149 lines
17 KiB
XML
149 lines
17 KiB
XML
<document id="6C3709DB028CC373BEC94B6970513A80" ID-DOI="http://dx.doi.org/10.3897/zookeys.665.11792" ID-GBIF-Dataset="f49697cc-fc0c-4a14-bba1-edf1268f59f0" ID-PMC="PMC5523169" ID-Pensoft-Pub="1313-2970-665-93" ID-PubMed="28769629" ID-ZooBank="4D5230AC67CB47ADAF2F051CA10D9724" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-665-93" ModsDocOrigin="ZooKeys 665" ModsDocTitle="Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1491378304537" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Wild, Michael, Sumoked, Bob & Balke, Michael" docDate="2017" docId="0F102D884A0043AA227C44A6C98DF1B5" docLanguage="en" docName="ZooKeys 665: 93-120" docOrigin="ZooKeys 665" docSource="http://dx.doi.org/10.3897/zookeys.665.11792" docTitle="Exocelina bagus Balke & Hendrich 2001" docType="treatment" docVersion="5" lastPageNumber="95" masterDocId="DC520217FF93FFF8FFDEFFCFFFA3777E" masterDocTitle="Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="120" masterPageNumber="93" pageNumber="95" updateTime="1701318038850" updateUser="plazi">
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<mods:title id="4F78787E27259E645B50ACF45537D0A6">Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart id="EC2B0973A9528B86A547A8B87C52F9AE">Shaverdo, Helena</mods:namePart>
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<mods:namePart id="5024F75C5B58E3E4871B5C02A691ED98">Wild, Michael</mods:namePart>
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<mods:namePart id="E4C1B789F52989768E404D0BB88A8035">Sumoked, Bob</mods:namePart>
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<mods:namePart id="0F0982F75F5ADC7531E6EF8B61E76BC3">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="5330AE8ED50DE249624F1C876BF9F70E">text</mods:typeOfResource>
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<mods:title id="69DA099A85A928B83E4C02A5560FF9DF">ZooKeys</mods:title>
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<mods:date id="0C518849D6912F22D38D3B159C4EF1FF">2017</mods:date>
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<mods:detail id="29D2139FE65D0A1BCF2B48590CF93468" type="volume">
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<mods:number id="011C92A6927DC80F51162BCE7DC2B185">665</mods:number>
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<mods:url id="BF219B9E9864C317CA26721CD8415EB0">http://dx.doi.org/10.3897/zookeys.665.11792</mods:url>
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<mods:classification id="6829B8F5BE5F54A8786CF69C9B2E4337">journal article</mods:classification>
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<mods:identifier id="13C80BE873964243EA19F1876EAA3196" type="DOI">http://dx.doi.org/10.3897/zookeys.665.11792</mods:identifier>
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<mods:identifier id="B3EF79C486AB6E29783122450E642306" type="Pensoft-Pub">1313-2970-665-93</mods:identifier>
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<mods:identifier id="94DB0CDDA35D0749DAC8EFDFA3C00A85" type="ZooBank">4D5230AC67CB47ADAF2F051CA10D9724</mods:identifier>
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<treatment id="0F102D884A0043AA227C44A6C98DF1B5" ID-GBIF-Taxon="128229139" LSID="urn:lsid:plazi:treatment:0F102D884A0043AA227C44A6C98DF1B5" httpUri="http://treatment.plazi.org/id/0F102D884A0043AA227C44A6C98DF1B5" lastPageNumber="95" pageId="2" pageNumber="95">
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<subSubSection id="91F0E6C184A43D46D7F42A2C7B67EAC0" pageId="2" pageNumber="95" type="nomenclature">
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<paragraph id="30E7C09B87103D04922E24DED5851CB1" pageId="2" pageNumber="95">
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3.
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<taxonomicName id="56D3A77F533310F62693DC4E3E0F77B2" ID-CoL="3DLP8" authority="Balke & Hendrich, 2001" authorityName="Balke & Hendrich" authorityYear="2001" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina bagus" order="Coleoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="bagus">Exocelina bagus (Balke & Hendrich, 2001)</taxonomicName>
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Figs 6, 14
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</paragraph>
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</subSubSection>
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<subSubSection id="922F72DB18FEBDAAFD23F90893D5E845" pageId="2" pageNumber="95" type="reference_group">
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<paragraph id="00A270D5CFB6A22AD253C22736C9A08A" pageId="2" pageNumber="95">
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<taxonomicName id="3B23586EE442C85DDAB949703FA652BB" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) speciosus" order="Coleoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="speciosus" subGenus="Papuadytes">Copelatus (Papuadytes) speciosus</taxonomicName>
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Balke & Hendrich, 1998: 336;
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<bibRefCitation id="6EAC1F39A43CEE7A76FA463798CC259F" author="Nilsson, AN" journalOrPublisher="Stenstrup, Apollo Books" pageId="12" pageNumber="105" title="Dytiscidae. World catalogue of insects. Vol. 3." year="2001">Nilsson 2001</bibRefCitation>
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: 77 (catalogue).
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</paragraph>
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<paragraph id="40A6F4E8A5541BC21D7B5AA3CEF34CD4" pageId="2" pageNumber="95">
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<taxonomicName id="B4E914BA5C0D113A6F24DDA8B881EAC4" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) bagus" order="Coleoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="bagus" subGenus="Papuadytes">Copelatus (Papuadytes) bagus</taxonomicName>
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Balke & Hendrich, in
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<bibRefCitation id="3247CB764A16E04F5D0F7024E323D19C" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="12" pageNumber="105" pagination="361 - 362" title="Replacement names for three New Guinea species of Copelatus, subgenus Papuadytes Balke, 1998 (Coleoptera: Dytiscidae)" volume="103 B" year="2001">Balke 2001</bibRefCitation>
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: 361.
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</paragraph>
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<paragraph id="A6DCEA1D0ABFAA984E64375B9764D5AA" pageId="2" pageNumber="95">
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<taxonomicName id="669F1B9CD7D188D057AA43247053B850" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina bagus" order="Coleoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="bagus">Exocelina bagus</taxonomicName>
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(Balke & Hendrich, 2001):
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<bibRefCitation id="A36F6ACC5134739ED4E0271F2A0E2658" author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="12" pageNumber="105" pagination="33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
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: 33 (as
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<taxonomicName id="BB92A72A43E00E7B6CC21C9D7028BB37" family="Dytiscidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. baga" order="Coleoptera" pageId="2" pageNumber="95" rank="species" species="baga">E. baga</taxonomicName>
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, comb.n.).
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</paragraph>
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<paragraph id="A570F3F0729E69F88A1965D2808AAD87" pageId="2" pageNumber="95">
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<taxonomicName id="1830542B8D2CC22D16166443009082D4" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina bagus" order="Coleoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="bagus">Exocelina bagus</taxonomicName>
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MB4915:
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<bibRefCitation id="0DB5E96C724058EC1A77055A043B6EC0" author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="13" pageNumber="106" pagination="1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity" url="https://doi.org/10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
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: Supplementary figs 1-4, Tab. 2.
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</paragraph>
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</subSubSection>
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<subSubSection id="F21906D792755816BC4ACE11D3003780" pageId="2" pageNumber="95" type="type locality">
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<paragraph id="7AD2A7D594C0210C14D1D8EE0B46556B" pageId="2" pageNumber="95">Type locality.</paragraph>
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<paragraph id="09CE7C338D04E89EE8B8EA9D4F03DFBB" pageId="2" pageNumber="95">
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Papua: Nabire Regency, 54-55 km of road Nabire to Enarotali, ca.
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<geoCoordinate id="13D89FCF1F9A9ABCA0A7D2F094B00D30" direction="south" orientation="latitude" precision="9" value="-3.4966667">03°29.80'S</geoCoordinate>
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;
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<geoCoordinate id="CF4E05785FCCA99BBE48A4853B4729B5" direction="east" orientation="longitude" precision="9" value="135.7315">135°43.89'E</geoCoordinate>
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. Note: the road only goes up to Enarotali, Ilaga is much further in the mountains, therefore, people now refer to the road as Nabire-Enarotali.
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</paragraph>
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</subSubSection>
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<subSubSection id="6B8860572859F63FD7631EFD5610FFF2" pageId="2" pageNumber="95" type="type material">
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<paragraph id="DB1BEDFD76CEA27C601B403ED620C785" pageId="2" pageNumber="95">Type material studied.</paragraph>
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<paragraph id="B842AD136499B74DD4C1D06A80FDCB53" pageId="2" pageNumber="95">
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Holotype: male "IR90-11: W. New Guinea, Trek Nabire-Ilaga, km55, 19.-25.ix.1990, Balke",
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<normalizedToken id="FA0F0EF7D4F714D7421F8C3AC3B17CEB" originalValue="“HOLOTYPUS”">"HOLOTYPUS"</normalizedToken>
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[red], "
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<taxonomicName id="F8C7F4AFD2F3E0D70E8D0C52F4F94D43" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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<taxonomicName id="6E0D46049775084390C6EF4D72801DA6" lsidName="speciosus" pageId="2" pageNumber="95" rank="species" species="speciosus">speciosus</taxonomicName>
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sp.n. Balke des. 1997" [red] (NHMW). Paratypes: 9 males, 12 females with the same label as the holotype (NHMW). 6 females "W.-Neuguinea/Paniai Prov. Strasse Nabire-Ilaga km 54 700m, 22.-25.9.1990/IR 11 leg: Balke & Hendrich" (NHMW). 7 males, 8 females "IR 20-W. New Guinea, track Nabire-Ilaga KM 59, ca.750m, 18.vii.1991, Balke & Hendrich leg." (NHMW). 1 female "IR 21-W. New Guinea track Nabire-Ilaga KM 65, Kali Utowa, 250 M, 18-19.vii.1991 Balke & Hendrich leg." (NHMW), note: the original description says "one male", it is probably a mistake in the sex identification or type error. 6 females "IR 23-W. New Guinea, track Nabire-Ilaga, KM 62, 250m, 24.vii.1991 Balke & Hendrich leg." (NHMW). 1 female "IR 24-W. New Guinea, track Nabire-Ilaga Km 54, basecamp, 750m, 25.vii.1991 Balke & Hendrich leg." (NHMW). All paratypes are additionally with red labels "Paratypus
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<taxonomicName id="886942AB870754375A23F2221BB41A2A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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<taxonomicName id="19732CFF2D5A0DF3EDEA534FB2AB1382" lsidName="speciosus" pageId="2" pageNumber="95" rank="species" species="speciosus">speciosus</taxonomicName>
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Balke des. 1997".
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</paragraph>
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</subSubSection>
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<subSubSection id="1C75C5EA0BD8D584E51C1E033F8470D4" pageId="2" pageNumber="95" type="materials_examined">
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<paragraph id="A75E84EC56E1CF44C425E5E4E55347CC" pageId="2" pageNumber="95">Additional material.</paragraph>
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<paragraph id="8CFD02C6075093C6454DFA3EC2599FFE" pageId="2" pageNumber="95">
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10 males, 15 females "IR #91-7 (IR 24). West New Guinea, Nabire-Ilaga km 54, 750m, 25.&27.1991 Balke" (NHMW). 8 males, 8 females "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 26./27.8.1996, 750-800m leg. M. Balke (96 # 2)" (NHMW). 6 males, 5 females "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 30.8.1996, 750m leg. M. Balke (96 # 9)" (NHMW). 2 males "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 10.9.1996, 900m leg. M. Balke (96 # 19)" (NHMW). 7 males, 3 females "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 10.9.1996, 800m leg. M. Balke (96 # 20)" (NHMW). Note: although most of these mentioned above specimens are with the paratype labels, they are not included in the type material of the original description in
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<bibRefCitation id="C9ADE8E77990D5948165A4B20A152E36" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="11" pageNumber="104" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I" volume="100 B" year="1998">Balke (1998</bibRefCitation>
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, p. 336). 1 female "Indonesia: Papua, Road Nabire-Enarotali KM 55, 774m, 22.x.2011,
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<geoCoordinate id="5B55C5E121E828BE3088E3B5B9FF3714" direction="south" orientation="latitude" precision="1" value="-3.4966">03 29.796S</geoCoordinate>
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<geoCoordinate id="793D458646BD025F50261CE5D9CBCB05" direction="east" orientation="longitude" precision="1" value="135.73141">135 43.885E</geoCoordinate>
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, Uncen (PAP09)" (NHMW). 2 males, 1 female "Indonesia: Papua, Road Nabire-Enarotali KM 60, 640m, 22.x.2011,
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<geoCoordinate id="321EA6C7EE5F78834E04477BABD9C914" direction="south" orientation="latitude" precision="1" value="-3.5079">03 30.474S</geoCoordinate>
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<geoCoordinate id="A0D5CE4921618E27A29C34A2B63465AA" direction="east" orientation="longitude" precision="1" value="135.71019">135 42.611E</geoCoordinate>
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, Uncen (PAP10)" (NHMW).
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</paragraph>
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</subSubSection>
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<subSubSection id="0487363132934EDCB5A9072154F66D69" pageId="2" pageNumber="95" type="diagnosis">
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<paragraph id="AD2B74A4A4AC20C1073796D2B873A8B1" pageId="2" pageNumber="95">Diagnosis.</paragraph>
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<paragraph id="3D12E097864149806B24070CCC5CF949" pageId="2" pageNumber="95">
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Beetle medium-sized (TL-H 3.8-4.8 mm), elongate; dark brown, with reddish brown pronotal sites and head anteriorly; submatt, with fine but evident punctation and rather strongly impressed microreticulation; pronotum without lateral bead (Fig. 6; fig. 3 in
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<bibRefCitation id="3E7F00CB4DE43BC200D5A7A578FBBAEF" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="11" pageNumber="104" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I" volume="100 B" year="1998">Balke (1998)</bibRefCitation>
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); male antennae strongly and modified: antennomeres 2-3 very strongly reduced, 4-6 excessively enlarged and 3 and 7 strongly enlarged (Fig. 6; fig. 16 in
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<bibRefCitation id="4A55B15DE110B97AB8E5E64813E291C2" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="11" pageNumber="104" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I" volume="100 B" year="1998">Balke (1998)</bibRefCitation>
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); male protarsomere 4 with large, strongly curved anterolateral hook-like setae; male protarsomere 5 slightly concave ventrally, with anterior band of more than 30 and posterior row of five relatively long, not pointed setae (Fig. 14D); median lobe strongly curved, evenly tapering to apex, apex straight, pointed, with short, thick lateral setae in lateral view, subdistal part of median lobe strongly broadened in ventral view; paramere with distinct dorsal notch and large, elongate subdistal part; subdistal setae very dense, strong, long, proximal setae very sparse, thin, small, weakly visible (Fig. 14
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<normalizedToken id="BF7EAF497EC3705DDC7C1A6E529AC528" originalValue="A–C">A-C</normalizedToken>
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; figs 39, 86, 92 in
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<bibRefCitation id="AC98B6F3B44A12D57170943571EAB2DE" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="11" pageNumber="104" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I" volume="100 B" year="1998">Balke (1998)</bibRefCitation>
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). For complete description, see
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<bibRefCitation id="6EB7B74C693E3CCBEC9559E5B3B6D522" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="11" pageNumber="104" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I" volume="100 B" year="1998">Balke (1998)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C093B83673E1F056BA47998D75B71B15" pageId="2" pageNumber="95" type="distribution">
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<paragraph id="F95085E88F9C434D6E5A418E4CB286F8" pageId="2" pageNumber="95">Distribution.</paragraph>
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<paragraph id="A8E4A1BC0D766BE0BBDC4FC3B7D7C9EF" pageId="2" pageNumber="95">Papua: Nabire Regency. The species is known only from the Mount Gamey area (Fig. 19).</paragraph>
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</subSubSection>
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</treatment>
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</document> |