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<document id="C3E0C0A3328D0D9BB5429ABC1D1EBA65" ID-CLB-Dataset="30315" ID-DOI="10.11646/zootaxa.4407.2.11" ID-GBIF-Dataset="533cfe8e-d57f-44d1-943f-3577ccc0c7e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1216312" ID-ZooBank="E98FFBCA-2CD1-4E86-9AC1-0B13A6A5E5DC" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1523431822613" checkinUser="plazi" docAuthor="Chen, Zhi-Teng" docDate="2018" docId="03B60E030203FF9DFF41A54786B8F815" docLanguage="en" docName="zootaxa.4407.2.11.pdf" docOrigin="Zootaxa 4407 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Podmosta Ricker 1952" docType="treatment" docVersion="7" lastPageNumber="294" masterDocId="FF8F767B0203FF9CFFD6A228865BFFC7" masterDocTitle="Females of the genus Podmosta (Plecoptera: Nemouridae): comparison of terminalia and a new female record in Baltic Amber" masterLastPageNumber="297" masterPageNumber="293" pageNumber="293" updateTime="1698535898913" updateUser="ExternalLinkService">
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<mods:title id="DBF06B4409D42AF327B5EE4EA6A76F7F">Females of the genus Podmosta (Plecoptera: Nemouridae): comparison of terminalia and a new female record in Baltic Amber</mods:title>
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<mods:namePart id="8F73954D0E651D1F4B41413E54596BA1">Chen, Zhi-Teng</mods:namePart>
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<mods:date id="1B7180E48CB2EB4285C93CA5D8A7FD6F">2018</mods:date>
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<treatment id="03B60E030203FF9DFF41A54786B8F815" ID-DOI="http://doi.org/10.5281/zenodo.5986739" ID-GBIF-Taxon="143536436" ID-Zenodo-Dep="5986739" LSID="urn:lsid:plazi:treatment:03B60E030203FF9DFF41A54786B8F815" httpUri="http://treatment.plazi.org/id/03B60E030203FF9DFF41A54786B8F815" lastPageId="1" lastPageNumber="294" pageId="0" pageNumber="293">
<subSubSection id="C305EC9E0203FF9CFF41A547875DF86C" pageId="0" pageNumber="293" type="nomenclature">
<paragraph id="8BA0BF150203FF9CFF41A5478763F84E" blockId="0.[151,312,1903,1963]" box="[151,312,1903,1929]" pageId="0" pageNumber="293">
<heading id="D0E808790203FF9CFF41A5478763F84E" bold="true" box="[151,312,1903,1929]" fontSize="11" level="1" pageId="0" pageNumber="293" reason="1">
<emphasis id="B96B63070203FF9CFF41A5478763F84E" bold="true" box="[151,312,1903,1929]" pageId="0" pageNumber="293">
<taxonomicName id="4C1FC4960203FF9CFF41A5478756F84E" ID-CoL="8MTKM" authorityName="Ricker" authorityYear="1952" box="[151,269,1903,1929]" class="Insecta" family="Nemouridae" genus="Podmosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B96B63070203FF9CFF41A5478756F84E" bold="true" box="[151,269,1903,1929]" italics="true" pageId="0" pageNumber="293">Podmosta</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA0BF150203FF9CFF41A5BA875DF86C" blockId="0.[151,312,1903,1963]" box="[151,262,1938,1963]" pageId="0" pageNumber="293">
<figureCitation id="1324A3900203FF9CFF41A5BA8684F86C" box="[151,223,1938,1963]" captionStart="FIGURE 1" captionStartId="1.[151,250,903,925]" captionText="FIGURE 1. Podmosta sp. Adult habitus, dorsolateral view;" pageId="0" pageNumber="293">Figs. 1</figureCitation>
7.
</paragraph>
</subSubSection>
<subSubSection id="C305EC9E0203FF9DFF41A5F186A8FA71" lastPageId="1" lastPageNumber="294" pageId="0" pageNumber="293" type="description">
<paragraph id="8BA0BF150203FF9CFF41A5F18368F834" blockId="0.[151,1331,2009,2035]" box="[151,1331,2009,2035]" pageId="0" pageNumber="293">
<emphasis id="B96B63070203FF9CFF41A5F18772F835" bold="true" box="[151,297,2009,2034]" pageId="0" pageNumber="293">Description.</emphasis>
Macropterous; body length ca. 5.7 mm, generally yellow and patterned with brown (
<figureCitation id="1324A3900203FF9CFB36A5F28379F834" box="[1248,1314,2010,2035]" captionStart="FIGURE 1" captionStartId="1.[151,250,903,925]" captionText="FIGURE 1. Podmosta sp. Adult habitus, dorsolateral view;" pageId="0" pageNumber="293">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF60EF9D0202FF9DFF41A1AF84AFFC5A" box="[151,756,903,925]" pageId="1" pageNumber="294">
<paragraph id="8BA0BF150202FF9DFF41A1AF84AFFC5A" blockId="1.[151,756,903,925]" box="[151,756,903,925]" pageId="1" pageNumber="294">
<emphasis id="B96B63070202FF9DFF41A1AF8748FC5B" bold="true" box="[151,275,903,925]" pageId="1" pageNumber="294">FIGURE 1.</emphasis>
<taxonomicName id="4C1FC4960202FF9DFECFA1AF8725FC5B" authorityName="Ricker" authorityYear="1952" box="[281,382,903,924]" class="Insecta" family="Nemouridae" genus="Podmosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B96B63070202FF9DFECFA1AF8725FC5B" box="[281,382,903,924]" italics="true" pageId="1" pageNumber="294">Podmosta</emphasis>
</taxonomicName>
sp. Adult habitus, dorsolateral view;
</paragraph>
</caption>
<paragraph id="8BA0BF150202FF9DFF11A1E1874DFBED" blockId="1.[151,1437,969,2002]" pageId="1" pageNumber="294">
Head (
<figureCitation id="1324A3900202FF9DFEC7A1E18704FC25" box="[273,351,969,994]" captionStart="FIGURE 1" captionStartId="1.[151,250,903,925]" captionText="FIGURE 1. Podmosta sp. Adult habitus, dorsolateral view;" pageId="1" pageNumber="294">Figs. 1</figureCitation>
,
<figureCitation id="1324A3900202FF9DFEBBA1E28725FC25" box="[365,382,970,994]" captionStart="FIGURE 2" captionStartId="2.[151,250,1392,1414]" captionTargetBox="[151,1436,219,1355]" captionTargetId="figure@2.[151,1436,219,1355]" captionTargetPageId="2" captionText="FIGURE 2. Podmosta sp. Antennae, lateral view;" httpUri="https://zenodo.org/record/1216314/files/figure.png" pageId="1" pageNumber="294">2</figureCitation>
): Head brown, wider than pronotum. Ocelli pale, compound eyes dark. Antennae filiform and mostly brown, almost equal with body length, with about 50 segments. Maxillary palp short, stout and foursegmented.
</paragraph>
<paragraph id="8BA0BF150202FF9DFF11A61D87F9FAC5" blockId="1.[151,1437,969,2002]" pageId="1" pageNumber="294">
Thorax (
<figureCitation id="1324A3900202FF9DFEF1A61D872FFB89" box="[295,372,1077,1102]" captionStart="FIGURE 1" captionStartId="1.[151,250,903,925]" captionText="FIGURE 1. Podmosta sp. Adult habitus, dorsolateral view;" pageId="1" pageNumber="294">Figs. 1</figureCitation>
,
<figureCitation id="1324A3900202FF9DFE55A61E87C8FB89" box="[387,403,1078,1102]" captionStart="FIGURE 3" captionStartId="2.[151,250,1990,2012]" captionText="FIGURE 3. Podmosta sp. Legs, lateral view;" pageId="1" pageNumber="294">3</figureCitation>
): Pronotum mostly brown, sub-quadrate, with sparse rugosities; cervical gills absent; cervix elongated, laterally with reduced oval nubs. Mesothorax and metathorax sclerotized, with similar length to prothorax. Forelegs shortest, hindlegs longest; femur and tibia mostly pale except for the darker joints; tibia longer and thinner than femur, ventroapically with two giant spurs; tarsus with three segments, second segment shortest, first and third segments subequal in length; claws sharp. Wings with the X-structure; the terminal costal crossvein Sc2 joins the radius R1.
</paragraph>
<paragraph id="8BA0BF150202FF9DFF11A72586A8FA71" blockId="1.[151,1437,969,2002]" pageId="1" pageNumber="294">
Abdomen (
<figureCitation id="1324A3900202FF9DFE9FA72587E1FAE1" box="[329,442,1293,1318]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="2.[151,250,1990,2012]" captionStartId-1="3.[151,250,1272,1294]" captionStartId-2="3.[151,250,2006,2028]" captionText-0="FIGURE 3. Podmosta sp. Legs, lateral view;" captionText-1="FIGURE 4. Podmosta sp. Terminalia, ventrolateral view;" captionText-2="FIGURE 5. Female terminalia of known Podmosta." pageId="1" pageNumber="294">Figs. 35</figureCitation>
): Abdomen near half of the body length; segments 17 pale. Posterior of sternum 7 slightly extended backwards. Sternum 8 indented posteromedially, with a “rabbit-shaped” sclerite, the sclerite with a bilobed anterior tip and widened posteriorly. Sterna 910 darkly sclerotized and shorter than sternum 8. Paraprocts subtriangular with blunt corners. Cerci sclerotized, unmodified and short, approximately twice as long as wide.
</paragraph>
</subSubSection>
<subSubSection id="C305EC9E0202FF9DFF11A7E9857FF9E5" pageId="1" pageNumber="294" type="materials_examined">
<paragraph id="8BA0BF150202FF9DFF11A7E983C6FA39" blockId="1.[151,1437,969,2002]" pageId="1" pageNumber="294">
<materialsCitation id="3B77B5480202FF9DFF11A7E983C6FA39" ID-GBIF-Occurrence="1837803818" collectorName="Chen Amber Collection, No." country="Lithuania" location="Lithuania" pageId="1" pageNumber="294" specimenCount="1">
<emphasis id="B96B63070202FF9DFF11A7E9876DFA1D" bold="true" box="[199,310,1473,1498]" pageId="1" pageNumber="294">Material.</emphasis>
Female, an almost complete specimen in a clear and well-preserved piece of Baltic amber from
<collectingCountry id="F308FF850202FF9DFF41A7CD875EFA39" box="[151,261,1509,1534]" name="Lithuania" pageId="1" pageNumber="294">Lithuania</collectingCountry>
, Eocene (ca. 4050 Mya). The specimen is deposited in the
<collectorName id="26EADAC30202FF9DFC74A7CD82BFFA39" box="[930,1252,1509,1534]" pageId="1" pageNumber="294">Chen Amber Collection, No.</collectorName>
CZT-PLE-BA1.
</materialsCitation>
</paragraph>
<paragraph id="8BA0BF150202FF9DFF11A421857FF9E5" blockId="1.[151,1437,969,2002]" box="[199,804,1545,1570]" pageId="1" pageNumber="294">
<emphasis id="B96B63070202FF9DFF11A421872AF9E5" bold="true" box="[199,369,1545,1570]" pageId="1" pageNumber="294">Syninclusions.</emphasis>
Plant remains and mineral substances.
</paragraph>
</subSubSection>
<subSubSection id="C305EC9E0202FF9DFF11A40686B8F815" pageId="1" pageNumber="294" type="discussion">
<paragraph id="8BA0BF150202FF9DFF11A4068288F885" blockId="1.[151,1437,969,2002]" pageId="1" pageNumber="294">
<emphasis id="B96B63070202FF9DFF11A4068760F981" bold="true" box="[199,315,1582,1606]" pageId="1" pageNumber="294">Remarks.</emphasis>
The new female is assigned to the extant genus
<taxonomicName id="4C1FC4960202FF9DFC89A4068594F981" authorityName="Ricker" authorityYear="1952" box="[863,975,1582,1606]" class="Insecta" family="Nemouridae" genus="Podmosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B96B63070202FF9DFC89A4068594F981" box="[863,975,1582,1606]" italics="true" pageId="1" pageNumber="294">Podmosta</emphasis>
</taxonomicName>
due to the following characters: cervical gills absent; sternum 8 narrow and shorter than sternum7; sternum 8 with a distinct median sclerite; the terminal costal crossvein Sc2 joins the radius R1 (
<bibRefCitation id="EF8EC2E40202FF9DFDA7A45E85D4F949" box="[625,911,1653,1678]" pageId="1" pageNumber="294" refId="ref1060" refString="Caruso, C. E. &amp; Wichard, W. I. (2010) Overview and descriptions of fossil stoneflies (Plecoptera) in Baltic Amber. Entomologie Heute, 22, 85 - 97." type="journal article">Caruso &amp; Wichard 2010</bibRefCitation>
). This specimen differs from females of other extant congeners by the posteromedially indented sternum 8 which has a rabbit-shaped sclerite (
<figureCitation id="1324A3900202FF9DFB2FA4B1831AF975" box="[1273,1345,1689,1714]" captionStart="FIGURE 5" captionStartId="3.[151,250,2006,2028]" captionText="FIGURE 5. Female terminalia of known Podmosta." pageId="1" pageNumber="294">Fig. 5</figureCitation>
). When compared to the two unnamed females proposed by
<bibRefCitation id="EF8EC2E40202FF9DFCD9A496821EF911" box="[783,1093,1725,1750]" pageId="1" pageNumber="294" refId="ref1060" refString="Caruso, C. E. &amp; Wichard, W. I. (2010) Overview and descriptions of fossil stoneflies (Plecoptera) in Baltic Amber. Entomologie Heute, 22, 85 - 97." type="journal article">Caruso &amp; Wichard (2010)</bibRefCitation>
from Baltic amber, the new specimen has a more indented sternum 8, a more modified median sclerite, but has no lateral sclerites (
<figureCitation id="1324A3900202FF9DFA91A4C983D5F93D" box="[1351,1422,1761,1786]" captionStart="FIGURE 5" captionStartId="3.[151,250,2006,2028]" captionText="FIGURE 5. Female terminalia of known Podmosta." pageId="1" pageNumber="294">Fig. 5</figureCitation>
). However, accurate species-level assignment of the new female and determination of the relationship between the new female and the fossil species
<taxonomicName id="4C1FC4960202FF9DFDC2A50285E3F885" authority="Caruso &amp; Wichard, 2010" authorityName="Caruso &amp; Wichard" authorityYear="2010" box="[532,952,1833,1858]" class="Insecta" family="Nemouridae" genus="Podmosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis id="B96B63070202FF9DFDC2A50284C2F886" box="[532,665,1834,1857]" italics="true" pageId="1" pageNumber="294">P. attenuata</emphasis>
<bibRefCitation id="EF8EC2E40202FF9DFD76A50285E3F885" box="[672,952,1833,1858]" pageId="1" pageNumber="294" refId="ref1060" refString="Caruso, C. E. &amp; Wichard, W. I. (2010) Overview and descriptions of fossil stoneflies (Plecoptera) in Baltic Amber. Entomologie Heute, 22, 85 - 97." type="journal article">Caruso &amp; Wichard, 2010</bibRefCitation>
</taxonomicName>
is not currently possible.
</paragraph>
<paragraph id="8BA0BF150202FF9DFF11A56586B8F815" blockId="1.[151,1437,969,2002]" pageId="1" pageNumber="294">
Currently, no extant species of the
<taxonomicName id="4C1FC4960202FF9DFDB3A566848EF8A1" authorityName="Ricker" authorityYear="1952" box="[613,725,1870,1894]" class="Insecta" family="Nemouridae" genus="Podmosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B96B63070202FF9DFDB3A566848EF8A1" box="[613,725,1870,1894]" italics="true" pageId="1" pageNumber="294">Podmosta</emphasis>
</taxonomicName>
has been reported from Europe, the source of Baltic amber. Historical dispersal of this genus may have occurred due to the climate change (
<bibRefCitation id="EF8EC2E40202FF9DFBF9A55A8310F84D" box="[1071,1355,1905,1930]" pageId="1" pageNumber="294" refId="ref1098" refString="Caruso, C. E. &amp; Wichard, W. I. (2011) Paleogeographic Distribution of Leuctridae and Nemouridae genera preserved in Baltic amber, with the description of Palaeopsole weiterschani n. gen., n. sp. (Plecoptera). Entomologie Heute, 23, 69 - 77." type="journal article">Caruso &amp; Wichard 2011</bibRefCitation>
). More morphological and genetic studies are needed to discovery evolutionary trends of stonefly taxa know from Baltic amber.
</paragraph>
</subSubSection>
</treatment>
</document>