ADDD958A2445CF15F72B">Phytotaxa</mods:title>
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<mods:date id="6ABB65C6264DADCEA1A6B71C3BC102B3">2022</mods:date>
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<mods:number id="8FA4919505B3FF8EC1A4FB8FF6638B2B">573</mods:number>
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<mods:classification id="0C6739F29EC492801FEBA65DB87535E3">journal article</mods:classification>
<mods:identifier id="43E9C87A5BE45345306A84598E60CF9C" type="CLB-Dataset">189256</mods:identifier>
<mods:identifier id="294DB8BD9168808574D9D5ACB6D72B9E" type="DOI">10.11646/phytotaxa.573.1.1</mods:identifier>
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<treatment id="226337481075433FFF6A84F5EA292E58" ID-DOI="http://doi.org/10.5281/zenodo.7334281" ID-GBIF-Taxon="205097852" ID-Zenodo-Dep="7334281" LSID="urn:lsid:plazi:treatment:226337481075433FFF6A84F5EA292E58" httpUri="http://treatment.plazi.org/id/226337481075433FFF6A84F5EA292E58" lastPageId="11" lastPageNumber="11" pageId="6" pageNumber="7">
<subSubSection id="E2D0D5D510754332FF6A84F5EB272E72" box="[136,799,244,271]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="AA75865E10754332FF6A84F5EB272E72" blockId="6.[136,799,244,271]" box="[136,799,244,271]" pageId="6" pageNumber="7">
<heading id="F13D313210754332FF6A84F5EB272E72" box="[136,799,244,271]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="3">
<taxonomicName id="6DCAFDDD10754332FF6A84F5EA342E72" ID-CoL="B2X4D" authority="Nilanthi" authorityName="Nilanthi" authorityYear="2022" box="[136,524,244,271]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="glandulata" status="sp. nov.">
<emphasis id="98BE5A4C10754332FF6A84F5E99D2E73" bold="true" box="[136,421,244,271]" italics="true" pageId="6" pageNumber="7">Strobilanthes glandulata</emphasis>
Nilanthi
</taxonomicName>
,
<emphasis id="98BE5A4C10754332FDF484F4EA532E72" box="[534,619,245,270]" italics="true" pageId="6" pageNumber="7">
<taxonomicNameLabel id="838DE73710754332FDF484F4EA532E72" box="[534,619,245,270]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Figures 5 &amp; 7)
</heading>
</paragraph>
</subSubSection>
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<paragraph id="AA75865E10754332FF6A851DECF42E4D" blockId="6.[136,1228,284,306]" box="[136,1228,284,306]" pageId="6" pageNumber="7">
<emphasis id="98BE5A4C10754332FF6A851DE8E22E4D" bold="true" box="[136,218,284,305]" pageId="6" pageNumber="7">Type:—</emphasis>
<materialsCitation id="1AA28C0310754332FF38851DECFF2E4D" ID-GBIF-Occurrence="3967649301" box="[218,1223,284,306]" collectingDate="1928-02-14" country="Sri Lanka" location="Madugoda" municipality="Kandy District" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Central" typeStatus="holotype">
<collectingCountry id="D2DDC6CE10754332FF38851DE9652E4D" box="[218,349,284,305]" name="Sri Lanka" pageId="6" pageNumber="7">SRI LANKA</collectingCountry>
.
<collectingRegion id="680E48BC10754332FE84851DEA322E4E" box="[358,522,284,306]" country="Sri Lanka" name="Central" pageId="6" pageNumber="7">Central province</collectingRegion>
:
<collectingMunicipality id="4A111C2410754332FDFB851DEA922E4E" box="[537,682,284,306]" pageId="6" pageNumber="7">Kandy District</collectingMunicipality>
,
<location id="AF15D08510754332FD51851DEB242E4E" LSID="urn:lsid:plazi:treatment:226337481075433FFF6A84F5EA292E58:AF15D08510754332FD51851DEB242E4E" box="[691,796,284,306]" country="Sri Lanka" municipality="Kandy District" name="Madugoda" pageId="6" pageNumber="7" stateProvince="Central">Madugoda</location>
,
<date id="DE74A09E10754332FCC4851DEB982E4D" box="[806,928,284,306]" pageId="6" pageNumber="7" value="1928-02-14">
<collectingDate id="CE30597610754332FCC4851DEB982E4D" box="[806,928,284,306]" pageId="6" pageNumber="7" value="1928-02-14">14 Feb 1928</collectingDate>
</date>
,
<emphasis id="98BE5A4C10754332FC48851DEC252E4D" box="[938,1053,284,305]" italics="true" pageId="6" pageNumber="7">Alston 1966</emphasis>
(
<typeStatus id="757138FC10754332FBCB851DEC472E4E" box="[1065,1151,284,306]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
PDA!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="FEB5D6D610754332FF6A83EBEA2C275F" ID-DOI="http://doi.org/10.5281/zenodo.7334289" ID-Zenodo-Dep="7334289" httpUri="https://zenodo.org/record/7334289/files/figure.png" pageId="6" pageNumber="7" startId="6.[136,229,2026,2048]" targetBox="[216,1372,364,1998]" targetPageId="6">
<paragraph id="AA75865E10754332FF6A83EBEA2C275F" blockId="6.[136,1452,2026,2084]" pageId="6" pageNumber="7">
<emphasis id="98BE5A4C10754332FF6A83EBE8DD277C" bold="true" box="[136,229,2026,2048]" pageId="6" pageNumber="7">FIGURE</emphasis>
.
<emphasis id="98BE5A4C10754332FEEA83EBE9C42883" box="[264,508,2026,2047]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="6DCAFDDD10754332FEEA83EBE9C02883" box="[264,504,2026,2047]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="glandulata">Strobilanthes glandulata</taxonomicName>
.
</emphasis>
<collectionCode id="CCDB1E9B10754332FDE383EBEA2D2883" box="[513,533,2026,2047]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="7" type="Herbarium">A</collectionCode>
. Habit; B. Flowering branch;
<collectionCode id="CCDB1E9B10754332FCD383EBEB7C2883" box="[817,836,2026,2047]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="7" type="Herbarium">C</collectionCode>
. Inflorescence; D &amp;
<collectionCode id="CCDB1E9B10754332FBEB83EBEC2F2883" box="[1033,1047,2026,2047]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="6" pageNumber="7" type="Herbarium">F</collectionCode>
. Leaves adaxial and abaxial view; E. front view of corolla; G. Internode; H. Bracts.
</paragraph>
</caption>
<caption id="FEB5D6D610744333FF6A83A0ED7228A7" ID-DOI="http://doi.org/10.5281/zenodo.7334291" ID-Zenodo-Dep="7334291" httpUri="https://zenodo.org/record/7334291/files/figure.png" pageId="7" pageNumber="8" startId="7.[136,229,1953,1975]" targetBox="[179,1408,190,1929]" targetPageId="7">
<paragraph id="AA75865E10744333FF6A83A0ED7228A7" blockId="7.[136,1451,1953,2011]" pageId="7" pageNumber="8">
<emphasis id="98BE5A4C10744333FF6A83A0E8DD28CB" bold="true" box="[136,229,1953,1975]" pageId="7" pageNumber="8">FIGURE</emphasis>
.
<emphasis id="98BE5A4C10744333FEE883A0E9DE28CA" box="[266,486,1953,1974]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="6DCAFDDD10744333FEE883A0E9DA28CA" box="[266,482,1953,1974]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="lupulina">Strobilanthes lupulina</taxonomicName>
.
</emphasis>
<collectionCode id="CCDB1E9B10744333FE0983A0E9C728CA" box="[491,511,1953,1974]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="7" pageNumber="8" type="Herbarium">A</collectionCode>
. Flowering branch; B–
<collectionCode id="CCDB1E9B10744333FD3983A0EAD328CA" box="[731,747,1953,1974]" country="Denmark" name="University of Copenhagen" pageId="7" pageNumber="8" type="Herbarium">C</collectionCode>
Inflorescence; D. Leaves adaxial and abaxial view; E. Bracts;
<collectionCode id="CCDB1E9B10744333FAAE83A0ED6228CA" box="[1356,1370,1953,1974]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="7" pageNumber="8" type="Herbarium">F</collectionCode>
. Corolla split open showing the stamens; G. front view of corolla; H. Close up of the leaf surface; I. Stamens; J. Internode; K. Capsule.
</paragraph>
</caption>
<caption id="FEB5D6D6107B433CFF6A83AAE998289A" ID-DOI="http://doi.org/10.5281/zenodo.7334293" ID-Zenodo-Dep="7334293" httpUri="https://zenodo.org/record/7334293/files/figure.png" pageId="8" pageNumber="9" startId="8.[136,229,1963,1985]" targetBox="[159,1428,210,1935]" targetPageId="8">
<paragraph id="AA75865E107B433CFF6A83AAE998289A" blockId="8.[136,1452,1963,2022]" pageId="8" pageNumber="9">
<emphasis id="98BE5A4C107B433CFF6A83AAE8DD28BD" bold="true" box="[136,229,1963,1985]" pageId="8" pageNumber="9">FIGURE</emphasis>
.
<taxonomicName id="6DCAFDDD107B433CFEEA83ADE9CF28BD" box="[264,503,1964,1985]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="glandulata">
<emphasis id="98BE5A4C107B433CFEEA83ADE9CF28BD" box="[264,503,1964,1985]" italics="true" pageId="8" pageNumber="9">Strobilanthes glandulata</emphasis>
</taxonomicName>
.
<collectionCode id="CCDB1E9B107B433CFDE183ADEA2F28BD" box="[515,535,1964,1985]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="8" pageNumber="9" type="Herbarium">A</collectionCode>
. Corolla split open showing the stamens; B. Inflorescence;
<collectionCode id="CCDB1E9B107B433CFB8683ADEC4F28BD" box="[1124,1143,1964,1985]" country="Denmark" name="University of Copenhagen" pageId="8" pageNumber="9" type="Herbarium">C</collectionCode>
. Leaf; D. Flowering branch; E. Close up of the leaf surfaces.
</paragraph>
</caption>
<caption id="FEB5D6D6107A433DFF6A83D4EAF22773" ID-DOI="http://doi.org/10.5281/zenodo.7334295" ID-Zenodo-Dep="7334295" httpUri="https://zenodo.org/record/7334295/files/figure.png" pageId="9" pageNumber="10" startId="9.[136,229,2005,2027]" targetBox="[155,1398,198,1980]" targetPageId="9">
<paragraph id="AA75865E107A433DFF6A83D4EAF22773" blockId="9.[136,1452,2005,2063]" pageId="9" pageNumber="10">
<emphasis id="98BE5A4C107A433DFF6A83D4E8DD2897" bold="true" box="[136,229,2005,2027]" pageId="9" pageNumber="10">FIGURE</emphasis>
.
<taxonomicName id="6DCAFDDD107A433DFEE783D4E9E42896" box="[261,476,2005,2026]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="lupulina">
<emphasis id="98BE5A4C107A433DFEE783D4E9E42896" box="[261,476,2005,2026]" italics="true" pageId="9" pageNumber="10">Strobilanthes lupulina</emphasis>
</taxonomicName>
.
<collectionCode id="CCDB1E9B107A433DFE0583D4E9C32896" box="[487,507,2005,2026]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="9" pageNumber="10" type="Herbarium">A</collectionCode>
. Capsule; B. Calyx;
<collectionCode id="CCDB1E9B107A433DFD2E83D4EAE72896" box="[716,735,2005,2026]" country="Denmark" name="University of Copenhagen" pageId="9" pageNumber="10" type="Herbarium">C</collectionCode>
. Bracts; D. Corolla split open showing the stamens; E. Inflorescences;
<collectionCode id="CCDB1E9B107A433DFA7883D7ED902897" box="[1434,1448,2006,2027]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="9" pageNumber="10" type="Herbarium">F</collectionCode>
. Flowering branch; G. Close up of the leaf surfaces; H. Leaf.
</paragraph>
</caption>
<subSubSection id="E2D0D5D51079433EFF6A849FEBF92E58" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="AA75865E1079433EFF6A849FEBF92E58" blockId="10.[136,1452,158,868]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF6A849FE9192FC4" bold="true" box="[136,289,158,184]" pageId="10" pageNumber="11">Diagnosis:—</emphasis>
<taxonomicName id="6DCAFDDD1079433EFEC3849FEA0F2FC4" box="[289,567,158,184]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="glandulata">
<emphasis id="98BE5A4C1079433EFEC3849FEA0F2FC4" box="[289,567,158,184]" italics="true" pageId="10" pageNumber="11">Strobilanthes glandulata</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="6DCAFDDD1079433EFCC8849EEB9E2FC4" box="[810,934,158,185]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="lupulina">
<emphasis id="98BE5A4C1079433EFCC8849EEB9E2FC4" box="[810,934,158,185]" italics="true" pageId="10" pageNumber="11">S. lupulina</emphasis>
</taxonomicName>
(Figure 6 &amp; 8).but differs by having slender rounded (not stout quadrangular) stem, ovate (not broadly elliptic) leaves, leaf margin dentate (not serrate), flowers in oval-elongated heads (not in short heads), bracts well-spaced (not compact), margin crenate (not entire), and 3-4 ribs which are completely encircle the poles of pollen (all ribs joined at poles)
</paragraph>
</subSubSection>
<subSubSection id="E2D0D5D51079433EFF5F852FED192DA9" pageId="10" pageNumber="11" type="description">
<paragraph id="AA75865E1079433EFF5F852FED192DA9" blockId="10.[136,1452,158,868]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF5F852FE9552E34" bold="true" box="[189,365,302,328]" pageId="10" pageNumber="11">Description:—</emphasis>
Shrubs,
<quantity id="6D322BBB1079433EFE27852FEA342E35" box="[453,524,302,329]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="10" pageNumber="11" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
high. Stem slender, rounded, glabrescent below, hispid-pubescent above, muchbranched, young shoots pale green, becoming dark brown when old. Leaves unequal, variable in size; blades ovate, 3–14 ×
<quantity id="6D322BBB1079433EFF3E8577E97E2EED" box="[220,326,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="8.0" metricValueMin="1.5" pageId="10" pageNumber="11" unit="cm" value="4.75" valueMax="8.0" valueMin="1.5">1.5–8 cm</quantity>
, apex acuminate, base attenuate, margin dentate, hispid on the upper surface, bullate above, cystoliths numerous, decurrent on the petiole; petioles
<quantity id="6D322BBB1079433EFD89859BEADD2EC9" box="[619,741,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="7.5" metricValueMin="1.3" pageId="10" pageNumber="11" unit="cm" value="4.4" valueMax="7.5" valueMin="1.3">1.3–7.5 cm</quantity>
long, hispid. Inflorescences terminal and axillary oval-elongated heads; peduncles
<quantity id="6D322BBB1079433EFEA985BFE98A2EA5" box="[331,434,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="10" pageNumber="11" unit="cm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 cm</quantity>
long, hispid; heads
<quantity id="6D322BBB1079433EFD6F85BFEB312EA5" box="[653,777,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="6.1" metricValueMin="4.5" pageId="10" pageNumber="11" unit="cm" value="5.3" valueMax="6.1" valueMin="4.5">4.5–6.1 cm</quantity>
long. Bracts distinct, broadly ovate, obtuse, concave, margin crenate with dotted glands, densely covered with glandular-hairs, lower bracts 3–10 ×
<quantity id="6D322BBB1079433EFBAB85E3EC902E81" box="[1097,1192,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="10" pageNumber="11" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
, smaller than the upper bracts 10–15 ×
<quantity id="6D322BBB1079433EFEDE8607E98C2D5D" box="[316,436,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="7.5" metricValueMin="4.0" pageId="10" pageNumber="11" unit="mm" value="5.75" valueMax="7.5" valueMin="4.0">4–7.5 mm</quantity>
. Calyx
<quantity id="6D322BBB1079433EFDEF8607EA552D5D" box="[525,621,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="10" pageNumber="11" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long, oblong, glabrous, lobes equal. Corolla
<quantity id="6D322BBB1079433EFB958607ECCB2D5D" box="[1143,1267,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="10" pageNumber="11" unit="mm" value="23.0" valueMax="25.0" valueMin="21.0">21–25 mm</quantity>
long,
<quantity id="6D322BBB1079433EFADC8607ED942D5D" box="[1342,1452,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="10" pageNumber="11" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
wide, pale purple, funnel-shaped, glabrous outside, glandular pubescent inside, lobes ovate, obtuse,
<quantity id="6D322BBB1079433EFB05862BED1F2D39" box="[1255,1319,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="mm" value="3.0">3 mm</quantity>
long,
<quantity id="6D322BBB1079433EFA8E862BED942D39" box="[1388,1452,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="11" unit="mm" value="4.0">4 mm</quantity>
wide. Stamens 4, included, 2-longer and 2-shorter, longer stamina filament
<quantity id="6D322BBB1079433EFC3B864FEC6B2D15" box="[985,1107,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="10" pageNumber="11" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">15–18 mm</quantity>
long, shorter stamina filament
<quantity id="6D322BBB1079433EFF6A8673E8DD2DF1" box="[136,229,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="10" pageNumber="11" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
long, stamina filament pubescent. Pollen ellipsoid, tricolporate, P/E = 1.3, ribs 18, ribs have solid aggregates of raised supratectal elements, endoaperture elongate (Figure 2). Ovary ca. 2×
<quantity id="6D322BBB1079433EFC118697EC0A2DCD" box="[1011,1074,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="1.0">1 mm</quantity>
, apex pubescent, style
<quantity id="6D322BBB1079433EFAD18697ED942DCD" box="[1331,1452,662,689]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="10" pageNumber="11" unit="mm" value="20.0" valueMax="22.0" valueMin="18.0">18–22 mm</quantity>
long, filiform, pubescent on lower part; stigma linear, ca.
<quantity id="6D322BBB1079433EFCEF86BBEB762DA9" box="[781,846,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="mm" value="2.0">2 mm</quantity>
long, simple, glabrous. Capsule not seen.
</paragraph>
</subSubSection>
<subSubSection id="E2D0D5D51079433EFF5F86DFEC6E2D84" box="[189,1110,734,761]" pageId="10" pageNumber="11" type="etymology">
<paragraph id="AA75865E1079433EFF5F86DFEC6E2D84" blockId="10.[136,1452,158,868]" box="[189,1110,734,761]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF5F86DFE95C2D84" bold="true" box="[189,356,734,760]" pageId="10" pageNumber="11">Etymology:—</emphasis>
The epithet “
<taxonomicName id="6DCAFDDD1079433EFE1486DFEA572D84" box="[502,623,734,760]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="glandulata">
<emphasis id="98BE5A4C1079433EFE1486DFEA572D84" box="[502,623,734,760]" italics="true" pageId="10" pageNumber="11">glandulata</emphasis>
</taxonomicName>
” refers to the glandular-hairs on the bracts.
</paragraph>
</subSubSection>
<subSubSection id="E2D0D5D51079433EFF5F8703E9072C18" pageId="10" pageNumber="11" type="biology_ecology">
<paragraph id="AA75865E1079433EFF5F8703EA852C61" blockId="10.[136,1452,158,868]" box="[189,701,770,797]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF5F8703E9672C60" bold="true" box="[189,351,770,796]" pageId="10" pageNumber="11">Phenology:—</emphasis>
Flowering from March to May.
</paragraph>
<paragraph id="AA75865E1079433EFF5F8727E9072C18" blockId="10.[136,1452,158,868]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF5F8727E9062C3C" bold="true" box="[189,318,806,832]" pageId="10" pageNumber="11">Habitat:—</emphasis>
The new species grows in the shady places among rocks in the banks of streams and edges of nature trails (Figure 9).
</paragraph>
</subSubSection>
<caption id="FEB5D6D61079433EFF6A81A5EA502AC5" ID-DOI="http://doi.org/10.5281/zenodo.7334297" ID-Zenodo-Dep="7334297" box="[136,616,1444,1466]" httpUri="https://zenodo.org/record/7334297/files/figure.png" pageId="10" pageNumber="11" startId="10.[136,229,1444,1466]" targetBox="[151,1436,893,1419]" targetPageId="10">
<paragraph id="AA75865E1079433EFF6A81A5EA502AC5" blockId="10.[136,616,1444,1466]" box="[136,616,1444,1466]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF6A81A5E8DD2AC6" bold="true" box="[136,229,1444,1466]" pageId="10" pageNumber="11">FIGURE</emphasis>
. Habitats of
<emphasis id="98BE5A4C1079433EFE9181A5EA502AC5" box="[371,616,1444,1465]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="6DCAFDDD1079433EFE9181A5EA5B2AC5" box="[371,611,1444,1465]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="glandulata">Strobilanthes glandulata</taxonomicName>
.
</emphasis>
</paragraph>
</caption>
<subSubSection id="E2D0D5D51079433EFF5F81E9EC062936" pageId="10" pageNumber="11" type="distribution">
<paragraph id="AA75865E1079433EFF5F81E9EC062936" blockId="10.[136,1452,1512,2079]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF5F81E9E94D297E" bold="true" box="[189,373,1512,1538]" pageId="10" pageNumber="11">Distribution:—</emphasis>
It is rather restricted in undisturbed habitat in Rambukoluwa, at an elevation of
<quantity id="6D322BBB1079433EFAE081E9ED72297F" box="[1282,1354,1512,1539]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="10" pageNumber="11" unit="m" value="282.0">282 m</quantity>
towards the eastern slope of the Knuckles Mountain and Madugoda in
<collectingCountry id="D2DDC6CE1079433EFCDC820DEB97295A" box="[830,943,1548,1574]" name="Sri Lanka" pageId="10" pageNumber="11">Sri Lanka</collectingCountry>
. Whereas
<taxonomicName id="6DCAFDDD1079433EFBC3820CECA1295A" box="[1057,1177,1548,1575]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="lupulina">
<emphasis id="98BE5A4C1079433EFBC3820CECA1295A" box="[1057,1177,1548,1575]" italics="true" pageId="10" pageNumber="11">S. lupulina</emphasis>
</taxonomicName>
shows quite widespread distributions at low and high altitudes of the wet zone at an elevation of
<quantity id="6D322BBB1079433EFC538231EC022936" box="[945,1082,1584,1610]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="18.0" metricValueMin="1.0" pageId="10" pageNumber="11" unit="m" value="950.0" valueMax="1800.0" valueMin="100.0">100–1800m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="E2D0D5D51079433EFF5F8255ECE8283A" pageId="10" pageNumber="11" type="conservation">
<paragraph id="AA75865E1079433EFF5F8255ECE8283A" blockId="10.[136,1452,1512,2079]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF5F8255E98D2912" bold="true" box="[189,437,1620,1646]" pageId="10" pageNumber="11">Conservation Status</emphasis>
:—From 2016 to 2021, we visited Madugoda for several times. However we were unable to rediscover
<taxonomicName id="6DCAFDDD1079433EFEE68278E9A129EE" box="[260,409,1656,1683]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="glandulata">
<emphasis id="98BE5A4C1079433EFEE68278E9A129EE" box="[260,409,1656,1683]" italics="true" pageId="10" pageNumber="11">S. glandulata</emphasis>
</taxonomicName>
from Madugoda. Destruction of natural habitats due to agriculture and development projects could be the main reason for the disappearance of this species from the
<typeStatus id="757138FC1079433EFC45829CEBEF29CB" box="[935,983,1693,1719]" pageId="10" pageNumber="11">type</typeStatus>
locality. As a result, the study determined that the only remaining population of
<taxonomicName id="6DCAFDDD1079433EFDDE82C0EAEB29A6" box="[572,723,1728,1755]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="glandulata">
<emphasis id="98BE5A4C1079433EFDDE82C0EAEB29A6" box="[572,723,1728,1755]" italics="true" pageId="10" pageNumber="11">S. glandulata</emphasis>
</taxonomicName>
is in Rambukoluwa. Approximately, 300 individuals have been noted in the Rambukoluwa area, and it seems that the species is well-adapted to the habitat. There is a threat to this population due to road construction for the Kaluganga project. Therefore,
<taxonomicName id="6DCAFDDD1079433EFC5A8308EC74285E" box="[952,1100,1800,1827]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="glandulata">
<emphasis id="98BE5A4C1079433EFC5A8308EC74285E" box="[952,1100,1800,1827]" italics="true" pageId="10" pageNumber="11">S. glandulata</emphasis>
</taxonomicName>
could be classified as Critically Endangered (B1ab, ii, iii, v; B2ab, ii, iii, v) following the IUCN Status Categories under Criterion
</paragraph>
</subSubSection>
<subSubSection id="E2D0D5D51079433FFF5F8351EA292E58" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="AA75865E1079433EFF5F8351ED3728F2" blockId="10.[136,1452,1512,2079]" pageId="10" pageNumber="11">
<materialsCitation id="1AA28C031079433EFF5F8351ED3428F2" ID-GBIF-Occurrence="3967649302" collectingDate="2019-04-24" collectorName="Rambukoluwa, E" county="Knuckles mountain" elevation="258834282" location="Matale District" municipality="Eastern" pageId="10" pageNumber="11" specimenCount="2" typeStatus="paratype">
<emphasis id="98BE5A4C1079433EFF5F8351EC022816" bold="true" box="[189,1082,1872,1899]" pageId="10" pageNumber="11">
Additional specimens of
<taxonomicName id="6DCAFDDD1079433EFE0A8351EB3E2817" box="[488,774,1872,1899]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="glandulata">
<emphasis id="98BE5A4C1079433EFE0A8351EB3E2817" bold="true" box="[488,774,1872,1899]" italics="true" pageId="10" pageNumber="11">Strobilanthes glandulata</emphasis>
</taxonomicName>
examined (
<typeStatus id="757138FC1079433EFC718351EC372816" box="[915,1039,1872,1898]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
):—
</emphasis>
<location id="AF15D0851079433EFBD88351ECDF2816" LSID="urn:lsid:plazi:treatment:226337481075433FFF6A84F5EA292E58:AF15D0851079433EFBD88351ECDF2816" box="[1082,1255,1872,1898]" county="Knuckles mountain" municipality="Eastern" name="Matale District" pageId="10" pageNumber="11">Matale District</location>
,
<collectingMunicipality id="4A111C241079433EFB168350ED702817" box="[1268,1352,1873,1899]" pageId="10" pageNumber="11">Eastern</collectingMunicipality>
slope of
<collectingCounty id="4314FED21079433EFF6A8375E95B28F2" box="[136,355,1908,1934]" pageId="10" pageNumber="11">Knuckles mountain</collectingCounty>
at
<collectorName id="073FE3881079433EFE678375EA7F28F3" box="[389,583,1908,1935]" pageId="10" pageNumber="11">Rambukoluwa, E</collectorName>
202704 N
<quantity id="6D322BBB1079433EFD268375EB5728F3" box="[708,879,1908,1935]" metricMagnitude="8" metricUnit="m" metricValue="2.58834282" pageId="10" pageNumber="11" unit="m" value="2.58834282E8">
<elevation id="21E7616D1079433EFD268375EB5728F3" box="[708,879,1908,1935]" metricMagnitude="8" metricUnit="m" metricValue="2.58834282" pageId="10" pageNumber="11" unit="m" value="2.58834282E8">258834, 282 m</elevation>
</quantity>
,
<date id="DE74A09E1079433EFC998375EC2F28F2" box="[891,1047,1908,1934]" pageId="10" pageNumber="11" value="2019-04-24">
<collectingDate id="CE3059761079433EFC998375EC2F28F2" box="[891,1047,1908,1934]" pageId="10" pageNumber="11" value="2019-04-24">24April 2019</collectingDate>
</date>
, RMRN_123 (PDA!)
</materialsCitation>
.
</paragraph>
<paragraph id="AA75865E1079433FFF5F8399EA292E58" blockId="10.[136,1452,1512,2079]" lastBlockId="11.[136,1451,158,293]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF5F8399EBEA28CE" bold="true" box="[189,978,1944,1971]" pageId="10" pageNumber="11">
<emphasis id="98BE5A4C1079433EFF5F8399E8F528CE" bold="true" box="[189,205,1944,1970]" italics="true" pageId="10" pageNumber="11">S</emphasis>
pecimens of
<taxonomicName id="6DCAFDDD1079433EFE858399EA5628CF" box="[359,622,1944,1971]" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="lupulina">
<emphasis id="98BE5A4C1079433EFE858399EA5628CF" bold="true" box="[359,622,1944,1971]" italics="true" pageId="10" pageNumber="11">Strobilanthes lupulina</emphasis>
</taxonomicName>
examined:—Matale District:
</emphasis>
Lagalla,
<date id="DE74A09E1079433EFBA08399EC8128CE" box="[1090,1209,1944,1970]" pageId="10" pageNumber="11" value="1893-09">Sept 1893</date>
,
<emphasis id="98BE5A4C1079433EFB258398ED7728CE" box="[1223,1359,1945,1970]" italics="true" pageId="10" pageNumber="11">Trimen s.n..</emphasis>
(PDA);
<emphasis id="98BE5A4C1079433EFF6A83BDE97628AA" bold="true" box="[136,334,1980,2006]" pageId="10" pageNumber="11">Kagalle District:</emphasis>
Kalugala,
<date id="DE74A09E1079433EFE2A83BDEA6328AA" box="[456,603,1980,2006]" pageId="10" pageNumber="11" value="1927-08-28">28 Aug 1927</date>
,
<emphasis id="98BE5A4C1079433EFD8483BDEAE628AA" box="[614,734,1980,2006]" italics="true" pageId="10" pageNumber="11">Alston 877</emphasis>
(PDA); Across river from Kitulgala,
<date id="DE74A09E1079433EFB9D83BDED3E28AA" box="[1151,1286,1980,2007]" pageId="10" pageNumber="11" value="1967-11-06">6 Nov 1967</date>
,
<emphasis id="98BE5A4C1079433EFAF383BCED9328AA" box="[1297,1451,1980,2007]" italics="true" pageId="10" pageNumber="11">Comanor 540</emphasis>
(PDA); Ambarawella,
<date id="DE74A09E1079433EFE6683E1EA2F2886" box="[388,535,2016,2042]" pageId="10" pageNumber="11" value="1968-08-23">23 Aug 1968</date>
,
<emphasis id="98BE5A4C1079433EFDC383E0EAD42886" box="[545,748,2016,2043]" italics="true" pageId="10" pageNumber="11">Amaratunga 1629</emphasis>
(PDA);
<emphasis id="98BE5A4C1079433EFCA983E1EC1A2886" bold="true" box="[843,1058,2016,2042]" pageId="10" pageNumber="11">Kalutara District:</emphasis>
Kalugala,
<date id="DE74A09E1079433EFB7E83E1ED1F2887" box="[1180,1319,2016,2043]" pageId="10" pageNumber="11" value="1975-06-28">28 Jun 1975</date>
,
<emphasis id="98BE5A4C1079433EFAD383E0ED932886" box="[1329,1451,2016,2042]" italics="true" pageId="10" pageNumber="11">Waas 1290</emphasis>
(PDA); Kalwana-Agalawatta,
<quantity id="6D322BBB1079433EFE398C05EA172762" box="[475,559,2052,2078]" metricMagnitude="4" metricUnit="m" metricValue="4.3452288" pageId="10" pageNumber="11" unit="mi" value="27.0">27 mile</quantity>
post,
<date id="DE74A09E1079433EFD908C05EAC92763" box="[626,753,2052,2079]" pageId="10" pageNumber="11" value="1975-09-09">9 Sep 1975</date>
,
<emphasis id="98BE5A4C1079433EFD1E8C05EC7A2762" box="[764,1090,2052,2078]" italics="true" pageId="10" pageNumber="11">Sumithraarachchi et al. 1010</emphasis>
(PDA); Kalugala Forest, Pahala Hewessa,
<date id="DE74A09E1078433FFF1A849FE9452FC5" box="[248,381,158,185]" pageId="11" pageNumber="12" value="1975-11-06">6 Nov 1975</date>
,
<emphasis id="98BE5A4C1078433FFE65849FEABD2FC4" box="[391,645,158,185]" italics="true" pageId="11" pageNumber="12">Sohmer &amp; Waas 10276</emphasis>
(PDA); half mile north of Pahala Hewessa,
<date id="DE74A09E1078433FFB88849FECCE2FC4" box="[1130,1270,158,184]" pageId="11" pageNumber="12" value="1977-02-17">17 Feb 1977</date>
.
<emphasis id="98BE5A4C1078433FFAE2849EE88A2FA0" italics="true" pageId="11" pageNumber="12">B. &amp; K. Bremer 810</emphasis>
(PDA), RATNAPURA DISTRICT: Kokawatte, Sinharaja Forest,
<date id="DE74A09E1078433FFC7D84C2EC372FA0" box="[927,1039,194,221]" pageId="11" pageNumber="12" value="1893-12">Dec 1893</date>
,
<emphasis id="98BE5A4C1078433FFBF984C2EC9A2FA1" box="[1051,1186,195,221]" italics="true" pageId="11" pageNumber="12">Trimen S.N.</emphasis>
(PDA);
<emphasis id="98BE5A4C1078433FFAE684C3E8D42E7C" bold="true" pageId="11" pageNumber="12">Nuwara Eliya District:</emphasis>
Gardner in
<emphasis id="98BE5A4C1078433FFE8C84E7EA0F2E7C" box="[366,567,230,257]" italics="true" pageId="11" pageNumber="12">Thwaites C.P. 254</emphasis>
(PDA); Road from Pundaluoya to highway A5 and Nuwara Eliya,
<date id="DE74A09E1078433FFAFA84E7ED9F2E7C" box="[1304,1447,230,257]" pageId="11" pageNumber="12" value="1970-12-26">26 Dec 1970</date>
,
<emphasis id="98BE5A4C1078433FFF6A850BE9802E58" box="[136,440,266,293]" italics="true" pageId="11" pageNumber="12">Theobald &amp; Krahulik 2818</emphasis>
(PDA).
</paragraph>
</subSubSection>
</treatment>
</document>
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