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<document id="7CAEA0026A712B82C6BEC9F652124C0E" ID-DOI="10.5252/cryptogamie-bryologie2022v43a2" ID-ISSN="1776-0992" ID-Zenodo-Dep="12215089" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719003595154" checkinUser="felipe" docAuthor="Bruggeman-Nannenga, Maria Alida" docDate="2022" docId="03AB87C7FF80FFA3502BFB9ADAEDFDEE" docLanguage="en" docName="CryptogamieBryologie.43.2.9-36.pdf" docOrigin="Cryptogamie, Bryologie 20 (2)" docSource="http://dx.doi.org/10.5252/cryptogamie-bryologie2022v43a2" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Fissidens lagenarius Mitt." docType="treatment" docVersion="2" lastPageNumber="23" masterDocId="FF92FFBFFF90FFB35110FFCEDE26FF9B" masterDocTitle="On the peristomes of the corticolous African species of Fissidens Hedw. (Fissidentaceae, Bryophyta)" masterLastPageNumber="36" masterPageNumber="9" pageNumber="23" updateTime="1719018102003" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6CF592D3B1F9E603AA2C1B0753D51EAC">On the peristomes of the corticolous African species of Fissidens Hedw. (Fissidentaceae, Bryophyta)</mods:title>
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<mods:namePart id="2975BF8FAD2B55580C0FA3B59419DBA2">Bruggeman-Nannenga, Maria Alida</mods:namePart>
<mods:affiliation id="B76EF20ED50890B6F53F3C7BB6189DC7">Griffensteijnseplein 23 NL- 3703 BE Zeist (The Netherlands)</mods:affiliation>
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<mods:title id="A036FAB32BA7901EA0881AACC4AF1CF4">Cryptogamie, Bryologie</mods:title>
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<mods:date id="7AF46E9F1C682957FA4CE9C5F1087E1E">2022</mods:date>
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<treatment id="03AB87C7FF80FFA3502BFB9ADAEDFDEE" LSID="urn:lsid:plazi:treatment:03AB87C7FF80FFA3502BFB9ADAEDFDEE" httpUri="http://treatment.plazi.org/id/03AB87C7FF80FFA3502BFB9ADAEDFDEE" lastPageNumber="23" pageId="16" pageNumber="23">
<subSubSection id="C318655AFF80FFA3502BFB9ADC77FBF5" box="[315,593,1108,1135]" pageId="16" pageNumber="23" type="nomenclature">
<paragraph id="8BBD36D1FF80FFA3502BFB9ADC77FBF5" blockId="16.[315,593,1108,1135]" box="[315,593,1108,1135]" pageId="16" pageNumber="23">
<heading id="D0F581BDFF80FFA3502BFB9ADC77FBF5" box="[315,593,1108,1135]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="23" reason="4">
<taxonomicName id="4C024D52FF80FFA3502BFB9ADC77FBF5" ID-CoL="6J4MS" authority="Mitt." authorityName="Mitt." box="[315,593,1108,1135]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="16" pageNumber="23" phylum="Bryophyta" rank="species" species="lagenarius">
<emphasis id="B976EAC3FF80FFA3502BFB9ADC36FBF4" bold="true" box="[315,528,1108,1135]" italics="true" pageId="16" pageNumber="23">Fissidens lagenarius</emphasis>
Mitt.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C318655AFF80FFA35086FBBADAEDFDEE" pageId="16" pageNumber="23" type="description">
<paragraph id="8BBD36D1FF80FFA35086FBBADFD3FB15" blockId="16.[406,501,1140,1166]" box="[406,501,1140,1166]" pageId="16" pageNumber="23">
<heading id="D0F581BDFF80FFA35086FBBADFD3FB15" box="[406,501,1140,1166]" centered="true" fontSize="11" level="3" pageId="16" pageNumber="23" reason="3">
(
<figureCitation id="13392A54FF80FFA3508FFBBADFCAFB15" box="[415,492,1140,1166]" captionStart="FIG" captionStartId="17.[132,143,1917,1935]" captionTargetBox="[168,1419,215,1904]" captionTargetId="figure-23@17.[704,1242,780,1349]" captionTargetPageId="17" captionText="FIG. 11. — Fissidens lagenarius Mitt. A, OPL, note the transparent OPL lamellae protruding beyond the narrower underlaying IPL lamellae (left below); B, OPL; C-F, IPL variations: C, IPL of two teeth, each composed of a single column of half cells (reduced situation); D, three teeth focused on the incrassate lamellar plates; in the middle tooth two columns are clearly recognizable (normal situation); E, IPL of three teeth focused on the trabeculae; F, IPL of single tooth; G, side view of tooth. A-D, G from Brazil, São Paulo, Yano 235, SP, private herbarium Bruggeman-Nannenga; E from Zambia, North-western Province, Mushitu Forest, Townsend 75/135, E, private herbarium Bruggeman-Nannenga; F from Brazil, Yano 560 235, SP, private herbarium Bruggeman-Nannenga. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12215123" httpUri="https://zenodo.org/record/12215123/files/figure.png" pageId="16" pageNumber="23">Fig. 11</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BBD36D1FF80FFA35194FB63DC02FAFF" blockId="16.[132,777,1197,1380]" pageId="16" pageNumber="23">
Pantropical, mammillose species with limbidia restricted to the upper leaves of perichaetial stems and ±divided peristomes with a characteristic ornamentation. OPL lamellae in mid teeth with spaced, irregular oblique to vertical ridges in the mid part of the teeth (
<figureCitation id="13392A54FF80FFA351D5FAD9DF1AFAB4" box="[197,316,1303,1327]" captionStart="FIG" captionStartId="17.[132,143,1917,1935]" captionTargetBox="[168,1419,215,1904]" captionTargetId="figure-23@17.[704,1242,780,1349]" captionTargetPageId="17" captionText="FIG. 11. — Fissidens lagenarius Mitt. A, OPL, note the transparent OPL lamellae protruding beyond the narrower underlaying IPL lamellae (left below); B, OPL; C-F, IPL variations: C, IPL of two teeth, each composed of a single column of half cells (reduced situation); D, three teeth focused on the incrassate lamellar plates; in the middle tooth two columns are clearly recognizable (normal situation); E, IPL of three teeth focused on the trabeculae; F, IPL of single tooth; G, side view of tooth. A-D, G from Brazil, São Paulo, Yano 235, SP, private herbarium Bruggeman-Nannenga; E from Zambia, North-western Province, Mushitu Forest, Townsend 75/135, E, private herbarium Bruggeman-Nannenga; F from Brazil, Yano 560 235, SP, private herbarium Bruggeman-Nannenga. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12215123" httpUri="https://zenodo.org/record/12215123/files/figure.png" pageId="16" pageNumber="23">Fig. 11A, B</figureCitation>
). Rare in Africa, known from two collections, both with rather poor peristomes. Part of the above description is therefore based on Neotropical specimens.
</paragraph>
<paragraph id="8BBD36D1FF80FFA35194FA5ADEC0FA36" blockId="16.[132,230,1428,1453]" box="[132,230,1428,1453]" pageId="16" pageNumber="23">
<heading id="D0F581BDFF80FFA35194FA5ADEC0FA36" box="[132,230,1428,1453]" fontSize="11" level="3" pageId="16" pageNumber="23" reason="9">
<emphasis id="B976EAC3FF80FFA35194FA5ADEC0FA36" box="[132,230,1428,1453]" italics="true" pageId="16" pageNumber="23">Peristome</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF80FFA35194FA7DDF83F9D7" blockId="16.[130,775,1459,1613]" pageId="16" pageNumber="23">
±erect when wet, stiff, undivided in the two African specimens (undivided or imperfectly and irregularly divided according
<bibRefCitation id="EF934B20FF80FFA35195FA3DDF31F996" author="PURSELL R. A." box="[133,279,1523,1549]" pageId="16" pageNumber="23" pagination="1 - 278" refId="ref14457" refString="PURSELL R. A. 2007. - Fissidentaceae. Flora Neotropica Monograph 101: 1 - 278." type="journal article" year="2007">Pursell (2007)</bibRefCitation>
in his neotropical monograph, teeth 66-200 µm long, tooth base 22-33 µm wide, distal parts papillose, often broken off or rudimentary.
</paragraph>
<paragraph id="8BBD36D1FF80FFA35194F9BDDF00F916" blockId="16.[132,294,1651,1677]" box="[132,294,1651,1677]" pageId="16" pageNumber="23">
<heading id="D0F581BDFF80FFA35194F9BDDF00F916" box="[132,294,1651,1677]" fontSize="11" level="3" pageId="16" pageNumber="23" reason="9">
<emphasis id="B976EAC3FF80FFA35194F9BDDF00F916" box="[132,294,1651,1677]" italics="true" pageId="16" pageNumber="23">Ornamentation</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF80FFA35194F95CDD21F870" blockId="16.[130,776,1682,2028]" pageId="16" pageNumber="23">
OPL basal ±3 cells with high, smooth trabeculae and thin lamellae that are wider than the IPL-lamellae, in distal direction the trabeculae gradually become indistinguishable, lamellae from about the 4-th cell onwards with an irregular pattern of spaced, vertical to diagonal ridges (
<figureCitation id="13392A54FF80FFA35084F8DCDC2CF8B7" box="[404,522,1810,1836]" captionStart="FIG" captionStartId="17.[132,143,1917,1935]" captionTargetBox="[168,1419,215,1904]" captionTargetId="figure-23@17.[704,1242,780,1349]" captionTargetPageId="17" captionText="FIG. 11. — Fissidens lagenarius Mitt. A, OPL, note the transparent OPL lamellae protruding beyond the narrower underlaying IPL lamellae (left below); B, OPL; C-F, IPL variations: C, IPL of two teeth, each composed of a single column of half cells (reduced situation); D, three teeth focused on the incrassate lamellar plates; in the middle tooth two columns are clearly recognizable (normal situation); E, IPL of three teeth focused on the trabeculae; F, IPL of single tooth; G, side view of tooth. A-D, G from Brazil, São Paulo, Yano 235, SP, private herbarium Bruggeman-Nannenga; E from Zambia, North-western Province, Mushitu Forest, Townsend 75/135, E, private herbarium Bruggeman-Nannenga; F from Brazil, Yano 560 235, SP, private herbarium Bruggeman-Nannenga. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12215123" httpUri="https://zenodo.org/record/12215123/files/figure.png" pageId="16" pageNumber="23">Fig. 11A, B</figureCitation>
), near the apex papillose; IPL in the basal part over a short distance with two columns (
<figureCitation id="13392A54FF80FFA3519DF89FDF37F8F7" box="[141,273,1873,1900]" captionStart="FIG" captionStartId="17.[132,143,1917,1935]" captionTargetBox="[168,1419,215,1904]" captionTargetId="figure-23@17.[704,1242,780,1349]" captionTargetPageId="17" captionText="FIG. 11. — Fissidens lagenarius Mitt. A, OPL, note the transparent OPL lamellae protruding beyond the narrower underlaying IPL lamellae (left below); B, OPL; C-F, IPL variations: C, IPL of two teeth, each composed of a single column of half cells (reduced situation); D, three teeth focused on the incrassate lamellar plates; in the middle tooth two columns are clearly recognizable (normal situation); E, IPL of three teeth focused on the trabeculae; F, IPL of single tooth; G, side view of tooth. A-D, G from Brazil, São Paulo, Yano 235, SP, private herbarium Bruggeman-Nannenga; E from Zambia, North-western Province, Mushitu Forest, Townsend 75/135, E, private herbarium Bruggeman-Nannenga; F from Brazil, Yano 560 235, SP, private herbarium Bruggeman-Nannenga. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12215123" httpUri="https://zenodo.org/record/12215123/files/figure.png" pageId="16" pageNumber="23">Fig. 11D, F</figureCitation>
), sometimes with a single column throughout (
<figureCitation id="13392A54FF80FFA3519DF8BFDEC8F810" box="[141,238,1905,1931]" captionStart="FIG" captionStartId="17.[132,143,1917,1935]" captionTargetBox="[168,1419,215,1904]" captionTargetId="figure-23@17.[704,1242,780,1349]" captionTargetPageId="17" captionText="FIG. 11. — Fissidens lagenarius Mitt. A, OPL, note the transparent OPL lamellae protruding beyond the narrower underlaying IPL lamellae (left below); B, OPL; C-F, IPL variations: C, IPL of two teeth, each composed of a single column of half cells (reduced situation); D, three teeth focused on the incrassate lamellar plates; in the middle tooth two columns are clearly recognizable (normal situation); E, IPL of three teeth focused on the trabeculae; F, IPL of single tooth; G, side view of tooth. A-D, G from Brazil, São Paulo, Yano 235, SP, private herbarium Bruggeman-Nannenga; E from Zambia, North-western Province, Mushitu Forest, Townsend 75/135, E, private herbarium Bruggeman-Nannenga; F from Brazil, Yano 560 235, SP, private herbarium Bruggeman-Nannenga. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12215123" httpUri="https://zenodo.org/record/12215123/files/figure.png" pageId="16" pageNumber="23">Fig. 11C</figureCitation>
); the ±5 basal-most cells with high, papillose trabeculae and ±smooth lamellae, more distal lamellae with close, oblique to horizontal close ridges; filaments papillose on both sides, often broken off; IPL thicker than the OPL (
<figureCitation id="13392A54FF80FFA35389F81FDCDCF870" box="[665,762,2001,2027]" captionStart="FIG" captionStartId="17.[132,143,1917,1935]" captionTargetBox="[168,1419,215,1904]" captionTargetId="figure-23@17.[704,1242,780,1349]" captionTargetPageId="17" captionText="FIG. 11. — Fissidens lagenarius Mitt. A, OPL, note the transparent OPL lamellae protruding beyond the narrower underlaying IPL lamellae (left below); B, OPL; C-F, IPL variations: C, IPL of two teeth, each composed of a single column of half cells (reduced situation); D, three teeth focused on the incrassate lamellar plates; in the middle tooth two columns are clearly recognizable (normal situation); E, IPL of three teeth focused on the trabeculae; F, IPL of single tooth; G, side view of tooth. A-D, G from Brazil, São Paulo, Yano 235, SP, private herbarium Bruggeman-Nannenga; E from Zambia, North-western Province, Mushitu Forest, Townsend 75/135, E, private herbarium Bruggeman-Nannenga; F from Brazil, Yano 560 235, SP, private herbarium Bruggeman-Nannenga. Scale bar: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12215123" httpUri="https://zenodo.org/record/12215123/files/figure.png" pageId="16" pageNumber="23">Fig. 11G</figureCitation>
).
</paragraph>
<paragraph id="8BBD36D1FF80FFA3523DFF19DDBAFF6A" blockId="16.[813,924,215,241]" box="[813,924,215,241]" pageId="16" pageNumber="23">
<heading id="D0F581BDFF80FFA3523DFF19DDBAFF6A" box="[813,924,215,241]" fontSize="11" level="3" pageId="16" pageNumber="23" reason="9">
<emphasis id="B976EAC3FF80FFA3523DFF19DDBAFF6A" box="[813,924,215,241]" italics="true" pageId="16" pageNumber="23">Sporophyte</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF80FFA3523DFF36DDF1FEE9" blockId="16.[810,1456,248,499]" pageId="16" pageNumber="23">
Seta
<quantity id="4CFA9B34FF80FFA35272FF36DDCFFE88" box="[866,1001,248,275]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="16" pageNumber="23" unit="mm" value="1.2" valueMax="1.5" valueMin="0.9">0.9-1.5 mm</quantity>
, smooth; capsule narrow and long, 0.45- 0.6 ×
<quantity id="4CFA9B34FF80FFA35273FED6DDCDFEA8" box="[867,1003,280,307]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="16" pageNumber="23" unit="mm" value="0.225" valueMax="0.25" valueMin="0.2">0.2-0.25mm</quantity>
, exothecial cell columns ±32, the cells oblong, operculum not observed, spores 17.5-20.5 × 16-18.0 µm, finely papillose.
</paragraph>
<paragraph id="8BBD36D1FF80FFA35253FEB7DA22FE68" blockId="16.[810,1456,248,499]" pageId="16" pageNumber="23">
A rather similar peristome is found in the Brazilian
<taxonomicName id="4C024D52FF80FFA35445FEB7DDCFFE2F" authority="Brugg." authorityName="Brugg." class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="16" pageNumber="23" phylum="Bryophyta" rank="species" species="scindulosus">
<emphasis id="B976EAC3FF80FFA35445FEB7DDBDFE28" italics="true" pageId="16" pageNumber="23">Fissidens scindulosus</emphasis>
Brugg.
</taxonomicName>
-Nann. This species differs from
<taxonomicName id="4C024D52FF80FFA35421FE54DB8BFE28" box="[1329,1453,409,435]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="16" pageNumber="23" phylum="Bryophyta" rank="species" species="lagenarius">
<emphasis id="B976EAC3FF80FFA35421FE54DB8BFE28" box="[1329,1453,409,435]" italics="true" pageId="16" pageNumber="23">F.lagenarius</emphasis>
</taxonomicName>
by scindulose cells with 1-3 scindulae. Scattered or in loose mats, hardly mixed.
</paragraph>
<paragraph id="8BBD36D1FF80FFA3523DFDD5DA6FFDAF" blockId="16.[813,1097,538,564]" box="[813,1097,538,564]" pageId="16" pageNumber="23">
<heading id="D0F581BDFF80FFA3523DFDD5DA6FFDAF" box="[813,1097,538,564]" fontSize="11" level="3" pageId="16" pageNumber="23" reason="9">
<emphasis id="B976EAC3FF80FFA3523DFDD5DA6FFDAF" box="[813,1097,538,564]" italics="true" pageId="16" pageNumber="23">Description and illustration</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF80FFA3523DFDF5DAEDFDEE" blockId="16.[813,1454,571,629]" pageId="16" pageNumber="23">
<bibRefCitation id="EF934B20FF80FFA3523DFDF5DD8CFDCE" author="BUCK W. R." box="[813,938,571,597]" pageId="16" pageNumber="23" pagination="1 - 67" refId="ref13841" refString="BUCK W. R. 2003. - Guide to the plants of Central French Guiana part 3. Mosses. Memoirs of the NewYork Botanical Garden 76 (3): 1 - 67." type="journal article" year="2003">Buck (2003</bibRefCitation>
: fig. 10G-I);
<bibRefCitation id="EF934B20FF80FFA35525FDF5DAE5FDCE" author="PURSELL R. A." box="[1077,1219,571,597]" pageId="16" pageNumber="23" pagination="1 - 278" refId="ref14457" refString="PURSELL R. A. 2007. - Fissidentaceae. Flora Neotropica Monograph 101: 1 - 278." type="journal article" year="2007">Pursell (2007</bibRefCitation>
: fig. 94);
<bibRefCitation id="EF934B20FF80FFA35438FDF5DDC2FDEE" author="BRUGGEMAN-NANNENGA M. A." pageId="16" pageNumber="23" pagination="40 - 109" refId="ref13383" refString="BRUGGEMAN-NANNENGA M. A. 2011. - Order Fissidentales, in JANSEN-JACOBS M. J. (ed.), Flora of the Guianas, series C: Bryophytes, fascicle 2. Musci IV, Royal Botanic Gardens, Kew: 40 - 109." type="book chapter" year="2011">Bruggeman-Nannenga (2011</bibRefCitation>
: fig. 26C, E, G, I, J).
</paragraph>
</subSubSection>
</treatment>
</document>
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