311 lines
36 KiB
XML
311 lines
36 KiB
XML
<document id="209D1BA6747D398DF8C9D6E2C8A75035" ID-DOI="10.5281/zenodo.5326990" ID-ISSN="0374-1036" ID-Zenodo-Dep="5326990" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630154894671" checkinUser="felipe" docAuthor="Hájek, Jiří, Hendrich, Lars, Hawlitschek, Oliver & Balke, Michael" docDate="2010" docId="03A487D6FFD7FFE2918DFF6798C91C48" docLanguage="en" docName="ActaEntMusNatPra.2010.50.2.437-443.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 50 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Copelatus sibelaemontis Hájek & Hendrich & Hawlitschek & Balke 2010, sp. nov." docType="treatment" docUuid="49FC9D3B-4AC7-434E-BC7C-69D5322151D5" docUuidSource="ZooBank" docVersion="4" lastPageNumber="442" masterDocId="FF9DFFAEFFD4FFE79028FFBC9A231A6E" masterDocTitle="Copelatus sibelaemontis sp. nov. (Coleoptera: Dytiscidae) from the Moluccas with generic assignment based on morphology and DNA sequence data" masterLastPageNumber="443" masterPageNumber="437" pageNumber="440" updateTime="1699166922473" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E5D389125F0B4820E8731C0ECEBC0F25">Copelatus sibelaemontis sp. nov. (Coleoptera: Dytiscidae) from the Moluccas with generic assignment based on morphology and DNA sequence data</mods:title>
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<mods:name id="C3B57661BE7F5D699AB3DB8BE27D1027" type="personal">
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<mods:roleTerm id="9D8218008A6D77DB84E3C54F186D251E">Author</mods:roleTerm>
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<mods:namePart id="257AF973DA5565CDA7B1C26E21045E2D">Hájek, Jiří</mods:namePart>
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<mods:affiliation id="586916B627A672DA14F5A66A86D8A8CB">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: jiri _ hajek @ nm. cz</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="662D80C3C30A40C1A69F1C9227D649EA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="864676CB506052C59C344E407C33F1AB">Hendrich, Lars</mods:namePart>
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<mods:affiliation id="866B65BF5B364C245792930479B078E6">Zoologische Staatssammlung, Münchhausenstrasse 21, D- 81247 München, Germany; e-mail: coleoptera-zsm @ zsm. mwn. de</mods:affiliation>
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</mods:name>
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<mods:name id="A0683C70CDF041B0D7F8DB6112597083" type="personal">
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<mods:roleTerm id="9BE61ACB2ECD974103D0A9CC62F1D8D3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="06C31F37163845FF428F77E1F2B0241E">Hawlitschek, Oliver</mods:namePart>
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<mods:affiliation id="EAF8AC43B4D84CA9E4E5191A8AAABFC5">Zoologische Staatssammlung, Münchhausenstrasse 21, D- 81247 München, Germany; e-mail: coleoptera-zsm @ zsm. mwn. de</mods:affiliation>
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<mods:roleTerm id="0A18FC8A47CA2E82FA45A7E140A99E2D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="31B4DF6C03E10E19529D01D74B2F26E6">Balke, Michael</mods:namePart>
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<mods:affiliation id="617AE2288B4C5569706477183D78DD72">Zoologische Staatssammlung, Münchhausenstrasse 21, D- 81247 München, Germany; e-mail: coleoptera-zsm @ zsm. mwn. de & GeoBioCenter, Ludwig-Maximilians-University, Richard-Wagner-Strasse 10, D- 80333 München, Germany</mods:affiliation>
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<mods:typeOfResource id="7954797594798E5A525675D7BA19519A">text</mods:typeOfResource>
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<mods:title id="E43B52E7246B8712E16EB0C0635A2824">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="7F063E49760E05BA65C81DA1ADFED1A1">
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<mods:date id="872D3874F90064525463BD0EC4DD782E">2010</mods:date>
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<mods:number id="9E153D9B5116FAAC34334797CE08F380">2010-12-15</mods:number>
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<mods:number id="4E8F87B6788CDB51734751698D2E7D86">50</mods:number>
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<mods:classification id="15A8BBAF222AD6515D90943F1A6027DC">journal article</mods:classification>
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<mods:identifier id="611B5B734CBE1C9987CEBF63BD742DD9" type="DOI">10.5281/zenodo.5326990</mods:identifier>
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<treatment id="03A487D6FFD7FFE2918DFF6798C91C48" LSID="urn:lsid:zoobank.org:act:49FC9D3B-4AC7-434E-BC7C-69D5322151D5" httpUri="http://treatment.plazi.org/id/03A487D6FFD7FFE2918DFF6798C91C48" lastPageId="5" lastPageNumber="442" pageId="3" pageNumber="440">
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<subSubSection id="C317654BFFD7FFE4918DFF67992E1A9D" box="[421,781,219,243]" pageId="3" pageNumber="440" type="nomenclature">
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<paragraph id="8BB236C0FFD7FFE4918DFF67992E1A9D" blockId="3.[421,781,219,243]" box="[421,781,219,243]" pageId="3" pageNumber="440">
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<heading id="D0FA81ACFFD7FFE4918DFF67992E1A9D" bold="true" box="[421,781,219,243]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="440" reason="3">
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<emphasis id="B979EAD2FFD7FFE4918DFF67992E1A9D" bold="true" box="[421,781,219,243]" pageId="3" pageNumber="440">
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<taxonomicName id="4C0D4D43FFD7FFE4918DFF6798931A9D" ID-CoL="Y3FR" authority="Hájek & Hendrich & Hawlitschek & Balke, 2010" authorityName="Hájek & Hendrich & Hawlitschek & Balke" authorityYear="2010" box="[421,688,219,243]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="sibelaemontis" status="sp. nov.">
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<emphasis id="B979EAD2FFD7FFE4918DFF6798931A9D" bold="true" box="[421,688,219,243]" italics="true" pageId="3" pageNumber="440">Copelatus sibelaemontis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24A57A9FFD7FFE4929EFF67992E1A9D" box="[694,781,219,243]" pageId="3" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C317654BFFD7FFE49202FF45999A1B3F" pageId="3" pageNumber="440" type="description">
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<paragraph id="8BB236C0FFD7FFE49202FF4598AB1B62" blockId="3.[554,648,249,268]" box="[554,648,249,268]" pageId="3" pageNumber="440">
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(
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<figureCitation id="13362A45FFD7FFE49218FF4598A21B62" box="[560,641,249,268]" captionStart="Figs" captionStartId="4.[151,190,1548,1567]" captionTargetBox="[202,996,227,1475]" captionTargetId="figure-10@4.[151,1026,219,1483]" captionTargetPageId="4" captionText="Figs. 1–4. Copelatus sibelaemontis sp. nov. 1 – habitus; 2 – median lobe in ventral view; 3 – the same in lateral view; 4 – lateral lobe (paramere)." figureDoi="http://doi.org/10.5281/zenodo.5326992" httpUri="https://zenodo.org/record/5326992/files/figure.png" pageId="3" pageNumber="440">Figs. 1–4</figureCitation>
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)
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</paragraph>
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<paragraph id="8BB236C0FFD7FFE49059FE9F99431B58" blockId="3.[113,953,290,337]" box="[113,864,290,310]" pageId="3" pageNumber="440">
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<emphasis id="B979EAD2FFD7FFE49059FE9F9B411B58" bold="true" box="[113,354,290,310]" pageId="3" pageNumber="440">WWW site on wikispecies.</emphasis>
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http://species.wikimedia.org/wiki/
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<taxonomicName id="4C0D4D43FFD7FFE492A0FE9F98FD1B58" authorityName="Erichson" authorityYear="1832" box="[648,734,291,310]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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||
_sibelaemontis
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||
</paragraph>
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<paragraph id="8BB236C0FFD7FFE49059FE81999A1B3F" blockId="3.[113,953,290,337]" box="[113,953,317,337]" pageId="3" pageNumber="440">
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<emphasis id="B979EAD2FFD7FFE49059FE819B631B3F" bold="true" box="[113,320,317,337]" pageId="3" pageNumber="440">Life Science Identifier.</emphasis>
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<uri id="FF9C3AC2FFD7FFE4916DFE82999A1B3F" box="[325,953,318,337]" pageId="3" pageNumber="440">
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urn:lsid:zoobank.org:act:
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<uuid id="FFAB0C15FFD7FFE4923EFE82999A1B3F" box="[534,953,318,337]" pageId="3" pageNumber="440">49FC9D3B-4AC7-434E-BC7C-69D5322151D5</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C317654BFFD7FFE49059FED49B401805" pageId="3" pageNumber="440" type="materials_examined">
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<paragraph id="8BB236C0FFD7FFE49059FED499701BFB" blockId="3.[113,1090,359,620]" pageId="3" pageNumber="440">
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<emphasis id="B979EAD2FFD7FFE49059FED49AC81B15" bold="true" box="[113,235,359,379]" pageId="3" pageNumber="440">
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<typeStatus id="54B68862FFD7FFE49059FED49ABC1B15" box="[113,159,360,379]" pageId="3" pageNumber="440">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F31A7650FFD7FFE490C6FED49B671B15" box="[238,324,360,379]" name="Indonesia" pageId="3" pageNumber="440">Indonesia</collectingCountry>
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:Northern
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<collectingRegion id="49C9F822FFD7FFE491B3FED49BFC1B15" box="[411,479,360,379]" country="Indonesia" name="Maluku Utara" pageId="3" pageNumber="440">Maluku</collectingRegion>
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||
, Bacan Island, ca.
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<quantity id="4CF59B25FFD7FFE49250FEDB98821B15" box="[632,673,359,379]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="440" unit="km" value="5.0">5 km</quantity>
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SE of Makian village, SE slopes of Mount Sibela,
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<quantity id="4CF59B25FFD7FFE49059FE3E9AF11BF8" box="[113,210,386,406]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="3" pageNumber="440" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">500–750 m</quantity>
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a.s.l. (approximate position from GoogleEarth:
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<geoCoordinate id="EE395007FFD7FFE49245FE3E98E81BFB" box="[621,715,385,405]" degrees="0" direction="south" minutes="44.388" orientation="latitude" pageId="3" pageNumber="440" precision="1" value="-0.73980004">0°44.388′S</geoCoordinate>
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<geoCoordinate id="EE395007FFD7FFE492F8FE3E996B1BFB" box="[720,840,385,405]" degrees="127" direction="east" minutes="33.984" orientation="longitude" pageId="3" pageNumber="440" precision="1" value="127.5664">127°33.984′E</geoCoordinate>
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).
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</paragraph>
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<paragraph id="8BB236C0FFD7FFE49059FE2198F11874" blockId="3.[113,1090,359,620]" pageId="3" pageNumber="440">
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<emphasis id="B979EAD2FFD7FFE49059FE219AD61BDE" bold="true" box="[113,245,412,432]" pageId="3" pageNumber="440">Type material.</emphasis>
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<typeStatus id="54B68862FFD7FFE490D1FE219B681BC1" box="[249,331,413,432]" pageId="3" pageNumber="440" type="holotype">HOLOTYPE</typeStatus>
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: J (
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<collectionCode id="ED1CAE05FFD7FFE49147FE219B921BDE" box="[367,433,413,432]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="440">NMPC</collectionCode>
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): ‘
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<collectingCountry id="F31A7650FFD7FFE491E8FE2198191BDE" box="[448,570,413,432]" name="Indonesia" pageId="3" pageNumber="440">INDONESIA</collectingCountry>
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, N
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<collectingCountry id="F31A7650FFD7FFE4927CFE21988F1BC1" box="[596,684,413,432]" name="Indonesia" pageId="3" pageNumber="440">MOLUCCAS</collectingCountry>
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/ Bacan Isl.,
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<quantity id="4CF59B25FFD7FFE4933EFE2199521BDE" box="[790,881,413,432]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="3" pageNumber="440" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">500-750 m</quantity>
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/ SE slopes of Mt. Sibela /
|
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<quantity id="4CF59B25FFD7FFE49055FE0B9A8B1BA5" box="[125,168,439,459]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="440" unit="km" value="5.0">5 km</quantity>
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SE of MAKIAN vill. / S. Jákl leg.,
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<date id="FFB31000FFD7FFE491E0FE04981B1BA5" box="[456,568,440,459]" pageId="3" pageNumber="440" value="2008-05-02" valueMax="2008-05-12" valueMin="2008-05-02">
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2.-12.
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<collectionCode id="ED1CAE05FFD7FFE491DCFE0498261BA5" box="[500,517,440,459]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="440" type="Museum">V</collectionCode>
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.2008
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</date>
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[printed] //
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<typeStatus id="54B68862FFD7FFE49288FE0499321BA5" box="[672,785,440,459]" pageId="3" pageNumber="440" type="holotype">HOLOTYPE</typeStatus>
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/
|
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<taxonomicName id="4C0D4D43FFD7FFE49309FE0499BF1BA5" authorityName="Erichson" authorityYear="1832" box="[801,924,440,459]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="genus">COPELATUS</taxonomicName>
|
||
/ sibelaemontis
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||
<taxonomicNameLabel id="A24A57A9FFD7FFE49400FE049ABB1B88" pageId="3" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
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/ J. Hájek et al. des. 2010 [red label, printed]’.
|
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<typeStatus id="54B68862FFD7FFE49203FE6F98A01B8A" box="[555,643,467,486]" pageId="3" pageNumber="440" type="paratype">PARATYPES</typeStatus>
|
||
: 8 JJ
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||
<specimenCount id="9D0BFD49FFD7FFE492ECFE6E98D01B88" box="[708,755,463,486]" pageId="3" pageNumber="440" type="female">4 ♀♀</specimenCount>
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, same label data as
|
||
<typeStatus id="54B68862FFD7FFE493B6FE6E99CA1B8B" box="[926,1001,466,485]" pageId="3" pageNumber="440" type="holotype">holotype</typeStatus>
|
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(
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||
<collectionCode id="ED1CAE05FFD7FFE493DCFE6E9E1E1B8B" box="[1012,1085,466,485]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="440" type="Museum">NHMW</collectionCode>
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||
,
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||
<collectionCode id="ED1CAE05FFD7FFE49059FE519A93186E" box="[113,176,493,512]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="440">NMPC</collectionCode>
|
||
,
|
||
<collectionCode id="ED1CAE05FFD7FFE4909FFE519AD0186E" box="[183,243,493,512]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="3" pageNumber="440">SMNS</collectionCode>
|
||
,
|
||
<collectionCode id="ED1CAE05FFD7FFE490D2FE519B1B186E" box="[250,312,493,512]" country="Germany" httpUri="http://grbio.org/cool/a6ms-py1z" name="Zoologische Staatssammlung" pageId="3" pageNumber="440">ZSMC</collectionCode>
|
||
). Each
|
||
<typeStatus id="54B68862FFD7FFE4915BFE519B9F186E" box="[371,444,493,512]" pageId="3" pageNumber="440" type="paratype">paratype</typeStatus>
|
||
is provided with the respective red printed label.One male
|
||
<typeStatus id="54B68862FFD7FFE49383FE5199D7186E" box="[939,1012,493,512]" pageId="3" pageNumber="440" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED1CAE05FFD7FFE493D6FE519E18186E" box="[1022,1083,493,512]" country="Germany" httpUri="http://grbio.org/cool/a6ms-py1z" name="Zoologische Staatssammlung" pageId="3" pageNumber="440">ZSMC</collectionCode>
|
||
) bears a green label with M. Balke’s DNA extraction number
|
||
<collectionCode id="ED1CAE05FFD7FFE49252FDB498B81875" box="[634,667,520,539]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="3" pageNumber="440">MB</collectionCode>
|
||
3456.
|
||
</paragraph>
|
||
<paragraph id="8BB236C0FFD7FFE49059FD9D9B401805" blockId="3.[113,1090,359,620]" pageId="3" pageNumber="440">
|
||
<emphasis id="B979EAD2FFD7FFE49059FD9D9BAA185B" bold="true" box="[113,393,545,565]" pageId="3" pageNumber="440">Additional material examined.</emphasis>
|
||
1 J
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||
<specimenCount id="9D0BFD49FFD7FFE4919DFD9E9BF51858" box="[437,470,543,566]" pageId="3" pageNumber="440" type="female">1 ♀</specimenCount>
|
||
, ‘
|
||
<collectingRegion id="49C9F822FFD7FFE491CDFD9E986A185B" box="[485,585,546,565]" country="Indonesia" name="Maluku Utara" pageId="3" pageNumber="440">MALUKU</collectingRegion>
|
||
:Is.Morotai / W Daruba, Raja /
|
||
<date id="FFB31000FFD7FFE4937DFD9E999A185B" box="[853,953,546,565]" pageId="3" pageNumber="440" value="1999-11-18">18.XI.1999</date>
|
||
, 50-
|
||
<quantity id="4CF59B25FFD7FFE493F6FD9E9E2C185B" box="[990,1039,546,565]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="440" unit="m" value="100.0">100m</quantity>
|
||
/ leg. A.RIEDEL’ (
|
||
<collectionCode id="ED1CAE05FFD7FFE490CEFD819B00183E" box="[230,291,573,592]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="3" pageNumber="440">SMNS</collectionCode>
|
||
,
|
||
<collectionCode id="ED1CAE05FFD7FFE49106FD819B4F183E" box="[302,364,573,592]" country="Germany" httpUri="http://grbio.org/cool/a6ms-py1z" name="Zoologische Staatssammlung" pageId="3" pageNumber="440">ZSMC</collectionCode>
|
||
). The specimens bear green labels with M. Balke’s DNA extraction numbers,
|
||
<collectionCode id="ED1CAE05FFD7FFE49437FD819E63183E" box="[1055,1088,573,592]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="3" pageNumber="440">MB</collectionCode>
|
||
3151 (J) and
|
||
<collectionCode id="ED1CAE05FFD7FFE490C1FDE49B291805" box="[233,266,600,619]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="3" pageNumber="440">MB</collectionCode>
|
||
3152 (
|
||
<specimenCount id="9D0BFD49FFD7FFE4916EFDE99B751802" box="[326,342,597,620]" pageId="3" pageNumber="440" type="female">♀</specimenCount>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317654BFFD7FFE29059FD3F9E081E68" lastPageId="5" lastPageNumber="442" pageId="3" pageNumber="440" type="description">
|
||
<paragraph id="8BB236C0FFD7FFE49059FD3F9BB51895" blockId="3.[113,1090,643,1595]" pageId="3" pageNumber="440">
|
||
<emphasis id="B979EAD2FFD7FFE49059FD3F9ADF18F5" bold="true" box="[113,252,643,667]" pageId="3" pageNumber="440">Description.</emphasis>
|
||
Body oblong-oval, broadest in basal third of elytra, moderately convex. Head relatively broad; clypeus rounded. Pronotum broadest between posterior angles, lateral margins moderately curved. Base of elytra as broad as pronotal base; lateral margins of elytra moderately curved (
|
||
<figureCitation id="13362A45FFD7FFE4916FFD5F9BA51895" box="[327,390,739,763]" captionStart="Figs" captionStartId="4.[151,190,1548,1567]" captionTargetBox="[202,996,227,1475]" captionTargetId="figure-10@4.[151,1026,219,1483]" captionTargetPageId="4" captionText="Figs. 1–4. Copelatus sibelaemontis sp. nov. 1 – habitus; 2 – median lobe in ventral view; 3 – the same in lateral view; 4 – lateral lobe (paramere)." figureDoi="http://doi.org/10.5281/zenodo.5326992" httpUri="https://zenodo.org/record/5326992/files/figure.png" pageId="3" pageNumber="440">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236C0FFD7FFE490B8FCB89B131935" blockId="3.[113,1090,643,1595]" pageId="3" pageNumber="440">
|
||
Measurements. Bacan: TL: 4.6–5.0 mm (
|
||
<typeStatus id="54B68862FFD7FFE4926CFCBF98801975" box="[580,675,771,795]" pageId="3" pageNumber="440" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CF59B25FFD7FFE49281FCBF98DD1975" box="[681,766,771,795]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="440" unit="mm" value="4.7">4.7 mm</quantity>
|
||
), TL-h:
|
||
<quantity id="4CF59B25FFD7FFE4937CFCBF99F71975" box="[852,980,771,795]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" pageId="3" pageNumber="440" unit="mm" value="4.3" valueMax="4.5" valueMin="4.1">4.1–4.5 mm</quantity>
|
||
(
|
||
<typeStatus id="54B68862FFD7FFE493CAFCBF9E621975" box="[994,1089,771,795]" pageId="3" pageNumber="440" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CF59B25FFD7FFE49059FC9F9AEA1955" box="[113,201,803,827]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="440" unit="mm" value="4.3">4.3 mm</quantity>
|
||
), TW:
|
||
<quantity id="4CF59B25FFD7FFE49130FC9F9BBA1955" box="[280,409,803,827]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="3" pageNumber="440" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
|
||
(
|
||
<typeStatus id="54B68862FFD7FFE49182FC9F982A1955" box="[426,521,803,827]" pageId="3" pageNumber="440" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CF59B25FFD7FFE4923AFC9F98491955" box="[530,618,803,827]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="440" unit="mm" value="2.3">2.3 mm</quantity>
|
||
). Morotai: TL:
|
||
<quantity id="4CF59B25FFD7FFE4933EFC9F994F1955" box="[790,876,803,827]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="440" unit="mm" value="4.6">4.6 mm</quantity>
|
||
, Tl-h:
|
||
<quantity id="4CF59B25FFD7FFE49390FC9F9E1F1955" box="[952,1084,803,827]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="3" pageNumber="440" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.1–4.2 mm</quantity>
|
||
, TW:
|
||
<quantity id="4CF59B25FFD7FFE49081FCFF9B0E1935" box="[169,301,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="3" pageNumber="440" unit="mm" value="2.25" valueMax="2.3" valueMin="2.2">2.2–2.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB236C0FFD7FFE490B8FCDF997019F5" blockId="3.[113,1090,643,1595]" pageId="3" pageNumber="440">Colouration. Body colour dark brown, head in front of eyes, sides of pronotum, appendages, and basal transverse band on elytra ferrugineous. Beetle rather shiny.</paragraph>
|
||
<paragraph id="8BB236C0FFD7FFE490B8FC1F9BE71E55" blockId="3.[113,1090,643,1595]" pageId="3" pageNumber="440">Surface sculpture. Head uniformly microreticulated, reticulation composed of moderately deeply impressed isodiametric meshes. Punctation composed of coarse setigerous punctures, and very small punctures spread sparsely on surface; rows of coarse punctures presented around inner margin of eyes and in small depression antero-laterally of eyes. Antenna with antennomeres long and slender.</paragraph>
|
||
<paragraph id="8BB236C0FFD7FFE490B8FBF898EF1ED5" blockId="3.[113,1090,643,1595]" pageId="3" pageNumber="440">Pronotum with lateral beading very thin and indistinct. Microreticulation similar to that of head. Punctation similar to that of head; row of coarse setigerous punctures presented along anterior margin, basal margin (except for baso-medially), and laterally close to sides. Indistinct longitudinal wrinkles presented baso-laterally.</paragraph>
|
||
<paragraph id="8BB236C0FFD7FFE490B8FB7F9ACA1F75" blockId="3.[113,1090,643,1595]" pageId="3" pageNumber="440">Elytra without striae or strioles, microreticulation similar to that of head and pronotum. Punctation similar to that of head; coarse setigerous punctures form four indistinct longitudinal rows.</paragraph>
|
||
<paragraph id="8BB236C0FFD7FFE490B8FA9F984E1F95" blockId="3.[113,1090,643,1595]" pageId="3" pageNumber="440">Ventral part. Finely microreticulated, with intermixed sparsely distributed very small punctures. Meshes isodiametric, except for metacoxae and abdominal ventrite I (longitudinal), abdominal ventrite II (diagonal), and abdominal ventrites III–IV (transverse). Prosternum obtusely keeled medially. Prosternal process lanceolate, bordered except for tip. Metepisterna (‘metasternal wings’) tongue-shaped, slender. Metacoxae and abdominal ventrites I–II with numerous striae, possibly used for stridulation. Indistinct transverse rows of large setigerous punctures are presented on abdominal sternites.</paragraph>
|
||
<paragraph id="8BB236C0FFD7FFE290B8F9BF9B8E1988" blockId="3.[113,1090,643,1595]" lastBlockId="5.[113,1089,910,1574]" lastPageId="5" lastPageNumber="442" pageId="3" pageNumber="440">
|
||
Male. Protarsomeres and mesotarsomeres 1–3 distinctly broadened, with adhesive discs on their ventral side. Median lobe of aedeagus slightly asymmetrical in ventral view, apically bifid; inner lobe well developed (
|
||
<figureCitation id="13362A45FFD1FFE291F5FC32980219C8" box="[477,545,910,934]" captionStart="Figs" captionStartId="4.[151,190,1548,1567]" captionTargetBox="[202,996,227,1475]" captionTargetId="figure-10@4.[151,1026,219,1483]" captionTargetPageId="4" captionText="Figs. 1–4. Copelatus sibelaemontis sp. nov. 1 – habitus; 2 – median lobe in ventral view; 3 – the same in lateral view; 4 – lateral lobe (paramere)." figureDoi="http://doi.org/10.5281/zenodo.5326992" httpUri="https://zenodo.org/record/5326992/files/figure.png" pageId="5" pageNumber="442">Fig. 2</figureCitation>
|
||
). In lateral view, median lobe broadest in middle; apex bent down (
|
||
<figureCitation id="13362A45FFD1FFE2910EFC129B4519A8" box="[294,358,942,966]" captionStart="Figs" captionStartId="4.[151,190,1548,1567]" captionTargetBox="[202,996,227,1475]" captionTargetId="figure-10@4.[151,1026,219,1483]" captionTargetPageId="4" captionText="Figs. 1–4. Copelatus sibelaemontis sp. nov. 1 – habitus; 2 – median lobe in ventral view; 3 – the same in lateral view; 4 – lateral lobe (paramere)." figureDoi="http://doi.org/10.5281/zenodo.5326992" httpUri="https://zenodo.org/record/5326992/files/figure.png" pageId="5" pageNumber="442">Fig. 3</figureCitation>
|
||
). Lateral lobe with broader base narrowing towards apex, setation on inner margin distinct (
|
||
<figureCitation id="13362A45FFD1FFE29176FC729B831988" box="[350,416,974,998]" captionStart="Figs" captionStartId="4.[151,190,1548,1567]" captionTargetBox="[202,996,227,1475]" captionTargetId="figure-10@4.[151,1026,219,1483]" captionTargetPageId="4" captionText="Figs. 1–4. Copelatus sibelaemontis sp. nov. 1 – habitus; 2 – median lobe in ventral view; 3 – the same in lateral view; 4 – lateral lobe (paramere)." figureDoi="http://doi.org/10.5281/zenodo.5326992" httpUri="https://zenodo.org/record/5326992/files/figure.png" pageId="5" pageNumber="442">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF726648FFD0FFE390BFF9B09B961C54" ID-DOI="http://doi.org/10.5281/zenodo.5326992" ID-Zenodo-Dep="5326992" httpUri="https://zenodo.org/record/5326992/files/figure.png" pageId="4" pageNumber="441" startId="4.[151,190,1548,1567]" targetBox="[202,996,227,1475]" targetPageId="4">
|
||
<paragraph id="8BB236C0FFD0FFE390BFF9B09B961C54" blockId="4.[151,1128,1548,1594]" pageId="4" pageNumber="441">
|
||
Figs. 1–4.
|
||
<taxonomicName id="4C0D4D43FFD0FFE390DCF9B19BE51C71" authority="Hájek & Hendrich & Hawlitschek & Balke, 2010" authorityName="Hájek & Hendrich & Hawlitschek & Balke" authorityYear="2010" box="[244,454,1549,1567]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="sibelaemontis" status="sp. nov.">
|
||
<emphasis id="B979EAD2FFD0FFE390DCF9B19BE51C71" box="[244,454,1549,1567]" italics="true" pageId="4" pageNumber="441">Copelatus sibelaemontis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24A57A9FFD0FFE391E5F9B098311C71" box="[461,530,1548,1567]" pageId="4" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
|
||
1 – habitus; 2 – median lobe in ventral view; 3 – the same in lateral view; 4 – lateral lobe (paramere).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF726648FFD1FFE2905BFC9A986D1957" ID-DOI="http://doi.org/10.5281/zenodo.5326994" ID-Zenodo-Dep="5326994" box="[115,590,806,825]" httpUri="https://zenodo.org/record/5326994/files/figure.png" pageId="5" pageNumber="442" startId="5.[115,146,806,825]" targetBox="[185,1017,219,776]" targetPageId="5">
|
||
<paragraph id="8BB236C0FFD1FFE2905BFC9A986D1957" blockId="5.[115,590,806,825]" box="[115,590,806,825]" pageId="5" pageNumber="442">
|
||
Fig. 5. Distribution of
|
||
<taxonomicName id="4C0D4D43FFD1FFE2911BFC9B98271957" authority="Hájek & Hendrich & Hawlitschek & Balke, 2010" authorityName="Hájek & Hendrich & Hawlitschek & Balke" authorityYear="2010" box="[307,516,807,825]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="sibelaemontis" status="sp. nov.">
|
||
<emphasis id="B979EAD2FFD1FFE2911BFC9B98271957" box="[307,516,807,825]" italics="true" pageId="5" pageNumber="442">Copelatus sibelaemontis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24A57A9FFD1FFE29222FC9A986D1957" box="[522,590,806,825]" pageId="5" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB236C0FFD1FFE290B8FC529E081E68" blockId="5.[113,1089,910,1574]" box="[144,1067,1006,1030]" pageId="5" pageNumber="442">Female. Similar to male in habitus. Protarsomeres and mesotarsomeres not broadened.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317654BFFD1FFE29059FBB29BA81E28" pageId="5" pageNumber="442" type="diagnosis">
|
||
<paragraph id="8BB236C0FFD1FFE29059FBB29BA81E28" blockId="5.[113,1089,910,1574]" pageId="5" pageNumber="442">
|
||
<emphasis id="B979EAD2FFD1FFE29059FBB29B4F1E48" bold="true" box="[113,364,1038,1062]" pageId="5" pageNumber="442">Differential diagnosis.</emphasis>
|
||
Readily characterized by the smooth elytra combined with shape of male genitalia (
|
||
<figureCitation id="13362A45FFD1FFE2913EFB929B5D1E28" box="[278,382,1070,1094]" captionStart="Figs" captionStartId="4.[151,190,1548,1567]" captionTargetBox="[202,996,227,1475]" captionTargetId="figure-10@4.[151,1026,219,1483]" captionTargetPageId="4" captionText="Figs. 1–4. Copelatus sibelaemontis sp. nov. 1 – habitus; 2 – median lobe in ventral view; 3 – the same in lateral view; 4 – lateral lobe (paramere)." figureDoi="http://doi.org/10.5281/zenodo.5326992" httpUri="https://zenodo.org/record/5326992/files/figure.png" pageId="5" pageNumber="442">Figs. 2–4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317654BFFD1FFE29059FBF298D11F68" pageId="5" pageNumber="442" type="etymology">
|
||
<paragraph id="8BB236C0FFD1FFE29059FBF298AF1EE8" blockId="5.[113,1089,910,1574]" pageId="5" pageNumber="442">
|
||
<emphasis id="B979EAD2FFD1FFE29059FBF29AD01E08" bold="true" box="[113,243,1102,1126]" pageId="5" pageNumber="442">Etymology.</emphasis>
|
||
The new species is named after the
|
||
<typeStatus id="54B68862FFD1FFE2925AFBF298831E08" box="[626,672,1102,1126]" pageId="5" pageNumber="442">type</typeStatus>
|
||
locality – Sibela, and Latin word
|
||
<emphasis id="B979EAD2FFD1FFE29420FBF39E631E08" box="[1032,1088,1103,1126]" italics="true" pageId="5" pageNumber="442">mons</emphasis>
|
||
(genitive
|
||
<emphasis id="B979EAD2FFD1FFE290FFFBD39B3D1EE8" box="[215,286,1135,1158]" italics="true" pageId="5" pageNumber="442">montis</emphasis>
|
||
, masculinum) meaning mountain.
|
||
</paragraph>
|
||
<paragraph id="8BB236C0FFD1FFE29059FB3298D11F68" blockId="5.[113,1089,910,1574]" pageId="5" pageNumber="442">
|
||
<emphasis id="B979EAD2FFD1FFE29059FB329BB91EC8" bold="true" box="[113,410,1166,1190]" pageId="5" pageNumber="442">Collection circumstances.</emphasis>
|
||
Specimens from Bacan were collected in a small stream (S. Jákl, pers. comm.). The species was associated in both localities with
|
||
<taxonomicName id="4C0D4D43FFD1FFE29373FB139B471E88" authority="J. Balfour-Browne, 1939" authorityName="J. Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="wallacei">
|
||
<emphasis id="B979EAD2FFD1FFE29373FB139E0B1EA8" box="[859,1064,1199,1222]" italics="true" pageId="5" pageNumber="442">Copelatus wallacei</emphasis>
|
||
J. Balfour-Browne, 1939
|
||
</taxonomicName>
|
||
, and on Bacan also with
|
||
<taxonomicName id="4C0D4D43FFD1FFE2924FFB7399921E88" authority="Regimbart, 1899" authorityName="Regimbart" authorityYear="1899" box="[615,945,1230,1254]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="ternatensis">
|
||
<emphasis id="B979EAD2FFD1FFE2924FFB7398D91E88" box="[615,762,1231,1254]" italics="true" pageId="5" pageNumber="442">C. ternatensis</emphasis>
|
||
Régimbart, 1899
|
||
</taxonomicName>
|
||
and a species from the
|
||
<taxonomicName id="4C0D4D43FFD1FFE290FBFB5398AB1F68" authority="Regimbart, 1899" authorityName="Regimbart" authorityYear="1899" box="[211,648,1262,1286]" class="Insecta" family="Dytiscidae" genus="Platynectes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="decastigma">
|
||
<emphasis id="B979EAD2FFD1FFE290FBFB539BED1F68" box="[211,462,1263,1286]" italics="true" pageId="5" pageNumber="442">Platynectes decastigma</emphasis>
|
||
Régimbart, 1899
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317654BFFD1FFE29059FAB298C91C48" pageId="5" pageNumber="442" type="distribution">
|
||
<paragraph id="8BB236C0FFD1FFE29059FAB298951F28" blockId="5.[113,1089,910,1574]" pageId="5" pageNumber="442">
|
||
<emphasis id="B979EAD2FFD1FFE29059FAB29B271F48" bold="true" box="[113,260,1294,1318]" pageId="5" pageNumber="442">Distribution.</emphasis>
|
||
So far known from Bacan and Morotai Islands of Northern
|
||
<collectingRegion id="49C9F822FFD1FFE29352FAB299F31F48" box="[890,976,1294,1318]" country="Indonesia" name="Maluku Utara" pageId="5" pageNumber="442">Maluku</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31A7650FFD1FFE293F1FAB29E621F48" box="[985,1089,1294,1318]" name="Indonesia" pageId="5" pageNumber="442">Indonesia</collectingCountry>
|
||
(
|
||
<figureCitation id="13362A45FFD1FFE29051FA929A991F28" box="[121,186,1326,1350]" captionStart="Fig" captionStartId="5.[115,146,806,825]" captionTargetBox="[185,1017,219,776]" captionTargetId="figure-260@5.[184,1017,218,776]" captionTargetPageId="5" captionText="Fig. 5. Distribution of Copelatus sibelaemontis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5326994" httpUri="https://zenodo.org/record/5326994/files/figure.png" pageId="5" pageNumber="442">Fig. 5</figureCitation>
|
||
). The occurrence in Halmahera Island is likely.
|
||
</paragraph>
|
||
<paragraph id="8BB236C0FFD1FFE29059FAF298C91C48" blockId="5.[113,1089,910,1574]" pageId="5" pageNumber="442">
|
||
<emphasis id="B979EAD2FFD1FFE29059FAF29A9B1F08" bold="true" box="[113,184,1358,1382]" pageId="5" pageNumber="442">Notes.</emphasis>
|
||
Specimens from Morotai and Bacan show> 4%
|
||
<emphasis id="B979EAD2FFD1FFE29294FAF398CD1F08" box="[700,750,1359,1382]" italics="true" pageId="5" pageNumber="442">cox1</emphasis>
|
||
divergence, suggestive of interrupted gene flow. This, together with their insular distribution, could be taken as indicators of ongoing speciation. The occurrence of local endemism would be typical of running water species, usually with limited dispersal abilities (
|
||
<bibRefCitation id="EF9C4B31FFD1FFE29244FA12990C1FA8" author="RIBERA I. & BARRACLOUGH T. G. & VOGLER A. P." box="[620,815,1454,1478]" pageId="5" pageNumber="442" pagination="721 - 735" refId="ref3176" refString="RIBERA I., BARRACLOUGH T. G. & VOGLER A. P. 2001: The effect of habitat type on speciation rates and range movements in aquatic beetles: inferences from species-level phylogenies. Molecular Ecology 10: 721 - 735." type="journal article" year="2001">RIBERA et al. 2001</bibRefCitation>
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), although in this case the lack of apparent morphological differentiation between the specimens from two islands about
|
||
<quantity id="4CF59B25FFD1FFE29059FA529AE21C68" box="[113,193,1518,1542]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="442" unit="km" value="300.0">300 km</quantity>
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distant from each other is noteworthy. We suggest to collect more material and study population-level processes in this very diverse archipelago.
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||
</paragraph>
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||
</subSubSection>
|
||
</treatment>
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||
</document> |