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<document id="167DB06A0F8753343EDF3B918B86AACD" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F11EFFEE75C6FE63511E7B79" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) kinabaluensis Pace 2008" docType="treatment" docVersion="12" lastPageNumber="231" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="231" updateTime="1699471357828" updateUser="ExternalLinkService">
<mods:mods id="612DC8AB67608C5E55A50D5C1AC99EDE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5C6EDF5D9160564666D62D6F4C643DF8">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="6A65195DBA84F332269734B29E27D015">Assing, Volker</mods:namePart>
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<mods:title id="0F2D77165AD67B8959AE6196302CA2E8">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="B1058D49659F0291CB69D665A4B4A7D2">
<mods:date id="D68E742C551CACC762AB50141EEFD13A">2017</mods:date>
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<mods:number id="A10868CA5B87046D36167DCC1A7D27A4">2017-12-08</mods:number>
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<mods:detail id="C5058A9B114B8A3CE0EB606123945D03" type="volume">
<mods:number id="3D2B01DB54288535DF43F796FFEC1D13">67</mods:number>
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<mods:classification id="378F35AF1F588AF999E2A22F225BFDAD">journal article</mods:classification>
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<mods:identifier id="D2978BE805415E208124BB9573D48682" type="DOI">10.21248/contrib.entomol.67.2.213-246</mods:identifier>
<mods:identifier id="FE0DBF377146EB0ACDFEF9511E24D5BC" type="GBIF-Dataset">43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924</mods:identifier>
<mods:identifier id="F254AF74FEE2D0BBC5BE517629041889" type="ISSN">0005-805X</mods:identifier>
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<mods:identifier id="F5A157B3181EAA87CFAFBC00D69440D7" type="ZooBank">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</mods:identifier>
</mods:mods>
<treatment id="03FD87A1F11EFFEE75C6FE63511E7B79" ID-DOI="http://doi.org/10.5281/zenodo.5888800" ID-GBIF-Taxon="192500449" ID-Zenodo-Dep="5888800" LSID="urn:lsid:plazi:treatment:03FD87A1F11EFFEE75C6FE63511E7B79" httpUri="http://treatment.plazi.org/id/03FD87A1F11EFFEE75C6FE63511E7B79" lastPageNumber="231" pageId="18" pageNumber="231">
<subSubSection id="C34E653CF11EFFEE75C6FE63561B7EF6" box="[227,623,435,462]" pageId="18" pageNumber="231" type="nomenclature">
<paragraph id="8BEB36B7F11EFFEE75C6FE63561B7EF6" blockId="18.[227,623,435,495]" box="[227,623,435,462]" pageId="18" pageNumber="231">
<heading id="D0A381DBF11EFFEE75C6FE63561B7EF6" box="[227,623,435,462]" centered="true" fontSize="12" level="2" pageId="18" pageNumber="231" reason="2">
<taxonomicName id="4C544D34F11EFFEE75C6FE63561B7EF6" ID-CoL="5DDDB" authority="PACE, 2008" authorityName="Pace" authorityYear="2008" box="[227,623,435,462]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="species" species="kinabaluensis" subGenus="Zyras">
<emphasis id="B920EAA5F11EFFEE75C6FE63556D7EF6" box="[227,281,435,462]" italics="true" pageId="18" pageNumber="231">Zyras</emphasis>
(
<emphasis id="B920EAA5F11EFFEE7402FE6355297EF6" box="[295,349,435,462]" italics="true" pageId="18" pageNumber="231">Zyras</emphasis>
)
<emphasis id="B920EAA5F11EFFEE744FFE6355827EF6" box="[362,502,435,462]" italics="true" pageId="18" pageNumber="231">kinabaluensis</emphasis>
<bibRefCitation id="EFC54B46F11EFFEE74D8FE63561B7EF6" author="PACE, R." box="[509,623,435,462]" pageId="18" pageNumber="231" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE, 2008</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11EFFEE75C6FE0B55257ED7" box="[227,337,475,495]" pageId="18" pageNumber="231" type="description">
<paragraph id="8BEB36B7F11EFFEE75C6FE0B55257ED7" blockId="18.[227,623,435,495]" box="[227,337,475,495]" pageId="18" pageNumber="231">
(
<figureCitation id="136F2A32F11EFFEE75C2FE0B55467ED7" box="[231,306,475,495]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="18" pageNumber="231">Figs 4, 21</figureCitation>
,
<figureCitation id="136F2A32F11EFFEE741CFE0B55397ED7" box="[313,333,475,495]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="18" pageNumber="231">44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11EFFEE7598FDDC56177D18" box="[189,611,523,545]" pageId="18" pageNumber="231" type="reference_group">
<paragraph id="8BEB36B7F11EFFEE7598FDDC56177D18" blockId="18.[189,611,523,545]" box="[189,611,523,545]" pageId="18" pageNumber="231">
<treatmentCitationGroup id="AB441199F11EFFEE7598FDDC56177D18" box="[189,611,523,545]" pageId="18" pageNumber="231">
<treatmentCitation id="0AF510A6F11EFFEE7598FDDC562B7D18" author="PACE, R." box="[189,607,523,545]" page="148" pageId="18" pageNumber="231" year="2008">
<taxonomicName id="4C544D34F11EFFEE7598FDDC562B7D18" ID-CoL="5DDDB" authority="PACE, 2008: 148" authorityName="Pace" authorityPageNumber="148" authorityYear="2008" box="[189,607,523,545]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="species" species="kinabaluensis" subGenus="Zyras">
<emphasis id="B920EAA5F11EFFEE7598FDDC54857D19" box="[189,241,524,545]" italics="true" pageId="18" pageNumber="231">Zyras</emphasis>
(
<emphasis id="B920EAA5F11EFFEE75DAFDDC55477D19" box="[255,307,524,545]" italics="true" pageId="18" pageNumber="231">Zyras</emphasis>
)
<emphasis id="B920EAA5F11EFFEE7464FDDB55B47D18" box="[321,448,523,544]" italics="true" pageId="18" pageNumber="231">kinabaluensis</emphasis>
<bibRefCitation id="EFC54B46F11EFFEE74E0FDDB562B7D18" author="PACE, R." box="[453,607,523,544]" pageId="18" pageNumber="231" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE, 2008: 148</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11EFFEE7598FD99555D7C39" pageId="18" pageNumber="231" type="materials_examined">
<paragraph id="8BEB36B7F11EFFEE7598FD99555D7C39" blockId="18.[189,794,585,769]" pageId="18" pageNumber="231">
<emphasis id="B920EAA5F11EFFEE7598FD9955B27D59" bold="true" box="[189,454,585,609]" pageId="18" pageNumber="231">Type material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF11EFFEE74F6FD99555D7C39" ID-GBIF-Occurrence="3450187341" collectingDate="1987-05-21" collectionCode="MHNG, ENTO" collectorName="Burckhardt &amp; Lobl" determinerName="V. Assing" elevation="1500" location="Mt. Kinabalu" municipality="Mt. Kinabalu" pageId="18" pageNumber="231" specimenCount="1" stateProvince="SABAH" typeStatus="holotype">
<typeStatus id="54EF8815F11EFFEE74F6FD9956427D59" box="[467,566,585,609]" pageId="18" pageNumber="231" type="holotype">Holotype</typeStatus>
<specimenCount id="9D52FD3EF11EFFEE7719FD9A56257D59" box="[572,593,586,609]" pageId="18" pageNumber="231" type="generic">♂</specimenCount>
: “♂ /
<collectingRegion id="4990F855F11EFFEE77B6FD99569E7D59" box="[659,746,585,609]" country="Malaysia" name="Sabah" pageId="18" pageNumber="231">SABAH</collectingRegion>
,
<collectingMunicipality id="6B8FACCDF11EFFEE77D0FD9A55547DB9" name="Mt. Kinabalu" pageId="18" pageNumber="231" stateProvince="Sabah">Mt. Kinabalu</collectingMunicipality>
,
<quantity id="4CAC9B52F11EFFEE7409FDB955097DB9" box="[300,381,617,641]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="231" unit="m" value="1500.0">
<elevation id="0079D184F11EFFEE7409FDB955097DB9" box="[300,381,617,641]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="231" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FFEA1077F11EFFEE74ACFDB955877DB9" box="[393,499,617,641]" pageId="18" pageNumber="231" value="1987-05-21">
<collectingDate id="EFAEE99FF11EFFEE74ACFDB955877DB9" box="[393,499,617,641]" pageId="18" pageNumber="231" value="1987-05-21">21.V.1987</collectingDate>
</date>
,
<collectorName id="26A15361F11EFFEE74DAFDB9560C7DB9" box="[511,632,617,641]" pageId="18" pageNumber="231">Burckhardt</collectorName>
-
<collectorName id="26A15361F11EFFEE77B7FDB956B67DB9" box="[658,706,617,641]" pageId="18" pageNumber="231">Löbl</collectorName>
/
<typeStatus id="54EF8815F11EFFEE77FEFDB954837D9A" pageId="18" pageNumber="231" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C544D34F11EFFEE75DBFD5A55B87D99" authorityName="Pace" authorityYear="2008" box="[254,460,649,673]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B920EAA5F11EFFEE75DBFD5A55B87D99" box="[254,460,649,673]" italics="true" pageId="18" pageNumber="231">Zyras kinabaluensis</emphasis>
</taxonomicName>
mihi, det. R. Pace 2000 /
<taxonomicName id="4C544D34F11EFFEE77C5FD5A553E7DF9" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="species" species="kinabaluensis" status="sp. nov.">
<emphasis id="B920EAA5F11EFFEE77C5FD5A553E7DF9" italics="true" pageId="18" pageNumber="231">Zyras kinabaluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF11EFFEE7471FD7A55FA7DF9" box="[340,398,682,705]" pageId="18" pageNumber="231" rank="species">n. sp.</taxonomicNameLabel>
, det. R. Pace 2000 /
<collectionCode id="ED45AE72F11EFFEE7753FD7956BD7DF9" box="[630,713,681,705]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="18" pageNumber="231">MHNG</collectionCode>
<collectionCode id="ED45AE72F11EFFEE77F6FD79576E7DF9" box="[723,794,681,705]" collectionName="ENTO" pageId="18" pageNumber="231">ENTO</collectionCode>
00010003 /
<taxonomicName id="4C544D34F11EFFEE7412FD1A56497DD9" authority="Pace" authorityName="Pace" authorityYear="2008" box="[311,573,713,737]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B920EAA5F11EFFEE7412FD1A56707DD9" box="[311,516,713,737]" italics="true" pageId="18" pageNumber="231">Zyras kinabaluensis</emphasis>
Pace
</taxonomicName>
, det.
<determinerName id="7E37DA73F11EFFEE7751FD1A56A27DD9" box="[628,726,713,737]" pageId="18" pageNumber="231">
V.
<bibRefCitation id="EFC54B46F11EFFEE77AAFD1956A27DD9" author="ASSING, V." box="[655,726,713,737]" pageId="18" pageNumber="231" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
</determinerName>
2017” (
<collectionCode id="ED45AE72F11EFFEE75E0FD39556A7C39" box="[197,286,745,769]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="18" pageNumber="231" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11EFFEE7598FCF956D87C59" pageId="18" pageNumber="231" type="discussion">
<paragraph id="8BEB36B7F11EFFEE7598FCF956D87C59" blockId="18.[189,794,809,865]" pageId="18" pageNumber="231">
<emphasis id="B920EAA5F11EFFEE7598FCF955587C79" bold="true" box="[189,300,809,833]" pageId="18" pageNumber="231">Comment</emphasis>
: The original description is based on a unique
<specimenCount id="9D52FD3EF11EFFEE7598FC9954857C59" box="[189,241,841,865]" pageId="18" pageNumber="231" type="male">male</specimenCount>
from “
<collectingRegion id="4990F855F11EFFEE741EFC9955097C59" box="[315,381,841,865]" country="Malaysia" name="Sabah" pageId="18" pageNumber="231">Sabah</collectingRegion>
,
<location id="8E8B606CF11EFFEE74A2FC9A56617C59" LSID="urn:lsid:plazi:treatment:03FD87A1F11EFFEE75C6FE63511E7B79:8E8B606CF11EFFEE74A2FC9A56617C59" box="[391,533,841,865]" name="Mt. Kinabalu" pageId="18" pageNumber="231" stateProvince="Sabah">Mt. Kinabalu</location>
” (
<bibRefCitation id="EFC54B46F11EFFEE7708FC9956D57C59" author="PACE, R." box="[557,673,841,865]" pageId="18" pageNumber="231" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">
<specimenCode id="DBF29ECCF11EFFEE7708FC9956D57C59" box="[557,673,841,865]" pageId="18" pageNumber="231">PACE 2008</specimenCode>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11EFFEE7598FC5950357CD9" pageId="18" pageNumber="231" type="description">
<paragraph id="8BEB36B7F11EFFEE7598FC5955BA7B79" blockId="18.[189,794,905,2081]" pageId="18" pageNumber="231">
<emphasis id="B920EAA5F11EFFEE7598FC5955227C99" bold="true" box="[189,342,905,929]" pageId="18" pageNumber="231">Redescription</emphasis>
: Moderately small species; body length
<quantity id="4CAC9B52F11EFFEE7598FC7955657CF9" box="[189,273,937,961]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="18" pageNumber="231" unit="mm" value="5.7">5.7 mm</quantity>
; length of forebody
<quantity id="4CAC9B52F11EFFEE7711FC7956FC7CF9" box="[564,648,937,961]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="18" pageNumber="231" unit="mm" value="2.4">2.4 mm</quantity>
. Coloration (
<figureCitation id="136F2A32F11EFFEE75E1FC1955587CD9" box="[196,300,969,993]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="18" pageNumber="231">Figs 4, 21</figureCitation>
,
<figureCitation id="136F2A32F11EFFEE7413FC1955207CD9" box="[310,340,969,993]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="18" pageNumber="231">44</figureCitation>
): body brown with the anterior portions of the paratergites slightly paler; legs yellow with the apical halves of the metafemora slightly darker; antennae and maxillary palpi yellowish.
</paragraph>
<paragraph id="8BEB36B7F11EFFEE7598FB9956557AB9" blockId="18.[189,794,905,2081]" pageId="18" pageNumber="231">
Head (
<figureCitation id="136F2A32F11EFFEE742FFB9955217B59" box="[266,341,1097,1121]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="18" pageNumber="231">Fig. 21</figureCitation>
) moderately transverse, median portion extensively impunctate; punctures in lateral portions coarse and sparse. Eyes moderately large, slightly longer than postocular region in dorsal view. Antenna (
<figureCitation id="136F2A32F11EFFEE77F1FB7957667BF9" box="[724,786,1193,1217]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="18" pageNumber="231">Fig. 4</figureCitation>
) moderately long and rather slender,
<quantity id="4CAC9B52F11EFFEE7764FB1956E67BD9" box="[577,658,1225,1249]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="231" unit="mm" value="2.2">2.2 mm</quantity>
long; antennomeres IV more than 1.5 times as long as broad, V–VIII of gradually decreasing length and decreasingly oblong, VIII weakly oblong, IX approximately as long as broad, X weakly transverse, and XI short, distinctly shorter than the combined length of IX and X.
</paragraph>
<paragraph id="8BEB36B7F11EFFEE7598FA5A56667959" blockId="18.[189,794,905,2081]" pageId="18" pageNumber="231">
Pronotum (
<figureCitation id="136F2A32F11EFFEE741EFA5955F17A99" box="[315,389,1417,1441]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="18" pageNumber="231">Fig. 21</figureCitation>
) slender, 1.07 times as broad as long and 1.11 times as broad as head, broadest near anterior angles, rather strongly convex in cross-section; lateral margins weakly sinuate in posterior third in dorsal view; posterior angles obtusely marked; punctation dense, coarse, and somewhat irregularly distributed; impunctate median band moderately broad.
</paragraph>
<paragraph id="8BEB36B7F11EFFEE7598F9B9556D7819" blockId="18.[189,794,905,2081]" pageId="18" pageNumber="231">
Elytra (
<figureCitation id="136F2A32F11EFFEE742BF9B9552D79B9" box="[270,345,1641,1665]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="18" pageNumber="231">Fig. 21</figureCitation>
) very short, 0.65 times as long as pronotum; punctation moderately dense, very coarse, slightly less dense posteriorly than anteriorly. Hind wings completely reduced. Legs long and slender; metatarsomere I slender, nearly as long as the combined length of II–IV.
</paragraph>
<paragraph id="8BEB36B7F11EFFEE7598F8F9579C7EF9" blockId="18.[189,794,905,2081]" lastBlockId="18.[831,1437,233,641]" pageId="18" pageNumber="231">
Abdomen (
<figureCitation id="136F2A32F11EFFEE7418F8F955FE7879" box="[317,394,1833,1857]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="18" pageNumber="231">Fig. 44</figureCitation>
) slightly broader than elytra, with deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of distinct non-setiferous punctures, with a lateral puncture on either side, and with six (tergites III and IV) or eight (tergite V) setiferous punctures bearing black setae at or near posterior margin; tergite VI with a narrow transverse band of non-setiferous punctures anteriorly and with two transverse series of setiferous punctures bearing dark setae posteriorly; tergite VII with nonsetiferous punctures anteriorly and with two transverse series of setiferous punctures posteriorly, posterior margin with narrow palisade fringe; tergite VIII with setiferous punctation only in posterior third, posterior margin convex.
</paragraph>
<paragraph id="8BEB36B7F11EFFEE761AFE1A50007D59" blockId="18.[831,1437,233,641]" pageId="18" pageNumber="231">
♂: median lobe of aedeagus
<quantity id="4CAC9B52F11EFFEE71BBFE1950897ED9" box="[1182,1277,457,481]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="231" unit="mm" value="0.75">0.75 mm</quantity>
long; ventral process of distinctive shape, rather broad in ventral view (
<bibRefCitation id="EFC54B46F11EFFEE766CFDD957CF7D19" author="PACE, R." box="[841,955,521,545]" pageId="18" pageNumber="231" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE 2008</bibRefCitation>
: figures 98–99); paramere
<quantity id="4CAC9B52F11EFFEE71D2FDD951227D19" box="[1271,1366,521,545]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="18" pageNumber="231" unit="mm" value="0.85">0.85 mm</quantity>
long, apical lobe rather short, flattened and apically broadly convex to obliquely truncate.
</paragraph>
<paragraph id="8BEB36B7F11EFFEE761AFDBA57BA7DB9" blockId="18.[831,1437,233,641]" box="[831,974,617,641]" pageId="18" pageNumber="231">♀: unknown.</paragraph>
<paragraph id="8BEB36B7F11EFFEE761AFD7950357CD9" blockId="18.[831,1437,681,993]" pageId="18" pageNumber="231">
<emphasis id="B920EAA5F11EFFEE761AFD7950607DF9" bold="true" box="[831,1044,681,705]" pageId="18" pageNumber="231">Comparative notes</emphasis>
: Based on the slender habitus, the slender antennae, and the morphology of the aedeagus,
<taxonomicName id="4C544D34F11EFFEE761AFD3A579D7C39" authorityName="Pace" authorityYear="2008" box="[831,1001,745,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B920EAA5F11EFFEE761AFD3A579D7C39" box="[831,1001,745,769]" italics="true" pageId="18" pageNumber="231">Z. kinabaluensis</emphasis>
</taxonomicName>
is closely allied to
<taxonomicName id="4C544D34F11EFFEE7181FD3A508A7C39" authorityName="CAMERON" authorityYear="1944" box="[1188,1278,746,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B920EAA5F11EFFEE7181FD3A508A7C39" box="[1188,1278,746,769]" italics="true" pageId="18" pageNumber="231">Z. nitens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C544D34F11EFFEE702CFD3A51E27C39" authorityName="Cameron" authorityYear="1943" box="[1289,1430,746,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="B920EAA5F11EFFEE702CFD3A51E27C39" box="[1289,1430,746,769]" italics="true" pageId="18" pageNumber="231">Z. nigerrimus</emphasis>
</taxonomicName>
, and related species. It is readily distinguished from all other
<taxonomicName id="4C544D34F11EFFEE7658FCFA57C37C79" authorityName="STEPHENS" authorityYear="1833" box="[893,951,810,833]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAA5F11EFFEE7658FCFA57C37C79" box="[893,951,810,833]" italics="true" pageId="18" pageNumber="231">Zyras</emphasis>
</taxonomicName>
sensu strictu species recorded from the Sunda Islands by much shorter elytra, completely reduced hind wings, and by the shape of the median lobe of the aedeagus. In addition, it is characterized by the coloration of the legs and the antennae and by the punctation pattern of the abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11EFFEE761AFBD9511E7B79" pageId="18" pageNumber="231" type="distribution">
<paragraph id="8BEB36B7F11EFFEE761AFBD9511E7B79" blockId="18.[831,1437,1033,1089]" pageId="18" pageNumber="231">
<emphasis id="B920EAA5F11EFFEE761AFBD957BC7B19" bold="true" box="[831,968,1033,1057]" pageId="18" pageNumber="231">Distribution</emphasis>
: The
<typeStatus id="54EF8815F11EFFEE7130FBDA50377B1A" box="[1045,1091,1034,1058]" pageId="18" pageNumber="231">type</typeStatus>
locality is situated in Mount
<location id="8E8B606CF11EFFEE761AFBF957D47B79" LSID="urn:lsid:plazi:treatment:03FD87A1F11EFFEE75C6FE63511E7B79:8E8B606CF11EFFEE761AFBF957D47B79" box="[831,928,1065,1089]" country="Malaysia" name="Kinabalu" pageId="18" pageNumber="231" stateProvince="Sabah">Kinabalu</location>
(
<collectingCountry id="F3437627F11EFFEE768BFBF950657B79" box="[942,1041,1065,1089]" name="Malaysia" pageId="18" pageNumber="231">Malaysia</collectingCountry>
:
<collectingRegion id="4990F855F11EFFEE713FFBF950287B79" box="[1050,1116,1065,1089]" country="Malaysia" name="Sabah" pageId="18" pageNumber="231">Sabah</collectingRegion>
) at an altitude of
<quantity id="4CAC9B52F11EFFEE7030FBF951127B79" box="[1301,1382,1065,1089]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="231" unit="m" value="1500.0">
<elevation id="0079D184F11EFFEE7030FBF951127B79" box="[1301,1382,1065,1089]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="231" unit="m" value="1500.0">1500 m</elevation>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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