714 lines
96 KiB
XML
714 lines
96 KiB
XML
<document id="9D14C268FBC598BF0D16E15AF8BB821D" ID-CLB-Dataset="46386" ID-DOI="10.5281/zenodo.205720" ID-GBIF-Dataset="8bff639d-baa3-4392-845a-e7aa97502f8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="205720" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460387095378" checkinUser="plazi" docAuthor="Kok, Philippe J. R." docDate="2011" docId="03A43A081D57FFF6B2F7EB93FCEAFAC8" docLanguage="en" docName="zt03018p012.pdf" docOrigin="Zootaxa 3018" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gonatodes timidus Kok, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="9" masterDocId="FF9D42701D56FFFEB260EE36FFADFFA5" masterDocTitle="A new species of the genus Gonatodes Fitzinger, 1843 (Reptilia: Sphaerodactylidae) from central Guyana, northern South America" masterLastPageNumber="12" masterPageNumber="1" pageNumber="2" updateTime="1698297759683" updateUser="ExternalLinkService">
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<mods:mods id="20DC940EFBBC01FE86160728B97D3AEC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EDF956908860E53FA4FA27C3CB32EB09">
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<mods:title id="D9D88DF5A9030619E0F56358830AE197">A new species of the genus Gonatodes Fitzinger, 1843 (Reptilia: Sphaerodactylidae) from central Guyana, northern South America</mods:title>
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</mods:titleInfo>
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<mods:name id="484B45874C633151C9C2CBEC070F8698" type="personal">
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<mods:roleTerm id="76A0B70B96BAE28AC7A9D22ABBF802D3">Author</mods:roleTerm>
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<mods:namePart id="05D87B956159C9640AA6A5566E131CA3">Kok, Philippe J. R.</mods:namePart>
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<mods:typeOfResource id="4372AA410996CF91F0A6384D8167D89E">text</mods:typeOfResource>
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<mods:title id="B376E8FA7C724067904A0A4B8724685C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F0988B9AAAE58E65C17404116F0423F8">
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<mods:date id="BB888844FD7C40FB72646714EA21E3BE">2011</mods:date>
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<mods:detail id="3789D39A0B0BD3D3A80527777A66861E" type="volume">
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<mods:number id="F67ADC5A0D8B6F2F38A2A65180E032BC">3018</mods:number>
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<mods:start id="A3DAD0A8F82BFFD76827B4EF3A506DB5">1</mods:start>
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<mods:classification id="E65CB65630503D131D48124E0122C75D">journal article</mods:classification>
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<mods:identifier id="661B594DAF48AD5ECEBE04D5EC1CAB9A" type="CLB-Dataset">46386</mods:identifier>
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<mods:identifier id="4C4543C45B7B57ABBFB33D3A196FFEB7" type="DOI">10.5281/zenodo.205720</mods:identifier>
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<mods:identifier id="83A5B5529EC4854F4BA892666B868838" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="295AD5767A3D2DE9C27E8FD6D07AB044" type="Zenodo-Dep">205720</mods:identifier>
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<treatment id="03A43A081D57FFF6B2F7EB93FCEAFAC8" ID-DOI="http://doi.org/10.5281/zenodo.5683198" ID-GBIF-Taxon="119415824" ID-Zenodo-Dep="5683198" LSID="urn:lsid:plazi:treatment:03A43A081D57FFF6B2F7EB93FCEAFAC8" httpUri="http://treatment.plazi.org/id/03A43A081D57FFF6B2F7EB93FCEAFAC8" lastPageId="8" lastPageNumber="9" pageId="1" pageNumber="2">
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<subSubSection id="C317D8951D57FFFFB2F7EB93FEDEFA44" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="8BB28B1E1D57FFFFB2F7EB93FE62FA18" blockId="1.[151,463,1445,1505]" box="[151,463,1445,1470]" pageId="1" pageNumber="2">
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<heading id="D0FA3C721D57FFFFB2F7EB93FE62FA18" bold="true" box="[151,463,1445,1470]" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
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<emphasis id="B979570C1D57FFFFB2F7EB93FE62FA18" bold="true" box="[151,463,1445,1470]" pageId="1" pageNumber="2">
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<taxonomicName id="4C0DF09D1D57FFFFB2F7EB93FEC3FA1B" box="[151,366,1445,1470]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="timidus" status="sp. nov.">
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<emphasis id="B979570C1D57FFFFB2F7EB93FEC3FA1B" bold="true" box="[151,366,1445,1470]" italics="true" pageId="1" pageNumber="2">Gonatodes timidus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24AEA771D57FFFFB315EB93FE62FA18" box="[373,463,1445,1469]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB28B1E1D57FFFFB2F7EBFFFEDEFA44" blockId="1.[151,463,1445,1505]" box="[151,371,1481,1505]" pageId="1" pageNumber="2">
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(
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<figureCitation id="1336979B1D57FFFFB2FFEBFFFEA7FA44" box="[159,266,1481,1505]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1930,1953]" captionStartId-1="4.[151,250,1904,1927]" captionStartId-2="5.[151,250,1963,1986]" captionStartId-3="6.[151,250,662,685]" captionTargetBox-0="[151,1436,561,1883]" captionTargetBox-1="[239,1352,193,1863]" captionTargetBox-2="[351,1231,193,1923]" captionTargetBox-3="[151,1436,193,630]" captionTargetId-0="figure@2.[151,1436,561,1884]" captionTargetId-1="figure@4.[239,1354,193,1864]" captionTargetId-2="figure@5.[351,1232,193,1924]" captionTargetId-3="figure@6.[151,1436,193,631]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Gonatodes timidus sp. nov. in life. Top: Male holotype (IRSNB 2664, 46.9 mm SVL). Middle: Male paratype (IRSNB 2665, 46.7 mm SVL). Bottom: Female paratype (IRSNB 2667, 51.2 mm SVL). Note the conspicuous difference in the colour of head ornamentation between the two males, and the sexual dichromatism." captionText-1="FIGURE 2. Gonatodes timidus sp. nov. Sexual dichromatism in life. Top left: Ventral face of male holotype (IRSNB 2664, 46.9 mm SVL). Lower left: Detail of the gular region in the same specimen. Top right: Ventral face of female paratype (IRSNB 2670, 51.6 mm SVL). Lower right: Detail of the gular region in the same specimen. Scale bars = 10 mm (top), and 5 mm (bottom)." captionText-2="FIGURE 3. Gonatodes timidus sp. nov. Details of lateral (top), dorsal (middle), and ventral (bottom) views of the head of the male holotype in life (IRSNB 2664, 46.9 mm SVL). Scale bar = 5 mm." captionText-3="FIGURE 4. Gonatodes timidus sp. nov. Detail of pericloacal region in preserved specimens. A: Male holotype (IRSNB 2664), note distinct cluster (lower belly) and rows (thighs) of escutcheon scales. B: Female paratype (IRSNB 2670)." httpUri-0="https://zenodo.org/record/205721/files/figure.png" httpUri-1="https://zenodo.org/record/205722/files/figure.png" httpUri-2="https://zenodo.org/record/205723/files/figure.png" httpUri-3="https://zenodo.org/record/205724/files/figure.png" pageId="1" pageNumber="2">Figs. 1–4</figureCitation>
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,
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<tableCitation id="C68FBEA51D57FFFFB376EBFFFECAFA44" box="[278,359,1481,1505]" captionStart="TABLE 1" captionStartId="7.[151,235,1187,1208]" captionTargetBox="[151,1435,1266,1727]" captionTargetPageId="7" captionText="TABLE 1. Morphometric measurements (in mm) of the type series of Gonatodes timidus sp. nov. M = male, F = female, JUV = juvenile, other abbreviations are defined in the text." httpUri="http://table.plazi.org/id/DF72DB961D51FFF9B2F7EA95FD1AFB72" pageId="1" pageNumber="2" tableUuid="DF72DB961D51FFF9B2F7EA95FD1AFB72">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C317D8951D57FFFFB2F7E824FBB9F880" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="8BB28B1E1D57FFFFB2F7E824FF43F9D4" blockId="1.[151,1436,1553,2009]" pageId="1" pageNumber="2">
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<emphasis id="B979570C1D57FFFFB2F7E824FEA6F98F" bold="true" box="[151,267,1554,1578]" pageId="1" pageNumber="2">
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<typeStatus id="54B635BC1D57FFFFB2F7E824FEABF98F" box="[151,262,1554,1578]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED1C13DB1D57FFFFB372E827FEC9F98C" box="[274,356,1553,1577]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="2">IRSNB</collectionCode>
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2664 (field number PK 3555), an adult male collected by Philippe J. R. Kok,
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<date id="FFB3ADDE1D57FFFFB6B5E827FA2DF98C" box="[1237,1408,1553,1577]" pageId="1" pageNumber="2" value="2011-03-22">22 March 2011</date>
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at 15h00, base of Iwokrama Mountains, Potaro-Siparuni District,
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<collectingCountry id="F31ACB8E1D57FFFFB13DE803FC19F9E8" box="[861,948,1589,1613]" name="Guyana" pageId="1" pageNumber="2">Guyana</collectingCountry>
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(
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<geoCoordinate id="EE39EDD91D57FFFFB1A4E803FBF2F9E8" box="[964,1119,1589,1613]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="15" value="4.331111">04° 19’ 52”N</geoCoordinate>
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,
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<geoCoordinate id="EE39EDD91D57FFFFB60BE803FABAF9E8" box="[1131,1303,1589,1613]" direction="west" orientation="longitude" pageId="1" pageNumber="2" precision="15" value="-58.799442">058° 47’ 58”W</geoCoordinate>
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,
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<quantity id="4CF526FB1D57FFFFB743E803FAC6F9E8" box="[1315,1387,1589,1613]" metricMagnitude="2" metricUnit="m" metricValue="2.09" pageId="1" pageNumber="2" unit="m" value="209.0">209 m</quantity>
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elevation).
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</paragraph>
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<paragraph id="8BB28B1E1D57FFFFB2A5E84BFBB9F880" blockId="1.[151,1436,1553,2009]" pageId="1" pageNumber="2">
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<typeStatus id="54B635BC1D57FFFFB2A5E84BFE90F930" box="[197,317,1661,1685]" pageId="1" pageNumber="2" type="paratype">
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<emphasis id="B979570C1D57FFFFB2A5E84BFE90F930" bold="true" box="[197,317,1661,1685]" pageId="1" pageNumber="2">Paratypes</emphasis>
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</typeStatus>
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(n=9). two adult males (
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<collectionCode id="ED1C13DB1D57FFFFB03EE84BFD1FF930" box="[606,690,1661,1685]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="2">IRSNB</collectionCode>
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<date id="FFB3ADDE1D57FFFFB0DBE84BFC8DF930" box="[699,800,1661,1685]" pageId="1" pageNumber="2" value="2665" valueMax="2666">2665–66</date>
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, field numbers PK 3538, PK 3557), four adult females (
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<collectionCode id="ED1C13DB1D57FFFFB2FEE897FF5FF91C" box="[158,242,1697,1721]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="2">IRSNB</collectionCode>
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<date id="FFB3ADDE1D57FFFFB299E897FEF3F91C" box="[249,350,1697,1721]" pageId="1" pageNumber="2" value="2667" valueMax="2670">2667–70</date>
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, field numbers PK 3539–41, PK 3556), two juveniles (
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<collectionCode id="ED1C13DB1D57FFFFB1A1E897FBB8F91C" box="[961,1045,1697,1721]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="2">IRSNB</collectionCode>
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<date id="FFB3ADDE1D57FFFFB67BE897FB2DF91C" box="[1051,1152,1697,1721]" pageId="1" pageNumber="2" value="2671" valueMax="2672">2671–72</date>
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, field numbers PK 3522– 23) with same data as
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<typeStatus id="54B635BC1D57FFFFB3EEE8F3FE5EF978" box="[398,499,1733,1757]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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, collected between
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<date id="FFB3ADDE1D57FFFFB0ACE8F3FC1AF978" box="[716,951,1733,1757]" pageId="1" pageNumber="2" value="2011-03-19" valueMax="2011-03-23" valueMin="2011-03-19">19 to 23 March 2011</date>
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, and one juvenile (
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<collectionCode id="ED1C13DB1D57FFFFB6E7E8F3FB76F978" box="[1159,1243,1733,1757]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="2">IRSNB</collectionCode>
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2673, field number PK 3553) collected by M. Wilkinson,
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<date id="FFB3ADDE1D57FFFFB008E8DFFCBCF8A4" box="[616,785,1769,1793]" pageId="1" pageNumber="2" value="2011-03-21">
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21 March
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<quantity id="4CF526FB1D57FFFFB0BAE8DFFCBCF8A4" box="[730,785,1769,1793]" metricMagnitude="1" metricUnit="m" metricValue="5.10794" pageId="1" pageNumber="2" unit="in" value="2011.0">2011</quantity>
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</date>
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in early afternoon, base of Iwokrama Mountains, opposite side of river of Tortoise camp (
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<geoCoordinate id="EE39EDD91D57FFFFB392E93BFD21F880" box="[498,652,1805,1829]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="15" value="4.33">04° 19’ 48”N</geoCoordinate>
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,
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<geoCoordinate id="EE39EDD91D57FFFFB0F8E93BFCEEF880" box="[664,835,1805,1829]" direction="west" orientation="longitude" pageId="1" pageNumber="2" precision="15" value="-58.798332">058° 47’ 54”W</geoCoordinate>
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,
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<quantity id="4CF526FB1D57FFFFB12FE93BFC3BF880" box="[847,918,1805,1829]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="1" pageNumber="2" unit="m" value="225.0">225 m</quantity>
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elevation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C317D8951D57FFFDB2A5E904FC32FC92" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="etymology">
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<paragraph id="8BB28B1E1D57FFFFB2A5E904FC0EF834" blockId="1.[151,1436,1553,2009]" pageId="1" pageNumber="2">
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<emphasis id="B979570C1D57FFFFB2A5E904FEE6F8EF" bold="true" box="[197,331,1842,1866]" pageId="1" pageNumber="2">Etymology.</emphasis>
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The specific name is considered a noun in apposition and is derived from the Latin adjective
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<taxonomicName id="4C0DF09D1D57FFFFB70FE904FF6BF8C8" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="timidus">
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<emphasis id="B979570C1D57FFFFB70FE904FF6BF8C8" italics="true" pageId="1" pageNumber="2">timidus</emphasis>
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</taxonomicName>
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meaning “shy” or “fearful”. It refers to the tendency of individuals of the new species to avoid to be seen by hastily escaping between rocks, making them very difficult to collect.
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</paragraph>
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<paragraph id="8BB28B1E1D57FFFCB2A5E9ABFC4EFE51" blockId="1.[151,1436,1553,2009]" lastBlockId="2.[151,1436,151,500]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<emphasis id="B979570C1D57FFFFB2A5E9ABFE42F813" bold="true" box="[197,495,1949,1974]" pageId="1" pageNumber="2">Definition and diagnosis.</emphasis>
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The new species differs from all known congeners by the following unique combination of characters: (1) moderate body size, with adult males ranging from 46.0–
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<quantity id="4CF526FB1D57FFFFB64AE9F7FB20F87C" box="[1066,1165,1985,2009]" metricMagnitude="-2" metricUnit="m" metricValue="4.6899999999999995" pageId="1" pageNumber="2" unit="mm" value="46.9">46.9 mm</quantity>
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SVL, and adult females from
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<quantity id="4CF526FB1D54FFFCB2B4EEA1FEDBFF0A" box="[212,374,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="4.785" metricValueMax="5.16" metricValueMin="4.41" pageId="2" pageNumber="3" unit="mm" value="47.85" valueMax="51.6" valueMin="44.1">44.1–51.6 mm</quantity>
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SVL; (2) absence of a clearly differentiated elongate supraciliary spine; (3) absence of clusters of distinctly enlarged conical scales on sides; (4) 89–97 scales around midbody; (5) 46–50 ventral scales counted in a longitudinal row; (6) escutcheon scales on posterior belly and ventral surfaces of thighs in males evident; (7) three to four lateral rows of scales on distal part of fingers and toes; (8) medial subcaudal scales distinctly differentiated from adjacent scales on non-regenerated tail, the pattern of scales consisting in repetitive series of a single midventral scale in contact laterodistally with one scale per side followed by a single midventral in contact with two scales per side (1’1”
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<emphasis id="B979570C1D54FFFCB3DAEF47FE55FE2D" box="[442,504,369,392]" italics="true" pageId="2" pageNumber="3">sensu</emphasis>
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<bibRefCitation id="EF9CF6EF1D54FFFCB061EF59FD68FE22" author="Avila-Pires" box="[513,709,367,391]" pageId="2" pageNumber="3" refString="Avila-Pires, T. S. C. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">Ávila-Pires 1995</bibRefCitation>
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||
); (9) sexual dichromatism obvious, males with upper surface of head black with bluish white to vivid yellow irregular stripes and blotches, and underside of head orange, females lack any colourful head ornamentation, but usually have two thin, black-edged white crescent-shaped collars on the neck region, and the underside of head black with bluish white reticulation.
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</paragraph>
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||
<caption id="DF72DB961D54FFFCB2F7E9BCFC4BF87B" httpUri="https://zenodo.org/record/205721/files/figure.png" pageId="2" pageNumber="3" targetBox="[151,1436,561,1883]" targetPageId="2">
|
||
<paragraph id="8BB28B1E1D54FFFCB2F7E9BCFC4BF87B" blockId="2.[151,1436,1930,2014]" pageId="2" pageNumber="3">
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<emphasis id="B979570C1D54FFFCB2F7E9BCFEB8F805" bold="true" box="[151,277,1930,1953]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
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<taxonomicName id="4C0DF09D1D54FFFCB37DE9BDFE72F804" box="[285,479,1931,1953]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="timidus" status="sp. nov.">
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<emphasis id="B979570C1D54FFFCB37DE9BDFE72F804" box="[285,479,1931,1953]" italics="true" pageId="2" pageNumber="3">Gonatodes timidus</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A24AEA771D54FFFCB387E9BDFD97F805" box="[487,570,1931,1952]" pageId="2" pageNumber="3" rank="species">
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<emphasis id="B979570C1D54FFFCB387E9BDFD97F805" bold="true" box="[487,570,1931,1952]" pageId="2" pageNumber="3">sp. nov.</emphasis>
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||
</taxonomicNameLabel>
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in life. Top: Male holotype (IRSNB 2664, 46.9 mm SVL). Middle: Male paratype (IRSNB 2665, 46.7 mm SVL). Bottom: Female paratype (IRSNB 2667, 51.2 mm SVL). Note the conspicuous difference in the colour of head ornamentation between the two males, and the sexual dichromatism.
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||
</paragraph>
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</caption>
|
||
<paragraph id="8BB28B1E1D55FFFDB2A5EEA1FEBAFD21" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="4">
|
||
Compared to the six
|
||
<taxonomicName id="4C0DF09D1D55FFFDB3CDEEAEFD8BFF15" box="[429,550,152,176]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="B979570C1D55FFFDB3CDEEAEFD8BFF15" box="[429,550,152,176]" italics="true" pageId="3" pageNumber="4">Gonatodes</emphasis>
|
||
</taxonomicName>
|
||
species known to occur in the
|
||
<collectingCountry id="F31ACB8E1D55FFFDB11EEEA1FC62FF0A" box="[894,975,151,175]" name="Guyana" pageId="3" pageNumber="4">Guiana</collectingCountry>
|
||
Shield,
|
||
<taxonomicName id="4C0DF09D1D55FFFDB64BEEAEFB53FF15" box="[1067,1278,152,176]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B979570C1D55FFFDB64BEEAEFB53FF15" box="[1067,1278,152,176]" italics="true" pageId="3" pageNumber="4">Gonatodes timidus</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished by its radically different male pattern and colouration. It further differs from
|
||
<taxonomicName id="4C0DF09D1D55FFFDB614EE8BFE92FF52" authority="Cole and Kok, 2010" authorityName="Cole and Kok" authorityYear="2010" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="alexandermendesi">
|
||
<emphasis id="B979570C1D55FFFDB614EE8BFAF2FF71" box="[1140,1375,188,212]" italics="true" pageId="3" pageNumber="4">G. alexandermendesi</emphasis>
|
||
Cole and Kok, 2010
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0DF09D1D55FFFDB32BEED6FA35FF52" authority="Schargel, Rivas, Makowsky, Senaris, Natera, Barros, Molina and Barrio-Amoros, 2010" authorityName="Schargel, Rivas, Makowsky, Senaris, Natera, Barros, Molina and Barrio-Amoros" authorityYear="2010" box="[331,1432,223,248]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="astralis">
|
||
<emphasis id="B979570C1D55FFFDB32BEED6FE10FF5D" box="[331,445,224,248]" italics="true" pageId="3" pageNumber="4">G. astralis</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D55FFFDB3A4EEE9FA35FF52" author="Schargel" box="[452,1432,223,247]" pageId="3" pageNumber="4" refString="Schargel, W. E., Rivas, G. A., Makowsky, R., Senaris, J. C., Natera, M. A., Barros, T. R., Molina, C. R. & Barrio-Amoros, C. L. (2010) Phylogenetic systematics of the genus Gonatodes (Squamata: Sphaerodactylidae) in the Guayana region, with description of a new species from Venezuela. Systematics and Biodiversity, 8, 321 - 339." type="journal article" year="2010">Schargel, Rivas, Makowsky, Señaris, Natera, Barros, Molina and Barrio-Amorós, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0DF09D1D55FFFDB2A8EF33FCE2FEB9" authority="Barrio-Amoros and Brewer-Carias, 2008" authorityName="Barrio-Amoros and Brewer-Carias" authorityYear="2008" box="[200,847,260,284]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="superciliaris">
|
||
<emphasis id="B979570C1D55FFFDB2A8EF33FED9FEB9" box="[200,372,260,284]" italics="true" pageId="3" pageNumber="4">G. superciliaris</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D55FFFDB31EEF32FCE2FEB9" author="Barrio-Amoros" box="[382,847,260,284]" pageId="3" pageNumber="4" refString="Barrio-Amoros, C. L. & Brewer-Carias, C. (2008) Herpetological results of the 2002 expedition to Sarisarinama, a tepui in Venezuelan Guayana, with the description of five new species. Zootaxa, 1942, 1 - 68." type="journal article" year="2008">Barrio-Amorós and Brewer-Carias, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
by the absence of a distinctly elongate supraciliary spine (present in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB336EF1EFDECFEE5" box="[342,577,296,320]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="alexandermendesi">
|
||
<emphasis id="B979570C1D55FFFDB336EF1EFDECFEE5" box="[342,577,296,320]" italics="true" pageId="3" pageNumber="4">G. alexandermendesi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0DF09D1D55FFFDB02EEF1EFD6CFEE5" box="[590,705,296,320]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="astralis">
|
||
<emphasis id="B979570C1D55FFFDB02EEF1EFD6CFEE5" box="[590,705,296,320]" italics="true" pageId="3" pageNumber="4">G. astralis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DF09D1D55FFFDB097EF1EFC0CFEE5" box="[759,929,296,320]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="superciliaris">
|
||
<emphasis id="B979570C1D55FFFDB097EF1EFC0CFEE5" box="[759,929,296,320]" italics="true" pageId="3" pageNumber="4">G. superciliaris</emphasis>
|
||
</taxonomicName>
|
||
), from
|
||
<taxonomicName id="4C0DF09D1D55FFFDB196EF1EFAEDFE9A" authority="Boulenger, 1887" authorityName="Boulenger" authorityYear="1887" box="[1014,1344,295,320]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="annularis">
|
||
<emphasis id="B979570C1D55FFFDB196EF1EFB2CFEE5" box="[1014,1153,296,320]" italics="true" pageId="3" pageNumber="4">G. annularis</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D55FFFDB6E7EF11FAEDFE9A" author="Boulenger" box="[1159,1344,295,319]" pageId="3" pageNumber="4" refString="Boulenger, G. A. (1887) On a new geckoid lizard from British Guiana. Proceedings of the Zoological Society of London, 1887, 153 - 154." type="proceedings paper" year="1887">Boulenger, 1887</bibRefCitation>
|
||
</taxonomicName>
|
||
by having the medial subcaudal scales distinctly differentiated from adjacent scales on non-regenerated tail (not distinctly differentiated in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB330EF46FE74FE2D" box="[336,473,368,392]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="annularis">
|
||
<emphasis id="B979570C1D55FFFDB330EF46FE74FE2D" box="[336,473,368,392]" italics="true" pageId="3" pageNumber="4">G. annularis</emphasis>
|
||
</taxonomicName>
|
||
) and a dark copper coloured iris in males (always blue in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB63FEF46FB45FE2D" box="[1119,1256,368,392]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="annularis">
|
||
<emphasis id="B979570C1D55FFFDB63FEF46FB45FE2D" box="[1119,1256,368,392]" italics="true" pageId="3" pageNumber="4">G. annularis</emphasis>
|
||
</taxonomicName>
|
||
males), from
|
||
<taxonomicName id="4C0DF09D1D55FFFDB7E4EF46FD81FE09" authority="Rivas and Schargel, 2008" authorityName="Rivas and Schargel" authorityYear="2008" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="infernalis">
|
||
<emphasis id="B979570C1D55FFFDB7E4EF46FEAEFE09" italics="true" pageId="3" pageNumber="4">G. infernalis</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D55FFFDB36AEFA2FD81FE09" author="Rivas" box="[266,556,404,428]" pageId="3" pageNumber="4" refString="Rivas, G. A. & Schargel, W. E. (2008) Gecko on the rocks: an enigmatic new species of Gonatodes (Sphaerodactylidae) from Inselbergs of the Venezuelan Guayana. Zootaxa, 1925, 39 - 50." type="journal article" year="2008">Rivas and Schargel, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
by its smaller size in female (maximum
|
||
<quantity id="4CF526FB1D55FFFDB661EFA2FBCAFE09" box="[1025,1127,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="5.16" pageId="3" pageNumber="4" unit="mm" value="51.6">51.6 mm</quantity>
|
||
in
|
||
<emphasis id="B979570C1D55FFFDB6EDEFA3FA8BFE09" box="[1165,1318,404,428]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C0DF09D1D55FFFDB6EDEFA3FB52FE09" box="[1165,1279,404,428]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="timidus">G. timidus</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
minimum
|
||
<quantity id="4CF526FB1D55FFFDB2F7EF81FF51FE6A" box="[151,252,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="5.56" pageId="3" pageNumber="4" unit="mm" value="55.6">55.6 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB342EF8EFE03FE75" box="[290,430,440,464]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="infernalis">
|
||
<emphasis id="B979570C1D55FFFDB342EF8EFE03FE75" box="[290,430,440,464]" italics="true" pageId="3" pageNumber="4">G. infernalis</emphasis>
|
||
</taxonomicName>
|
||
), the presence of markings on head and body in females (absence of dorsal pattern in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB7E4EF8EFEAFFE51" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="infernalis">
|
||
<emphasis id="B979570C1D55FFFDB7E4EF8EFEAFFE51" italics="true" pageId="3" pageNumber="4">G. infernalis</emphasis>
|
||
</taxonomicName>
|
||
), and in having the throat black with bluish white irregular stripes and blotches in females (immaculate in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB2F7EC36FE8CFDBD" box="[151,289,512,536]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="infernalis">
|
||
<emphasis id="B979570C1D55FFFDB2F7EC36FE8CFDBD" box="[151,289,512,536]" italics="true" pageId="3" pageNumber="4">G. infernalis</emphasis>
|
||
</taxonomicName>
|
||
), from
|
||
<emphasis id="B979570C1D55FFFDB314EC36FE53FDBD" box="[372,510,512,536]" italics="true" pageId="3" pageNumber="4">G. h u m e r a l i s</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9CF6EF1D55FFFDB070EFC9FD60FDB2" author="Guichenot" box="[528,717,511,535]" pageId="3" pageNumber="4" refString="Guichenot, A. (1855) Reptiles. In: Castelnau F. (Ed), Animaux Nouveaux ou Rares Recueillis pendant l'Expedition dans les Parties Centrales de l'Amerique du Sud, de Rio de Janeiro a Lima, et de Lima au Para; Executee par Ordre du Gouvernement Francais pendant les Annees 1843 a 1847, sous la Direction du Comte Francis de Castelnau. Reptiles. Tome second. P. Bertrand, Paris, pp. 1 - 95." type="book chapter" year="1855">Guichenot, 1855</bibRefCitation>
|
||
) by its larger size in females (minimum
|
||
<quantity id="4CF526FB1D55FFFDB6EFEFC9FB5EFDB2" box="[1167,1267,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="4.41" pageId="3" pageNumber="4" unit="mm" value="44.1">44.1 mm</quantity>
|
||
SVL in
|
||
<emphasis id="B979570C1D55FFFDB731EC36FF46FD99" italics="true" pageId="3" pageNumber="4">G. t i m - idus vs.</emphasis>
|
||
maximum
|
||
<quantity id="4CF526FB1D55FFFDB30CEC12FE7DFD99" box="[364,464,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" pageId="3" pageNumber="4" unit="mm" value="40.5">40.5 mm</quantity>
|
||
SVL in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB051EC13FD6DFD99" box="[561,704,548,572]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="humeralis">
|
||
<emphasis id="B979570C1D55FFFDB051EC13FD6DFD99" box="[561,704,548,572]" italics="true" pageId="3" pageNumber="4">G. humeralis</emphasis>
|
||
</taxonomicName>
|
||
), the presence of 3–4 lateral rows of scales at each side on distal part of fingers and toes (
|
||
<quantity id="4CF526FB1D55FFFDB3D8EC71FE4BFDFA" box="[440,486,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="4" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="4C0DF09D1D55FFFDB390EC7EFD2FFDC5" box="[496,642,584,608]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="humeralis">
|
||
<emphasis id="B979570C1D55FFFDB390EC7EFD2FFDC5" box="[496,642,584,608]" italics="true" pageId="3" pageNumber="4">G. humeralis</emphasis>
|
||
</taxonomicName>
|
||
), and the absence of a black antehumeral spot (always present in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB7E4EC7EFEAAFD21" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="humeralis">
|
||
<emphasis id="B979570C1D55FFFDB7E4EC7EFEAAFD21" italics="true" pageId="3" pageNumber="4">G. humeralis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D55FFFDB2A5ECB9FC32FC92" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="4">
|
||
Although absent from the
|
||
<collectingCountry id="F31ACB8E1D55FFFDB392ECB9FDEEFD02" box="[498,579,655,679]" name="Guyana" pageId="3" pageNumber="4">Guiana</collectingCountry>
|
||
Shield region,
|
||
<taxonomicName id="4C0DF09D1D55FFFDB093ECA6FBD3FD02" authority="Griffin, 1917" authorityName="Griffin" authorityYear="1917" box="[755,1150,655,680]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="hasemani">
|
||
<emphasis id="B979570C1D55FFFDB093ECA6FC4CFD0D" box="[755,993,656,680]" italics="true" pageId="3" pageNumber="4">Gonatodes hasemani</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D55FFFDB189ECB9FBD3FD02" author="Griffin" box="[1001,1150,655,679]" pageId="3" pageNumber="4" refString="Griffin, L. E. (1917) A list of the South American lizards of the Carnegie Museum, with descriptions of four new species. Annals of the Carnegie Museum, 11, 304 - 320." type="journal article" year="1917">Griffin, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
(a southwestern Amazonian species) was shown to be the sister species of
|
||
<taxonomicName id="4C0DF09D1D55FFFDB0BAEC83FCCBFD69" box="[730,870,692,716]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="annularis">
|
||
<emphasis id="B979570C1D55FFFDB0BAEC83FCCBFD69" box="[730,870,692,716]" italics="true" pageId="3" pageNumber="4">G. annularis</emphasis>
|
||
</taxonomicName>
|
||
and to be part of the “
|
||
<collectingCountry id="F31ACB8E1D55FFFDB603EC82FB1BFD69" box="[1123,1206,692,716]" name="Guyana" pageId="3" pageNumber="4">Guiana</collectingCountry>
|
||
Shield clade”
|
||
<emphasis id="B979570C1D55FFFDB73EEC83FA31FD69" box="[1374,1436,693,716]" italics="true" pageId="3" pageNumber="4">sensu</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D55FFFDB2F7ECE1FE3EFD4A" box="[151,403,727,752]" pageId="3" pageNumber="4" refString="Schargel, W. E., Rivas, G. A., Makowsky, R., Senaris, J. C., Natera, M. A., Barros, T. R., Molina, C. R. & Barrio-Amoros, C. L. (2010) Phylogenetic systematics of the genus Gonatodes (Squamata: Sphaerodactylidae) in the Guayana region, with description of a new species from Venezuela. Systematics and Biodiversity, 8, 321 - 339." type="journal article">
|
||
Schargel
|
||
<emphasis id="B979570C1D55FFFDB361ECEFFE96FD55" box="[257,315,728,752]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
;
|
||
<taxonomicName id="4C0DF09D1D55FFFDB3C0ECEEFD81FD55" box="[416,556,728,752]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="hasemani">
|
||
<emphasis id="B979570C1D55FFFDB3C0ECEEFD81FD55" box="[416,556,728,752]" italics="true" pageId="3" pageNumber="4">G. hasemani</emphasis>
|
||
</taxonomicName>
|
||
is immediately distinguished from
|
||
<emphasis id="B979570C1D55FFFDB1A5ECEEFB99FD55" box="[965,1076,728,752]" italics="true" pageId="3" pageNumber="4">G. t i m i d u s</emphasis>
|
||
in having a distinctly elongate supraciliary spine (absent in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB3B7ECCBFDE5FCB1" box="[471,584,764,788]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B979570C1D55FFFDB3B7ECCBFDE5FCB1" box="[471,584,764,788]" italics="true" pageId="3" pageNumber="4">G. timidus</emphasis>
|
||
</taxonomicName>
|
||
), and in having the medial subcaudal scales not differentiated from adjacent scales on non-regenerated tail (distinctly differentiated in
|
||
<taxonomicName id="4C0DF09D1D55FFFDB17EED16FC22FC9D" box="[798,911,800,824]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B979570C1D55FFFDB17EED16FC22FC9D" box="[798,911,800,824]" italics="true" pageId="3" pageNumber="4">G. timidus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317D8951D55FFF6B2A5ED75FEBFFCF4" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="8BB28B1E1D55FFFDB2A5ED75FCCBFBAA" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="4">
|
||
<emphasis id="B979570C1D55FFFDB2A5ED75FDB6FCF9" bold="true" box="[197,539,835,860]" pageId="3" pageNumber="4">
|
||
Description of the
|
||
<typeStatus id="54B635BC1D55FFFDB3C7ED72FDBBFCF9" box="[423,534,836,860]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
An adult male in very good condition (
|
||
<figureCitation id="1336979B1D55FFFDB186ED72FBFEFCF9" box="[998,1107,836,860]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1930,1953]" captionStartId-1="4.[151,250,1904,1927]" captionStartId-2="5.[151,250,1963,1986]" captionStartId-3="6.[151,250,662,685]" captionTargetBox-0="[151,1436,561,1883]" captionTargetBox-1="[239,1352,193,1863]" captionTargetBox-2="[351,1231,193,1923]" captionTargetBox-3="[151,1436,193,630]" captionTargetId-0="figure@2.[151,1436,561,1884]" captionTargetId-1="figure@4.[239,1354,193,1864]" captionTargetId-2="figure@5.[351,1232,193,1924]" captionTargetId-3="figure@6.[151,1436,193,631]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Gonatodes timidus sp. nov. in life. Top: Male holotype (IRSNB 2664, 46.9 mm SVL). Middle: Male paratype (IRSNB 2665, 46.7 mm SVL). Bottom: Female paratype (IRSNB 2667, 51.2 mm SVL). Note the conspicuous difference in the colour of head ornamentation between the two males, and the sexual dichromatism." captionText-1="FIGURE 2. Gonatodes timidus sp. nov. Sexual dichromatism in life. Top left: Ventral face of male holotype (IRSNB 2664, 46.9 mm SVL). Lower left: Detail of the gular region in the same specimen. Top right: Ventral face of female paratype (IRSNB 2670, 51.6 mm SVL). Lower right: Detail of the gular region in the same specimen. Scale bars = 10 mm (top), and 5 mm (bottom)." captionText-2="FIGURE 3. Gonatodes timidus sp. nov. Details of lateral (top), dorsal (middle), and ventral (bottom) views of the head of the male holotype in life (IRSNB 2664, 46.9 mm SVL). Scale bar = 5 mm." captionText-3="FIGURE 4. Gonatodes timidus sp. nov. Detail of pericloacal region in preserved specimens. A: Male holotype (IRSNB 2664), note distinct cluster (lower belly) and rows (thighs) of escutcheon scales. B: Female paratype (IRSNB 2670)." httpUri-0="https://zenodo.org/record/205721/files/figure.png" httpUri-1="https://zenodo.org/record/205722/files/figure.png" httpUri-2="https://zenodo.org/record/205723/files/figure.png" httpUri-3="https://zenodo.org/record/205724/files/figure.png" pageId="3" pageNumber="4">Figs. 1–4</figureCitation>
|
||
), with SVL of
|
||
<quantity id="4CF526FB1D55FFFDB764ED72FAC0FCF9" box="[1284,1389,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="4.6899999999999995" pageId="3" pageNumber="4" unit="mm" value="46.9">46.9 mm</quantity>
|
||
. TL
|
||
<quantity id="4CF526FB1D55FFFDB2F7ED51FF50FC25" box="[151,253,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" pageId="3" pageNumber="4" unit="mm" value="52.5">52.5 mm</quantity>
|
||
, tail complete, but with distal part regenerated. Head 1.2 times longer than wide (HL:
|
||
<quantity id="4CF526FB1D55FFFDB6AAED51FA9DFC25" box="[1226,1328,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="3" pageNumber="4" unit="mm" value="11.4">11.4 mm</quantity>
|
||
; HW:
|
||
<quantity id="4CF526FB1D55FFFDB719ED51FF6AFC01" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="4" unit="mm" value="9.6">9.6 mm</quantity>
|
||
). Snout short (
|
||
<quantity id="4CF526FB1D55FFFDB313EDBAFE7FFC01" box="[371,466,908,932]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="4" unit="mm" value="2.2">2.2 mm</quantity>
|
||
), 0.19 times HL, acutely rounded in dorsal view, rounded in profile, gently sloping towards top of head. Neck slightly narrower than head and body. Body nearly cylindrical, but wider than high; AXG
|
||
<quantity id="4CF526FB1D55FFFDB2BCEDE2FEE9FC49" box="[220,324,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="3" pageNumber="4" unit="mm" value="18.8">18.8 mm</quantity>
|
||
. Limbs well developed with moderately long digits, ToIV
|
||
<quantity id="4CF526FB1D55FFFDB1BEEDE2FB9AFC49" box="[990,1079,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="4" unit="mm" value="6.5">6.5 mm</quantity>
|
||
, 0.74 times ShL (
|
||
<quantity id="4CF526FB1D55FFFDB760EDE2FAF0FC49" box="[1280,1373,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="3" pageNumber="4" unit="mm" value="8.8">8.8 mm</quantity>
|
||
). Tail round in section, tapering towards tip, which is bluntly rounded.
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D55FFFDB2A5EA2BFBD8FBD9" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="4">Tongue relatively wide, narrowing anteriorly, covered by small scale-like papillae that become much less defined posteriorly; a bump bearing larger papillae is present in the middle of the tongue; posterior lateral margins indented; tip rounded with a short median cleft. Teeth small, subequal in length, conical.</paragraph>
|
||
<paragraph id="8BB28B1E1D55FFFDB2A5EABEFAEEF97A" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="4">
|
||
Rostral large, visible from above, roughly heptagonal, with a small indentation on top margin from which a short cleft extends forward about halfway through rostral. Three postrostrals, lateral ones (supranasals) distinctly larger than median, which is partially contained in the indentation of the rostral scale, and slightly larger than adjacent posterior scales on snout. Rostrals and postrostrals with many minute tubercles. Nostril bordered by rostral, lateral postrostral, three postnasals, and first supralabial. Dorsal postnasal slightly larger than lower two on the left side, noticeably larger than lower two on the right side; dorsal postnasals noticeably larger than scales in the loreal region; lower two postnasals about same size as scales in the loreal region. Scales on top of snout roughly round and granular, juxtaposed, gradually becoming more oval-shaped and conical towards loreal region. Loreal scales number about 14 (both sides) in a straight line between postnasals and anterior margin of orbit. Scales decrease slightly in size from the posterorostrals towards posterior part of the head. Scales on supraorbital region similar (in size and shape) to and continuous with those on top of head. Supraciliary flap without a distinctly elongate supraciliary spine, but three (both sides) larger conical supraciliary scales projecting posterolaterally. Pupil round. Supralabials 6 (both sides), first largest, last smallest, followed by 10/10 much smaller scales along the lip to rictus of mouth, all granular and similar in size to scales on temporal region. Fifth supralabial below centre of eye. Scales on temporal region similar (in size and shape) to those on posterior top of head, but scales contained in white stripes and blotches on head slightly larger and sometimes more spiny (especially on snout) than adjacent scales. Ear opening deep, very small (
|
||
<quantity id="4CF526FB1D55FFFDB38DE8F1FDE5F97A" box="[493,584,1735,1759]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="4" unit="mm" value="0.3">0.3 mm</quantity>
|
||
), much smaller than eye horizontal length (
|
||
<quantity id="4CF526FB1D55FFFDB64CE8F1FB25F97A" box="[1068,1160,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="4" unit="mm" value="2.5">2.5 mm</quantity>
|
||
), obliquely oval.
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D55FFFDB2A5E8DAFA31F8D4" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="4">Mental large with a round posterior margin. Postmentals three, distinctly larger than adjacent posterior scales. Scales on chin located directly behind postmentals small and polygonal, granular and tiny posteriorly, a few larger, polygonal, juxtaposed scales adjacent to infralabials. Infralabials 5 (right side)/4 (left side), decreasing in size posteriorly, first two very large and projecting onto ventral plane, suture between third and fourth below centre of eye.</paragraph>
|
||
<paragraph id="8BB28B1E1D55FFFDB2A5E94AFC75F879" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="4">Scales on nape and sides of neck continuous with those on head, granular, slightly conical anteriorly, becoming more conical posteriorly. Scales on throat smooth, imbricate, with round posterior margin, transition between juxtaposed granular, conical gular scales and scales on throat somewhat abrupt.</paragraph>
|
||
<caption id="DF72DB961D52FFFAB2F7E946FF63F847" httpUri="https://zenodo.org/record/205722/files/figure.png" pageId="4" pageNumber="5" targetBox="[239,1352,193,1863]" targetPageId="4">
|
||
<paragraph id="8BB28B1E1D52FFFAB2F7E946FF63F847" blockId="4.[151,1436,1904,2018]" pageId="4" pageNumber="5">
|
||
<emphasis id="B979570C1D52FFFAB2F7E946FEB9F820" bold="true" box="[151,276,1904,1927]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C0DF09D1D52FFFAB37CE946FE70F823" box="[284,477,1904,1926]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="timidus" status="sp. nov.">
|
||
<emphasis id="B979570C1D52FFFAB37CE946FE70F823" box="[284,477,1904,1926]" italics="true" pageId="4" pageNumber="5">Gonatodes timidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AEA771D52FFFAB385E947FD95F823" box="[485,568,1905,1926]" pageId="4" pageNumber="5" rank="species">
|
||
<emphasis id="B979570C1D52FFFAB385E947FD95F823" bold="true" box="[485,568,1905,1926]" pageId="4" pageNumber="5">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Sexual dichromatism in life. Top left: Ventral face of male holotype (IRSNB 2664, 46.9 mm SVL). Lower left: Detail of the gular region in the same specimen. Top right: Ventral face of female paratype (IRSNB 2670, 51.6 mm SVL). Lower right: Detail of the gular region in the same specimen. Scale bars = 10 mm (top), and 5 mm (bottom).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF72DB961D53FFFBB2F7E99DFCC5F845" httpUri="https://zenodo.org/record/205723/files/figure.png" pageId="5" pageNumber="6" targetBox="[351,1231,193,1923]" targetPageId="5">
|
||
<paragraph id="8BB28B1E1D53FFFBB2F7E99DFCC5F845" blockId="5.[151,1436,1963,2017]" pageId="5" pageNumber="6">
|
||
<emphasis id="B979570C1D53FFFBB2F7E99DFEBEF864" bold="true" box="[151,275,1963,1986]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C0DF09D1D53FFFBB37AE99AFE74F867" box="[282,473,1964,1986]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="timidus" status="sp. nov.">
|
||
<emphasis id="B979570C1D53FFFBB37AE99AFE74F867" box="[282,473,1964,1986]" italics="true" pageId="5" pageNumber="6">Gonatodes timidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AEA771D53FFFBB3BFE99AFD9DF864" box="[479,560,1964,1985]" pageId="5" pageNumber="6" rank="species">
|
||
<emphasis id="B979570C1D53FFFBB3BFE99AFD9DF864" bold="true" box="[479,560,1964,1985]" pageId="5" pageNumber="6">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Details of lateral (top), dorsal (middle), and ventral (bottom) views of the head of the male holotype in life (IRSNB 2664, 46.9 mm SVL). Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF72DB961D50FFF8B2F7ECA0FB45FD6E" httpUri="https://zenodo.org/record/205724/files/figure.png" pageId="6" pageNumber="7" targetBox="[151,1436,193,630]" targetPageId="6">
|
||
<paragraph id="8BB28B1E1D50FFF8B2F7ECA0FB45FD6E" blockId="6.[151,1436,662,715]" pageId="6" pageNumber="7">
|
||
<emphasis id="B979570C1D50FFF8B2F7ECA0FEBFFD0E" bold="true" box="[151,274,662,685]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C0DF09D1D50FFF8B378ECA0FE7AFD09" box="[280,471,662,684]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="timidus" status="sp. nov.">
|
||
<emphasis id="B979570C1D50FFF8B378ECA0FE7AFD09" box="[280,471,662,684]" italics="true" pageId="6" pageNumber="7">Gonatodes timidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AEA771D50FFF8B3BDECA1FD83FD09" box="[477,558,663,684]" pageId="6" pageNumber="7" rank="species">
|
||
<emphasis id="B979570C1D50FFF8B3BDECA1FD83FD09" bold="true" box="[477,558,663,684]" pageId="6" pageNumber="7">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Detail of pericloacal region in preserved specimens. A: Male holotype (IRSNB 2664), note distinct cluster (lower belly) and rows (thighs) of escutcheon scales. B: Female paratype (IRSNB 2670).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB28B1E1D50FFF8B2A5ECC1FA30FCF2" blockId="6.[151,1437,759,2008]" pageId="6" pageNumber="7">Dorsals subconical to conical, round to oval-shaped at base, slightly larger than scales on top of head, somewhat projecting posteriorly; dorsolaterally and on flanks slightly larger than dorsally, distinctly larger proximal to ventrolateral region. Transition between scales on flanks and ventrals somewhat abrupt, but not clearly demarcated.</paragraph>
|
||
<paragraph id="8BB28B1E1D50FFF8B2A7ED55FB93FB3E" blockId="6.[151,1437,759,2008]" pageId="6" pageNumber="7">
|
||
Ventral region with scales distinctly larger than dorsals, slightly smaller on chest than on belly, smooth, with round to slightly acuminate posterior margin; ventrals in oblique rows also forming rather regular longitudinal rows on belly, with 46 scales along the midventral line between an imaginary line between the anterior part of the arm insertions with the body to the vent (not including small granules lining the edge of the cloaca). Scales around midbody about 94, of which about 15 are ventrals. Escutcheon scales evident: posterior belly (between thighs) composed of a cluster of about 45 scales distinctly different from other ventrals, larger, unpigmented except slightly on their lateral margins (under high magnification all ventrals are covered by tiny flecks of dark pigments, except scales on chest, which are also unpigmented). The same unpigmented scales are present under the thighs where they are arranged in three transverse rows of about 9–11 scales each (
|
||
<figureCitation id="1336979B1D50FFF8B18BEAB5FB9DFB3E" box="[1003,1072,1155,1179]" captionStart="FIGURE 4" captionStartId="6.[151,250,662,685]" captionTargetBox="[151,1436,193,630]" captionTargetId="figure@6.[151,1436,193,631]" captionTargetPageId="6" captionText="FIGURE 4. Gonatodes timidus sp. nov. Detail of pericloacal region in preserved specimens. A: Male holotype (IRSNB 2664), note distinct cluster (lower belly) and rows (thighs) of escutcheon scales. B: Female paratype (IRSNB 2670)." httpUri="https://zenodo.org/record/205724/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D50FFF8B2A5EA9EFE55F9A6" blockId="6.[151,1437,759,2008]" pageId="6" pageNumber="7">
|
||
Scales on dorsum of tail slightly larger than on body, flat, smooth, imbricate, with posterior margin rather acuminate; transition area between the conical body scales and the caudal scales somewhat abrupt, but not clearly demarcated. Scales on underside of tail smooth, flat, imbricate, with round posterior margin, increasing in size towards midventral line; first six small subcaudals posterior to vent on midventral line increasing in size posteriorly, but not clearly differentiated from adjacent scales, followed by a midventral line of transversely enlarged scales clearly differentiated from adjacent scales, the pattern of these scales consisting in repetitive series of a single midventral scale in contact laterodistally with one scale per side followed by a single midventral in contact with two scales per side (1’1” sequence
|
||
<emphasis id="B979570C1D50FFF8B07FEB93FDF0FA19" box="[543,605,1445,1468]" italics="true" pageId="6" pageNumber="7">sensu</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D50FFF8B004EB95FC8BFA1E" author="Avila-Pires" box="[612,806,1443,1467]" pageId="6" pageNumber="7" refString="Avila-Pires, T. S. C. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">Ávila-Pires 1995</bibRefCitation>
|
||
), although the first enlarged subcaudal is divided; scales on that midventral line entirely cover the underside of the distal portion of the tail (about
|
||
<quantity id="4CF526FB1D50FFF8B61AEBFEFB73FA45" box="[1146,1246,1480,1504]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="6" pageNumber="7" unit="mm" value="18.7">18.7 mm</quantity>
|
||
long, which corresponds to a regenerated part).
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D50FFF8B2A5E826FBF9F97E" blockId="6.[151,1437,759,2008]" pageId="6" pageNumber="7">
|
||
Scales on limbs conical and juxtaposed, except ventrally and on anterior surface of forearms and thighs where they are smooth, flat, with round to acuminate posterior margin. Lamellae under first (
|
||
<emphasis id="B979570C1D50FFF8B614E802FBD2F9E9" bold="true" box="[1140,1151,1588,1612]" pageId="6" pageNumber="7">I</emphasis>
|
||
) through fifth (
|
||
<emphasis id="B979570C1D50FFF8B753E802FAEAF9E9" bold="true" box="[1331,1351,1588,1612]" pageId="6" pageNumber="7">V</emphasis>
|
||
) finger (right/left side):
|
||
<emphasis id="B979570C1D50FFF8B330E86EFEF6F9D5" bold="true" box="[336,347,1624,1648]" pageId="6" pageNumber="7">I</emphasis>
|
||
: 14/- (first finger missing on left side),
|
||
<emphasis id="B979570C1D50FFF8B17EE86EFC99F9D5" bold="true" box="[798,820,1624,1648]" pageId="6" pageNumber="7">II</emphasis>
|
||
: 15/12 (claw, and possibly distal part of second finger missing on left side),
|
||
<emphasis id="B979570C1D50FFF8B3E9E84AFE07F931" bold="true" box="[393,426,1660,1684]" pageId="6" pageNumber="7">III</emphasis>
|
||
: 18/19,
|
||
<emphasis id="B979570C1D50FFF8B066E84AFD8EF931" bold="true" box="[518,547,1660,1684]" pageId="6" pageNumber="7">IV</emphasis>
|
||
: 21/21,
|
||
<emphasis id="B979570C1D50FFF8B0E0E84AFD3FF931" bold="true" box="[640,658,1660,1684]" pageId="6" pageNumber="7">V</emphasis>
|
||
: 17/17. Lamellae under first (
|
||
<emphasis id="B979570C1D50FFF8B183E84AFC43F931" bold="true" box="[995,1006,1660,1684]" pageId="6" pageNumber="7">I</emphasis>
|
||
) through fifth (
|
||
<emphasis id="B979570C1D50FFF8B6FDE84AFB1CF931" bold="true" box="[1181,1201,1660,1684]" pageId="6" pageNumber="7">V</emphasis>
|
||
) toe (right/left side):
|
||
<emphasis id="B979570C1D50FFF8B2F7E896FF0FF91D" bold="true" box="[151,162,1696,1720]" pageId="6" pageNumber="7">I</emphasis>
|
||
: 11/12,
|
||
<emphasis id="B979570C1D50FFF8B29CE896FEBFF91D" bold="true" box="[252,274,1696,1720]" pageId="6" pageNumber="7">II</emphasis>
|
||
: 16/17,
|
||
<emphasis id="B979570C1D50FFF8B30DE896FE23F91D" bold="true" box="[365,398,1696,1720]" pageId="6" pageNumber="7">III</emphasis>
|
||
: 20/20,
|
||
<emphasis id="B979570C1D50FFF8B389E896FDABF91D" bold="true" box="[489,518,1696,1720]" pageId="6" pageNumber="7">IV</emphasis>
|
||
: 22/23,
|
||
<emphasis id="B979570C1D50FFF8B002E896FDD9F91D" bold="true" box="[610,628,1696,1720]" pageId="6" pageNumber="7">V</emphasis>
|
||
: 20/20. Fingers and toes with three to four lateral rows of scales distally. Claws exposed, non-retractile, between two basal scales (one dorsal and one ventral).
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D50FFF9B2A5E8D1FC88FDB2" blockId="6.[151,1437,759,2008]" lastBlockId="7.[151,1437,151,1140]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<emphasis id="B979570C1D50FFF8B2A5E8D1FD81F8A5" bold="true" box="[197,556,1767,1792]" pageId="6" pageNumber="7">
|
||
Colour of the
|
||
<typeStatus id="54B635BC1D50FFF8B30DE8DEFE77F8A5" box="[365,474,1768,1792]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
|
||
in life.
|
||
</emphasis>
|
||
The dorsal surface of the head is black with bluish white irregular stripes and blotches, which are distributed as follows: one broken (left side) or incomplete (right side) stripe on each
|
||
<emphasis id="B979570C1D50FFF8B75EE93AFA38F881" box="[1342,1429,1804,1828]" italics="true" pageId="6" pageNumber="7">canthus</emphasis>
|
||
, one small blotch located medially between the posterior corner of the eyes, one large oblique blotch located posterodorsally to eye on each side, and one small blotch above ear opening on each side; a very small yellowish white spot is present about
|
||
<quantity id="4CF526FB1D50FFF8B3E2E94EFE6FF835" box="[386,450,1912,1936]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="mm" value="1.0">1 mm</quantity>
|
||
posterior to rictus on each side, three similar tiny spots are found on the side of the neck on each side, and another tiny yellowish white spot is found below the last infralabials on each side. The supralabials are iridescent black. Two bluish white broken, incomplete crescent-shaped collars are present on the neck region, which is black dorsally and orange ventrolaterally. The underside of the head, including the infralabials and the chin, is orange with some scales partially or completely dark pigmented resulting in a faint reticulated pattern; the imbricate scales on the throat and the chest are orange, as well as most of the anterior surface of arms; all other ventral parts, including palms, soles, legs and tail are grey, except escutcheon scales, which are paler. A faint greyish brown, moderately broad vertebral stripe extends from posterior neck to near tail insertion. This stripe is irregularly margined by blackish blotches (five on the right side, three on the left side) and a few inconspicuous greenish brown to light brown suffusions. The flanks are chestnut brown with a few black flecks and small dark blotches. A darker patch is located on the right anterior flank, just posterior to arm insertion, and might correspond to a regenerated patch of skin. The limbs are similar in colour to the flanks, but with more dark markings. The tail is similar in colour to the dorsum, but slightly darker and without any longitudinal stripe. The iris is dark copper with a thin golden ring around the pupil. The claws are greyish white.
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D51FFF9B2A5EC15FD7DFD21" blockId="7.[151,1437,151,1140]" pageId="7" pageNumber="8">
|
||
<emphasis id="B979570C1D51FFF9B2A5EC15FD35FD99" bold="true" box="[197,664,547,572]" pageId="7" pageNumber="8">
|
||
Colour of the
|
||
<typeStatus id="54B635BC1D51FFF9B30BEC12FE75FD99" box="[363,472,548,572]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
After one month in preservative, patterns are essentially the same as in life, but the orange colour became white and the bluish tint in the white blotches and stripes disappeared. The tongue is whitish, with its first one-third dark grey.
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D51FFF9B2A5ECB8FAD7FBD6" blockId="7.[151,1437,151,1140]" pageId="7" pageNumber="8">
|
||
<emphasis id="B979570C1D51FFF9B2A5ECB8FE7AFD03" bold="true" box="[197,471,654,679]" pageId="7" pageNumber="8">
|
||
Variation in
|
||
<typeStatus id="54B635BC1D51FFF9B33AECB8FE7FFD03" box="[346,466,654,678]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="54B635BC1D51FFF9B070ECB9FDD6FD02" box="[528,635,655,679]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
consist in two adult males (SVL 46.0 and
|
||
<quantity id="4CF526FB1D51FFF9B636ECB8FB13FD03" box="[1110,1214,654,678]" metricMagnitude="-2" metricUnit="m" metricValue="4.67" pageId="7" pageNumber="8" unit="mm" value="46.7">46.7 mm</quantity>
|
||
), four adult females (SVL from
|
||
<quantity id="4CF526FB1D51FFF9B37AEC87FE7AFD6C" box="[282,471,689,713]" metricMagnitude="-2" metricUnit="m" metricValue="4.785" metricValueMax="5.16" metricValueMin="4.41" pageId="7" pageNumber="8" unit="mm" value="47.85" valueMax="51.6" valueMin="44.1">44.1 to 51.6 mm</quantity>
|
||
), and three unsexed juveniles (SVL from
|
||
<quantity id="4CF526FB1D51FFF9B1CAEC87FBCAFD6C" box="[938,1127,689,713]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.29" metricValueMin="2.91" pageId="7" pageNumber="8" unit="mm" value="31.0" valueMax="32.9" valueMin="29.1">29.1 to 32.9 mm</quantity>
|
||
).
|
||
<tableCitation id="C68FBEA51D51FFF9B61AEC87FB61FD6C" box="[1146,1228,689,713]" captionStart="TABLE 1" captionStartId="7.[151,235,1187,1208]" captionTargetBox="[151,1435,1266,1727]" captionTargetPageId="7" captionText="TABLE 1. Morphometric measurements (in mm) of the type series of Gonatodes timidus sp. nov. M = male, F = female, JUV = juvenile, other abbreviations are defined in the text." httpUri="http://table.plazi.org/id/DF72DB961D51FFF9B2F7EA95FD1AFB72" pageId="7" pageNumber="8" tableUuid="DF72DB961D51FFF9B2F7EA95FD1AFB72">Table 1</tableCitation>
|
||
provides morphometric measurements of the
|
||
<typeStatus id="54B635BC1D51FFF9B3B5ECE0FDA8FD4B" box="[469,517,726,750]" pageId="7" pageNumber="8">type</typeStatus>
|
||
series, and
|
||
<figureCitation id="1336979B1D51FFF9B0E9ECE3FD5CFD48" box="[649,753,725,749]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1930,1953]" captionStartId-1="4.[151,250,1904,1927]" captionTargetBox-0="[151,1436,561,1883]" captionTargetBox-1="[239,1352,193,1863]" captionTargetId-0="figure@2.[151,1436,561,1884]" captionTargetId-1="figure@4.[239,1354,193,1864]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. Gonatodes timidus sp. nov. in life. Top: Male holotype (IRSNB 2664, 46.9 mm SVL). Middle: Male paratype (IRSNB 2665, 46.7 mm SVL). Bottom: Female paratype (IRSNB 2667, 51.2 mm SVL). Note the conspicuous difference in the colour of head ornamentation between the two males, and the sexual dichromatism." captionText-1="FIGURE 2. Gonatodes timidus sp. nov. Sexual dichromatism in life. Top left: Ventral face of male holotype (IRSNB 2664, 46.9 mm SVL). Lower left: Detail of the gular region in the same specimen. Top right: Ventral face of female paratype (IRSNB 2670, 51.6 mm SVL). Lower right: Detail of the gular region in the same specimen. Scale bars = 10 mm (top), and 5 mm (bottom)." httpUri-0="https://zenodo.org/record/205721/files/figure.png" httpUri-1="https://zenodo.org/record/205722/files/figure.png" pageId="7" pageNumber="8">Figs. 1–2</figureCitation>
|
||
show variation and sexual dichromatism. Supralabial counts vary from 6 (n=7) to 7 (n=2, one side only), and infralabials counts vary from 4 (n=1, one side only) to 5 (n=8). Loreal scales counted in straight line between postnasals and anterior border of orbit range from 12 to 15, 14 being the most common state (n=5). Postrostrals range from 3 (n=7) to 4 (n=2). Postmentals range from 2 (n=7) to 3 (n=2). Scales around midbody range from 89 to 97, of which 16 to 19 are ventrals. There are 46 to 50 scales between an imaginary line between the anterior part of the arm insertions with the body to the vent. Clusters of escutcheon scales on lower belly range from about 45 to 50 scales; under the thighs these scales are arranged in three to four transverse rows of about 2 (most anterior row) to 13 scales each, these scales are absent in females (see
|
||
<figureCitation id="1336979B1D51FFF9B2ABEDC7FEA2FBAC" box="[203,271,1009,1033]" captionStart="FIGURE 4" captionStartId="6.[151,250,662,685]" captionTargetBox="[151,1436,193,630]" captionTargetId="figure@6.[151,1436,193,631]" captionTargetPageId="6" captionText="FIGURE 4. Gonatodes timidus sp. nov. Detail of pericloacal region in preserved specimens. A: Male holotype (IRSNB 2664), note distinct cluster (lower belly) and rows (thighs) of escutcheon scales. B: Female paratype (IRSNB 2670)." httpUri="https://zenodo.org/record/205724/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
). The pattern of subcaudals (
|
||
<emphasis id="B979570C1D51FFF9B034EDC4FD3FFBAC" box="[596,658,1010,1033]" italics="true" pageId="7" pageNumber="8">sensu</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D51FFF9B0F9EDC7FCF4FBAC" author="Avila-Pires" box="[665,857,1009,1033]" pageId="7" pageNumber="8" refString="Avila-Pires, T. S. C. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">Ávila-Pires 1995</bibRefCitation>
|
||
) is consistent, with the first distinctly enlarged subcaudal divided only in the
|
||
<typeStatus id="54B635BC1D51FFF9B3A2EA22FD89FB89" box="[450,548,1044,1068]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
and in one juvenile. Variation in the number of lamellae under first (
|
||
<emphasis id="B979570C1D51FFF9B749EA23FA99FB88" bold="true" box="[1321,1332,1045,1069]" pageId="7" pageNumber="8">I</emphasis>
|
||
) through fifth (
|
||
<emphasis id="B979570C1D51FFF9B2BAEA0EFF43FBF5" bold="true" box="[218,238,1080,1104]" pageId="7" pageNumber="8">V</emphasis>
|
||
) finger is as follows:
|
||
<emphasis id="B979570C1D51FFF9B391EA0EFE51FBF5" bold="true" box="[497,508,1080,1104]" pageId="7" pageNumber="8">I</emphasis>
|
||
: 12–16,
|
||
<emphasis id="B979570C1D51FFF9B004EA0EFDD7FBF5" bold="true" box="[612,634,1080,1104]" pageId="7" pageNumber="8">II</emphasis>
|
||
: 16–19,
|
||
<emphasis id="B979570C1D51FFF9B081EA0EFCAFFBF5" bold="true" box="[737,770,1080,1104]" pageId="7" pageNumber="8">III</emphasis>
|
||
: 17–22,
|
||
<emphasis id="B979570C1D51FFF9B10AEA0EFC2AFBF5" bold="true" box="[874,903,1080,1104]" pageId="7" pageNumber="8">IV</emphasis>
|
||
: 19–23,
|
||
<emphasis id="B979570C1D51FFF9B18FEA0EFBACFBF5" bold="true" box="[1007,1025,1080,1104]" pageId="7" pageNumber="8">V</emphasis>
|
||
: 16–20. Variation in the number of lamellae under first (
|
||
<emphasis id="B979570C1D51FFF9B3E1EA6AFE21FBD1" bold="true" box="[385,396,1116,1140]" pageId="7" pageNumber="8">I</emphasis>
|
||
) through fifth (
|
||
<emphasis id="B979570C1D51FFF9B05BEA6AFDE2FBD1" bold="true" box="[571,591,1116,1140]" pageId="7" pageNumber="8">V</emphasis>
|
||
) toe is as follows:
|
||
<emphasis id="B979570C1D51FFF9B143EA6AFC83FBD1" bold="true" box="[803,814,1116,1140]" pageId="7" pageNumber="8">I</emphasis>
|
||
: 11–14,
|
||
<emphasis id="B979570C1D51FFF9B1F0EA6AFC0BFBD1" bold="true" box="[912,934,1116,1140]" pageId="7" pageNumber="8">II</emphasis>
|
||
: 16–18,
|
||
<emphasis id="B979570C1D51FFF9B668EA6AFB84FBD1" bold="true" box="[1032,1065,1116,1140]" pageId="7" pageNumber="8">III</emphasis>
|
||
: 19–23,
|
||
<emphasis id="B979570C1D51FFF9B6ECEA6AFB04FBD1" bold="true" box="[1164,1193,1116,1140]" pageId="7" pageNumber="8">IV</emphasis>
|
||
: 22–25,
|
||
<emphasis id="B979570C1D51FFF9B76CEA6AFAB3FBD1" bold="true" box="[1292,1310,1116,1140]" pageId="7" pageNumber="8">V</emphasis>
|
||
: 19–23.
|
||
</paragraph>
|
||
<caption id="DF72DB961D51FFF9B2F7EA95FD1AFB72" ID-Table-UUID="DF72DB961D51FFF9B2F7EA95FD1AFB72" httpUri="http://table.plazi.org/id/DF72DB961D51FFF9B2F7EA95FD1AFB72" pageId="7" pageNumber="8" targetBox="[151,1435,1266,1727]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="8BB28B1E1D51FFF9B2F7EA95FD1AFB72" blockId="7.[151,1435,1186,1239]" pageId="7" pageNumber="8">
|
||
<emphasis id="B979570C1D51FFF9B2F7EA95FEA8FB1D" bold="true" box="[151,261,1187,1208]" pageId="7" pageNumber="8">TABLE 1.</emphasis>
|
||
Morphometric measurements (in mm) of the type series of
|
||
<taxonomicName id="4C0DF09D1D51FFF9B104EA95FB8FFB1C" box="[868,1058,1187,1209]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="timidus" status="sp. nov.">
|
||
<emphasis id="B979570C1D51FFF9B104EA95FB8FFB1C" box="[868,1058,1187,1209]" italics="true" pageId="7" pageNumber="8">Gonatodes timidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AEA771D51FFF9B64AEA95FBD7FB1C" box="[1066,1146,1187,1209]" pageId="7" pageNumber="8" rank="species">
|
||
<emphasis id="B979570C1D51FFF9B64AEA95FBD8FB1D" bold="true" box="[1066,1141,1187,1208]" pageId="7" pageNumber="8">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
M = male, F = female, JUV = juvenile, other abbreviations are defined in the text.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB28B1E1D51FFF9B2FFEAC4FAF3F91A" pageId="7" pageNumber="8">
|
||
<table id="F90D79BE1D510001B2F7EAC4FA36F91A" box="[151,1435,1266,1727]" gridcols="10" gridrows="9" pageId="7" pageNumber="8">
|
||
<tr id="353D895C1D510001B2F7EAC4FA36FA91" box="[151,1435,1266,1332]" gridrow="0" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7EAC4FEFBFA91" box="[151,342,1266,1332]" gridcol="0" gridrow="0" pageId="7" pageNumber="8">Specimen number IRSNB 2664</th>
|
||
<th id="76ECE0201D510001B38BEAC4FDB7FA91" box="[491,538,1266,1332]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">Sex M</th>
|
||
<th id="76ECE0201D510001B034EAC4FD29FA91" box="[596,644,1266,1332]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">SVL 46.9</th>
|
||
<th id="76ECE0201D510001B0A1EAC4FD40FA91" box="[705,749,1266,1332]" gridcol="3" gridrow="0" pageId="7" pageNumber="8">TL 52.5</th>
|
||
<th id="76ECE0201D510001B147EAC4FCFEFA91" box="[807,851,1266,1332]" gridcol="4" gridrow="0" pageId="7" pageNumber="8">HL 11.4</th>
|
||
<th id="76ECE0201D510001B1ECEAC4FC15FA91" box="[908,952,1266,1332]" gridcol="5" gridrow="0" pageId="7" pageNumber="8">HW 9.6</th>
|
||
<th id="76ECE0201D510001B193EAC4FBB8FA91" box="[1011,1045,1266,1332]" gridcol="6" gridrow="0" pageId="7" pageNumber="8">EN 3.6</th>
|
||
<th id="76ECE0201D510001B62BEAC4FB2FFA91" box="[1099,1154,1266,1332]" gridcol="7" gridrow="0" pageId="7" pageNumber="8">AXG 18.8</th>
|
||
<th id="76ECE0201D510001B6A3EAC4FB55FA91" box="[1219,1272,1266,1332]" gridcol="8" gridrow="0" pageId="7" pageNumber="8">ToIV 6.5</th>
|
||
<th id="76ECE0201D510001B75DEAC4FA36FA91" box="[1341,1435,1266,1332]" gridcol="9" gridrow="0" pageId="7" pageNumber="8">ShL 8.8</th>
|
||
</tr>
|
||
<tr id="353D895C1D510001B2F7EB7CFA36FAC5" box="[151,1435,1354,1376]" gridrow="1" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7EB7CFEFBFAC5" box="[151,342,1354,1376]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">IRSNB 2665</th>
|
||
<td id="76ECE0201D510001B38BEB7CFDB7FAC5" box="[491,538,1354,1376]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">M</td>
|
||
<td id="76ECE0201D510001B034EB7CFD29FAC5" box="[596,644,1354,1376]" gridcol="2" gridrow="1" pageId="7" pageNumber="8">46.7</td>
|
||
<td id="76ECE0201D510001B0A1EB7CFD40FAC5" box="[705,749,1354,1376]" gridcol="3" gridrow="1" pageId="7" pageNumber="8">-</td>
|
||
<td id="76ECE0201D510001B147EB7CFCFEFAC5" box="[807,851,1354,1376]" gridcol="4" gridrow="1" pageId="7" pageNumber="8">11.3</td>
|
||
<td id="76ECE0201D510001B1ECEB7CFC15FAC5" box="[908,952,1354,1376]" gridcol="5" gridrow="1" pageId="7" pageNumber="8">8.9</td>
|
||
<td id="76ECE0201D510001B193EB7CFBB8FAC5" box="[1011,1045,1354,1376]" gridcol="6" gridrow="1" pageId="7" pageNumber="8">3.8</td>
|
||
<td id="76ECE0201D510001B62BEB7CFB2FFAC5" box="[1099,1154,1354,1376]" gridcol="7" gridrow="1" pageId="7" pageNumber="8">17.8</td>
|
||
<td id="76ECE0201D510001B6A3EB7CFB55FAC5" box="[1219,1272,1354,1376]" gridcol="8" gridrow="1" pageId="7" pageNumber="8">6.6</td>
|
||
<td id="76ECE0201D510001B75DEB7CFA36FAC5" box="[1341,1435,1354,1376]" gridcol="9" gridrow="1" pageId="7" pageNumber="8">9.3</td>
|
||
</tr>
|
||
<tr id="353D895C1D510001B2F7EB40FA36FA29" box="[151,1435,1398,1420]" gridrow="2" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7EB40FEFBFA29" box="[151,342,1398,1420]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">IRSNB 2666</th>
|
||
<td id="76ECE0201D510001B38BEB40FDB7FA29" box="[491,538,1398,1420]" gridcol="1" gridrow="2" pageId="7" pageNumber="8">M</td>
|
||
<td id="76ECE0201D510001B034EB40FD29FA29" box="[596,644,1398,1420]" gridcol="2" gridrow="2" pageId="7" pageNumber="8">46.0</td>
|
||
<td id="76ECE0201D510001B0A1EB40FD40FA29" box="[705,749,1398,1420]" gridcol="3" gridrow="2" pageId="7" pageNumber="8">-</td>
|
||
<td id="76ECE0201D510001B147EB40FCFEFA29" box="[807,851,1398,1420]" gridcol="4" gridrow="2" pageId="7" pageNumber="8">11.1</td>
|
||
<td id="76ECE0201D510001B1ECEB40FC15FA29" box="[908,952,1398,1420]" gridcol="5" gridrow="2" pageId="7" pageNumber="8">8.7</td>
|
||
<td id="76ECE0201D510001B193EB40FBB8FA29" box="[1011,1045,1398,1420]" gridcol="6" gridrow="2" pageId="7" pageNumber="8">3.5</td>
|
||
<td id="76ECE0201D510001B62BEB40FB2FFA29" box="[1099,1154,1398,1420]" gridcol="7" gridrow="2" pageId="7" pageNumber="8">18.9</td>
|
||
<td id="76ECE0201D510001B6A3EB40FB55FA29" box="[1219,1272,1398,1420]" gridcol="8" gridrow="2" pageId="7" pageNumber="8">5.9</td>
|
||
<td id="76ECE0201D510001B75DEB40FA36FA29" box="[1341,1435,1398,1420]" gridcol="9" gridrow="2" pageId="7" pageNumber="8">9.2</td>
|
||
</tr>
|
||
<tr id="353D895C1D510001B2F7EB94FA36FA1D" box="[151,1435,1442,1464]" gridrow="3" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7EB94FEFBFA1D" box="[151,342,1442,1464]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">IRSNB 2667</th>
|
||
<td id="76ECE0201D510001B38BEB94FDB7FA1D" box="[491,538,1442,1464]" gridcol="1" gridrow="3" pageId="7" pageNumber="8">F</td>
|
||
<td id="76ECE0201D510001B034EB94FD29FA1D" box="[596,644,1442,1464]" gridcol="2" gridrow="3" pageId="7" pageNumber="8">51.2</td>
|
||
<td id="76ECE0201D510001B0A1EB94FD40FA1D" box="[705,749,1442,1464]" gridcol="3" gridrow="3" pageId="7" pageNumber="8">54.6</td>
|
||
<td id="76ECE0201D510001B147EB94FCFEFA1D" box="[807,851,1442,1464]" gridcol="4" gridrow="3" pageId="7" pageNumber="8">11.9</td>
|
||
<td id="76ECE0201D510001B1ECEB94FC15FA1D" box="[908,952,1442,1464]" gridcol="5" gridrow="3" pageId="7" pageNumber="8">9.7</td>
|
||
<td id="76ECE0201D510001B193EB94FBB8FA1D" box="[1011,1045,1442,1464]" gridcol="6" gridrow="3" pageId="7" pageNumber="8">4.0</td>
|
||
<td id="76ECE0201D510001B62BEB94FB2FFA1D" box="[1099,1154,1442,1464]" gridcol="7" gridrow="3" pageId="7" pageNumber="8">21.2</td>
|
||
<td id="76ECE0201D510001B6A3EB94FB55FA1D" box="[1219,1272,1442,1464]" gridcol="8" gridrow="3" pageId="7" pageNumber="8">6.6</td>
|
||
<td id="76ECE0201D510001B75DEB94FA36FA1D" box="[1341,1435,1442,1464]" gridcol="9" gridrow="3" pageId="7" pageNumber="8">9.4</td>
|
||
</tr>
|
||
<tr id="353D895C1D510001B2F7EBF8FA36FA41" box="[151,1435,1486,1508]" gridrow="4" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7EBF8FEFBFA41" box="[151,342,1486,1508]" gridcol="0" gridrow="4" pageId="7" pageNumber="8">IRSNB 2668</th>
|
||
<td id="76ECE0201D510001B38BEBF8FDB7FA41" box="[491,538,1486,1508]" gridcol="1" gridrow="4" pageId="7" pageNumber="8">F</td>
|
||
<td id="76ECE0201D510001B034EBF8FD29FA41" box="[596,644,1486,1508]" gridcol="2" gridrow="4" pageId="7" pageNumber="8">44.1</td>
|
||
<td id="76ECE0201D510001B0A1EBF8FD40FA41" box="[705,749,1486,1508]" gridcol="3" gridrow="4" pageId="7" pageNumber="8">-</td>
|
||
<td id="76ECE0201D510001B147EBF8FCFEFA41" box="[807,851,1486,1508]" gridcol="4" gridrow="4" pageId="7" pageNumber="8">10.6</td>
|
||
<td id="76ECE0201D510001B1ECEBF8FC15FA41" box="[908,952,1486,1508]" gridcol="5" gridrow="4" pageId="7" pageNumber="8">8.2</td>
|
||
<td id="76ECE0201D510001B193EBF8FBB8FA41" box="[1011,1045,1486,1508]" gridcol="6" gridrow="4" pageId="7" pageNumber="8">3.1</td>
|
||
<td id="76ECE0201D510001B62BEBF8FB2FFA41" box="[1099,1154,1486,1508]" gridcol="7" gridrow="4" pageId="7" pageNumber="8">19.0</td>
|
||
<td id="76ECE0201D510001B6A3EBF8FB55FA41" box="[1219,1272,1486,1508]" gridcol="8" gridrow="4" pageId="7" pageNumber="8">5.8</td>
|
||
<td id="76ECE0201D510001B75DEBF8FA36FA41" box="[1341,1435,1486,1508]" gridcol="9" gridrow="4" pageId="7" pageNumber="8">8.6</td>
|
||
</tr>
|
||
<tr id="353D895C1D510001B2F7EBCCFA36F9B5" box="[151,1435,1530,1552]" gridrow="5" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7EBCCFEFBF9B5" box="[151,342,1530,1552]" gridcol="0" gridrow="5" pageId="7" pageNumber="8">IRSNB 2669</th>
|
||
<td id="76ECE0201D510001B38BEBCCFDB7F9B5" box="[491,538,1530,1552]" gridcol="1" gridrow="5" pageId="7" pageNumber="8">F</td>
|
||
<td id="76ECE0201D510001B034EBCCFD29F9B5" box="[596,644,1530,1552]" gridcol="2" gridrow="5" pageId="7" pageNumber="8">46.0</td>
|
||
<td id="76ECE0201D510001B0A1EBCCFD40F9B5" box="[705,749,1530,1552]" gridcol="3" gridrow="5" pageId="7" pageNumber="8">-</td>
|
||
<td id="76ECE0201D510001B147EBCCFCFEF9B5" box="[807,851,1530,1552]" gridcol="4" gridrow="5" pageId="7" pageNumber="8">11.3</td>
|
||
<td id="76ECE0201D510001B1ECEBCCFC15F9B5" box="[908,952,1530,1552]" gridcol="5" gridrow="5" pageId="7" pageNumber="8">8.0</td>
|
||
<td id="76ECE0201D510001B193EBCCFBB8F9B5" box="[1011,1045,1530,1552]" gridcol="6" gridrow="5" pageId="7" pageNumber="8">3.5</td>
|
||
<td id="76ECE0201D510001B62BEBCCFB2FF9B5" box="[1099,1154,1530,1552]" gridcol="7" gridrow="5" pageId="7" pageNumber="8">20.6</td>
|
||
<td id="76ECE0201D510001B6A3EBCCFB55F9B5" box="[1219,1272,1530,1552]" gridcol="8" gridrow="5" pageId="7" pageNumber="8">5.4</td>
|
||
<td id="76ECE0201D510001B75DEBCCFA36F9B5" box="[1341,1435,1530,1552]" gridcol="9" gridrow="5" pageId="7" pageNumber="8">8.8</td>
|
||
</tr>
|
||
<tr id="353D895C1D510001B2F7E810FA36F999" box="[151,1435,1574,1596]" gridrow="6" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7E810FEFBF999" box="[151,342,1574,1596]" gridcol="0" gridrow="6" pageId="7" pageNumber="8">IRSNB 2670</th>
|
||
<td id="76ECE0201D510001B38BE810FDB7F999" box="[491,538,1574,1596]" gridcol="1" gridrow="6" pageId="7" pageNumber="8">F</td>
|
||
<td id="76ECE0201D510001B034E810FD29F999" box="[596,644,1574,1596]" gridcol="2" gridrow="6" pageId="7" pageNumber="8">51.6</td>
|
||
<td id="76ECE0201D510001B0A1E810FD40F999" box="[705,749,1574,1596]" gridcol="3" gridrow="6" pageId="7" pageNumber="8">46.6</td>
|
||
<td id="76ECE0201D510001B147E810FCFEF999" box="[807,851,1574,1596]" gridcol="4" gridrow="6" pageId="7" pageNumber="8">12.0</td>
|
||
<td id="76ECE0201D510001B1ECE810FC15F999" box="[908,952,1574,1596]" gridcol="5" gridrow="6" pageId="7" pageNumber="8">10.0</td>
|
||
<td id="76ECE0201D510001B193E810FBB8F999" box="[1011,1045,1574,1596]" gridcol="6" gridrow="6" pageId="7" pageNumber="8">4.2</td>
|
||
<td id="76ECE0201D510001B62BE810FB2FF999" box="[1099,1154,1574,1596]" gridcol="7" gridrow="6" pageId="7" pageNumber="8">23.1</td>
|
||
<td id="76ECE0201D510001B6A3E810FB55F999" box="[1219,1272,1574,1596]" gridcol="8" gridrow="6" pageId="7" pageNumber="8">6.8</td>
|
||
<td id="76ECE0201D510001B75DE810FA36F999" box="[1341,1435,1574,1596]" gridcol="9" gridrow="6" pageId="7" pageNumber="8">9.7</td>
|
||
</tr>
|
||
<tr id="353D895C1D510001B2F7E864FA36F936" box="[151,1435,1618,1683]" gridrow="7" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7E864FEFBF936" box="[151,342,1618,1683]" gridcol="0" gridrow="7" pageId="7" pageNumber="8">IRSNB 2671 IRSNB 2672</th>
|
||
<td id="76ECE0201D510001B38BE864FDB7F936" box="[491,538,1618,1683]" gridcol="1" gridrow="7" pageId="7" pageNumber="8">JUV JUV</td>
|
||
<td id="76ECE0201D510001B034E864FD29F936" box="[596,644,1618,1683]" gridcol="2" gridrow="7" pageId="7" pageNumber="8">29.1 32.9</td>
|
||
<td id="76ECE0201D510001B0A1E864FD40F936" box="[705,749,1618,1683]" gridcol="3" gridrow="7" pageId="7" pageNumber="8">- -</td>
|
||
<td id="76ECE0201D510001B147E864FCFEF936" box="[807,851,1618,1683]" gridcol="4" gridrow="7" pageId="7" pageNumber="8">7.1 8.6</td>
|
||
<td id="76ECE0201D510001B1ECE864FC15F936" box="[908,952,1618,1683]" gridcol="5" gridrow="7" pageId="7" pageNumber="8">6.0 6.4</td>
|
||
<td id="76ECE0201D510001B193E864FBB8F936" box="[1011,1045,1618,1683]" gridcol="6" gridrow="7" pageId="7" pageNumber="8">2.3 2.5</td>
|
||
<td id="76ECE0201D510001B62BE864FB2FF936" box="[1099,1154,1618,1683]" gridcol="7" gridrow="7" pageId="7" pageNumber="8">12.7 13.7</td>
|
||
<td id="76ECE0201D510001B6A3E864FB55F936" box="[1219,1272,1618,1683]" gridcol="8" gridrow="7" pageId="7" pageNumber="8">3.8 4.7</td>
|
||
<td id="76ECE0201D510001B75DE864FA36F936" box="[1341,1435,1618,1683]" gridcol="9" gridrow="7" pageId="7" pageNumber="8">5.1 6.5</td>
|
||
</tr>
|
||
<tr id="353D895C1D510001B2F7E89FFA36F91A" box="[151,1435,1705,1727]" gridrow="8" pageId="7" pageNumber="8">
|
||
<th id="76ECE0201D510001B2F7E89FFEFBF91A" box="[151,342,1705,1727]" gridcol="0" gridrow="8" pageId="7" pageNumber="8">IRSNB 2673</th>
|
||
<td id="76ECE0201D510001B38BE89FFDB7F91A" box="[491,538,1705,1727]" gridcol="1" gridrow="8" pageId="7" pageNumber="8">JUV</td>
|
||
<td id="76ECE0201D510001B034E89FFD29F91A" box="[596,644,1705,1727]" gridcol="2" gridrow="8" pageId="7" pageNumber="8">30.2</td>
|
||
<td id="76ECE0201D510001B0A1E89FFD40F91A" box="[705,749,1705,1727]" gridcol="3" gridrow="8" pageId="7" pageNumber="8">-</td>
|
||
<td id="76ECE0201D510001B147E89FFCFEF91A" box="[807,851,1705,1727]" gridcol="4" gridrow="8" pageId="7" pageNumber="8">8.0</td>
|
||
<td id="76ECE0201D510001B1ECE89FFC15F91A" box="[908,952,1705,1727]" gridcol="5" gridrow="8" pageId="7" pageNumber="8">6.1</td>
|
||
<td id="76ECE0201D510001B193E89FFBB8F91A" box="[1011,1045,1705,1727]" gridcol="6" gridrow="8" pageId="7" pageNumber="8">2.4</td>
|
||
<td id="76ECE0201D510001B62BE89FFB2FF91A" box="[1099,1154,1705,1727]" gridcol="7" gridrow="8" pageId="7" pageNumber="8">12.3</td>
|
||
<td id="76ECE0201D510001B6A3E89FFB55F91A" box="[1219,1272,1705,1727]" gridcol="8" gridrow="8" pageId="7" pageNumber="8">3.9</td>
|
||
<td id="76ECE0201D510001B75DE89FFA36F91A" box="[1341,1435,1705,1727]" gridcol="9" gridrow="8" pageId="7" pageNumber="8">6.3</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D51FFF9B2A5E931FF4BF8C0" blockId="7.[151,1436,1799,2036]" pageId="7" pageNumber="8">
|
||
All
|
||
<typeStatus id="54B635BC1D51FFF9B292E93EFEF0F885" box="[242,349,1800,1824]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
have a few larger conical supraciliary scales projecting posterolaterally, but a distinct elongate supraciliary spine as in
|
||
<taxonomicName id="4C0DF09D1D51FFF9B3D4E91CFCA8F8E7" box="[436,773,1834,1858]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="alexandermendesi">
|
||
<emphasis id="B979570C1D51FFF9B3D4E91CFCA8F8E7" box="[436,773,1834,1858]" italics="true" pageId="7" pageNumber="8">Gonatodes alexandermendesi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0DF09D1D51FFF9B178E91DFC3DF8E7" box="[792,912,1834,1858]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="astralis">
|
||
<emphasis id="B979570C1D51FFF9B178E91DFC3DF8E7" box="[792,912,1834,1858]" italics="true" pageId="7" pageNumber="8">G. astralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0DF09D1D51FFF9B1C3E91DFB9EF8E7" box="[931,1075,1834,1858]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="hasemani">
|
||
<emphasis id="B979570C1D51FFF9B1C3E91DFB9EF8E7" box="[931,1075,1834,1858]" italics="true" pageId="7" pageNumber="8">G. hasemani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DF09D1D51FFF9B613E91DFA8EF8E7" box="[1139,1315,1834,1858]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="superciliaris">
|
||
<emphasis id="B979570C1D51FFF9B613E91DFA8EF8E7" box="[1139,1315,1834,1858]" italics="true" pageId="7" pageNumber="8">G. superciliaris</emphasis>
|
||
</taxonomicName>
|
||
is always absent.
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D51FFF6B2A5E947FEBFFCF4" blockId="7.[151,1436,1799,2036]" lastBlockId="8.[151,1437,151,1389]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
There is an evident sexual dichromatism (see
|
||
<figureCitation id="1336979B1D51FFF9B0AAE947FC83F82C" box="[714,814,1905,1929]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1930,1953]" captionStartId-1="4.[151,250,1904,1927]" captionTargetBox-0="[151,1436,561,1883]" captionTargetBox-1="[239,1352,193,1863]" captionTargetId-0="figure@2.[151,1436,561,1884]" captionTargetId-1="figure@4.[239,1354,193,1864]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. Gonatodes timidus sp. nov. in life. Top: Male holotype (IRSNB 2664, 46.9 mm SVL). Middle: Male paratype (IRSNB 2665, 46.7 mm SVL). Bottom: Female paratype (IRSNB 2667, 51.2 mm SVL). Note the conspicuous difference in the colour of head ornamentation between the two males, and the sexual dichromatism." captionText-1="FIGURE 2. Gonatodes timidus sp. nov. Sexual dichromatism in life. Top left: Ventral face of male holotype (IRSNB 2664, 46.9 mm SVL). Lower left: Detail of the gular region in the same specimen. Top right: Ventral face of female paratype (IRSNB 2670, 51.6 mm SVL). Lower right: Detail of the gular region in the same specimen. Scale bars = 10 mm (top), and 5 mm (bottom)." httpUri-0="https://zenodo.org/record/205721/files/figure.png" httpUri-1="https://zenodo.org/record/205722/files/figure.png" pageId="7" pageNumber="8">Figs 1–2</figureCitation>
|
||
). In life, the two male
|
||
<typeStatus id="54B635BC1D51FFF9B650E944FB36F82F" box="[1072,1179,1906,1930]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
are very similar to the
|
||
<typeStatus id="54B635BC1D51FFF9B2F7E9A2FF54F809" box="[151,249,1940,1964]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
in colour and pattern, except that IRSNB 2665 has vivid yellow stripes and blotches on head (instead of bluish white), a less contrasted pattern on body, and a paler body colour. Stripes on
|
||
<emphasis id="B979570C1D51FFF9B65CE98EFB3EF875" box="[1084,1171,1976,2000]" italics="true" pageId="7" pageNumber="8">canthus</emphasis>
|
||
extend from postnasals to near the eye in both male
|
||
<typeStatus id="54B635BC1D51FFF9B3B1E9EAFD91F851" box="[465,572,2012,2036]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
(broken and incomplete in the
|
||
<typeStatus id="54B635BC1D51FFF9B1F7E9EDFC53F856" box="[919,1022,2011,2035]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
), and the two crescent-shaped collars are more complete. Sides of neck are less orangish in the male
|
||
<typeStatus id="54B635BC1D5EFFF6B101EEAEFC61FF15" box="[865,972,152,176]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
|
||
than in the
|
||
<typeStatus id="54B635BC1D5EFFF6B633EEA1FB15FF0A" box="[1107,1208,151,175]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
. In preservative, the male
|
||
<typeStatus id="54B635BC1D5EFFF6B2B4EE8AFE92FF71" box="[212,319,188,212]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
|
||
are almost identical to the
|
||
<typeStatus id="54B635BC1D5EFFF6B00EEE8DFD7EFF76" box="[622,723,187,211]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
, except in the shape of blotches and stripes on the head. IRSNB 2666 has a slightly pinkish throat (instead of white in the other
|
||
<typeStatus id="54B635BC1D5EFFF6B111EED6FC7CFF5D" box="[881,977,224,248]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
and in the
|
||
<typeStatus id="54B635BC1D5EFFF6B635EEE9FB11FF52" box="[1109,1212,223,247]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
). In life, the female
|
||
<typeStatus id="54B635BC1D5EFFF6B2F7EF35FEAFFEBE" box="[151,258,259,283]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
|
||
are brownish dorsally and laterally, with some whitish markings on tail, flanks, and limbs. Like in males a faint greyish brown, moderately broad vertebral stripe extends from posterior neck to near tail insertion and is irregularly margined by inconspicuous blackish blotches (more conspicuous in IRSNB 2668 and IRSNB 2669, which are the two smallest adult females). That vertebral stripe is also much more conspicuous in IRSNB 2668 and IRSNB 2669, and probably faints with age. The four female
|
||
<typeStatus id="54B635BC1D5EFFF6B124EFA7FC02FE0C" box="[836,943,401,425]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
|
||
lack any colourful head ornamentation, but have two (only one in IRSNB 2670) thin, black-edged white crescent-shaped collars on the neck region, and usually a faint oblique stripe/blotch located posterodorsally to eye on each side (inconspicuous in IRSNB 2667). The dorsal surface of the head is slightly lighter than the body (distinctly lighter in IRSNB 2670). The supralabials are iridescent light grey, sometimes whitish posteriorly. The underside of the head, including the throat and the upper chest is black with bluish white reticulation (see
|
||
<figureCitation id="1336979B1D5EFFF6B0D4EC77FD55FDFC" box="[692,760,577,601]" captionStart="FIGURE 2" captionStartId="4.[151,250,1904,1927]" captionTargetBox="[239,1352,193,1863]" captionTargetId="figure@4.[239,1354,193,1864]" captionTargetPageId="4" captionText="FIGURE 2. Gonatodes timidus sp. nov. Sexual dichromatism in life. Top left: Ventral face of male holotype (IRSNB 2664, 46.9 mm SVL). Lower left: Detail of the gular region in the same specimen. Top right: Ventral face of female paratype (IRSNB 2670, 51.6 mm SVL). Lower right: Detail of the gular region in the same specimen. Scale bars = 10 mm (top), and 5 mm (bottom)." httpUri="https://zenodo.org/record/205722/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
|
||
). The infralabials are dark grey, usually with a variably large white area posteriorly. The ventral face is light grey with a suffusion of light orangish yellow (see
|
||
<figureCitation id="1336979B1D5EFFF6B687EC53FA81FDD8" box="[1255,1324,613,637]" captionStart="FIGURE 2" captionStartId="4.[151,250,1904,1927]" captionTargetBox="[239,1352,193,1863]" captionTargetId="figure@4.[239,1354,193,1864]" captionTargetPageId="4" captionText="FIGURE 2. Gonatodes timidus sp. nov. Sexual dichromatism in life. Top left: Ventral face of male holotype (IRSNB 2664, 46.9 mm SVL). Lower left: Detail of the gular region in the same specimen. Top right: Ventral face of female paratype (IRSNB 2670, 51.6 mm SVL). Lower right: Detail of the gular region in the same specimen. Scale bars = 10 mm (top), and 5 mm (bottom)." httpUri="https://zenodo.org/record/205722/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
|
||
); the ventral surface of the tail is dark grey. In preservative the female
|
||
<typeStatus id="54B635BC1D5EFFF6B127ECBFFC1FFD04" box="[839,946,649,673]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
|
||
are dorsally darker than in life, but patterns remain essentially the same as in life. The bluish white colour became white, and the venter became very light grey. The three unsexed juveniles have essentially the same pattern and colouration as the adult females, including on throat, but patterns are more conspicuous and usually more contrasted. Two juveniles, IRSNB 2671 and IRSNB 2673, have a less contrasted throat pattern compared to the other juvenile. Females and juveniles usually have lighter iris.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317D8951D5EFFF6B2A5ED6BFCEAFAC8" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="8BB28B1E1D5EFFF6B2A5ED6BFE0FFC18" blockId="8.[151,1437,151,1389]" pageId="8" pageNumber="9">
|
||
<emphasis id="B979570C1D5EFFF6B2A5ED6BFE5FFCD0" bold="true" box="[197,498,861,886]" pageId="8" pageNumber="9">Distribution and ecology.</emphasis>
|
||
The new species is currently known only from the
|
||
<typeStatus id="54B635BC1D5EFFF6B65CED68FBC1FCD3" box="[1084,1132,862,886]" pageId="8" pageNumber="9">type</typeStatus>
|
||
locality, between
|
||
<quantity id="4CF526FB1D5EFFF6B75AED6BFF00FC3C" metricMagnitude="2" metricUnit="m" metricValue="2.17" metricValueMax="2.25" metricValueMin="2.09" pageId="8" pageNumber="9" unit="m" value="217.0" valueMax="225.0" valueMin="209.0">209–225 m</quantity>
|
||
elevation, in lowland primary rainforest at the base of the Iwokrama Mountains in the Iwokrama Forest Reserve, central
|
||
<collectingCountry id="F31ACB8E1D5EFFF6B28AED93FEECFC18" box="[234,321,933,957]" name="Guyana" pageId="8" pageNumber="9">Guyana</collectingCountry>
|
||
(
|
||
<figureCitation id="1336979B1D5EFFF6B330ED93FE38FC18" box="[336,405,933,957]" captionStart="FIGURE 5" captionStartId="9.[151,250,1936,1959]" captionTargetBox="[151,1436,195,1907]" captionTargetId="figure@9.[151,1436,195,1907]" captionTargetPageId="9" captionText="FIGURE 5. Map of the Iwokrama Forest Reserve. The black dot indicates the type locality of Gonatodes timidus sp. nov., at the foot of the Iwokrama Mountains. The red line crossing the Reserve is the road between Linden and Lethem. Map courtesy Iwokrama International Centre for Rainforest Conservation and Development." httpUri="https://zenodo.org/record/205725/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D5EFFF6B2A5EDFFFCCCFB30" blockId="8.[151,1437,151,1389]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C0DF09D1D5EFFF6B2A5EDFFFE3AFC44" box="[197,407,969,993]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B979570C1D5EFFF6B2A5EDFFFE3AFC44" box="[197,407,969,993]" italics="true" pageId="8" pageNumber="9">Gonatodes timidus</emphasis>
|
||
</taxonomicName>
|
||
is diurnal, locally abundant, but difficult to collect because specimens are very shy and tend to quickly escape in cracks and holes between boulders when disturbed. Most adult specimens have been collected on, or between relatively large boulders located about
|
||
<quantity id="4CF526FB1D5EFFF6B09FEA27FCCEFB8C" box="[767,867,1041,1065]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="9" unit="m" value="20.0" valueMax="30.0" valueMin="10.0">10–30 m</quantity>
|
||
from a stream (
|
||
<figureCitation id="1336979B1D5EFFF6B679EA27FBF3FB8C" box="[1049,1118,1041,1065]" captionStart="FIGURE 6" captionStartId="10.[151,250,1509,1532]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@10.[151,1436,193,1488]" captionTargetPageId="10" captionText="FIGURE 6. Habitat of Gonatodes timidus sp. nov. at the type locality. A: Macrohabitat, the white arrow indicates a cluster of large boulders on the primary forest floor. B: Microhabitat, the white arrow indicates the exact place where the holotype of G. timidus sp. nov. was first spotted. It quickly escaped in the hole between the boulders indicated by a black arrow. C: The stream running at the type locality (22 March 2011)." httpUri="https://zenodo.org/record/205726/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
|
||
). One adult female (IRSNB 2669) was collected between cavities on a tree trunk, close to large boulders. Two juveniles (IRSNB
|
||
<date id="FFB3ADDE1D5EFFF6B696EA03FAF4FBE8" box="[1270,1369,1077,1101]" pageId="8" pageNumber="9" value="2671" valueMax="2672">2671–72</date>
|
||
) have been collected among the leaf litter close to boulders, an additional juvenile (IRSNB 2673) was collected in a rotten fallen tree trunk. No juveniles have been observed on rocks.
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D5EFFF6B2A5EA94FF4CFA80" blockId="8.[151,1437,151,1389]" pageId="8" pageNumber="9">
|
||
<collectingCountry id="F31ACB8E1D5EFFF6B2A5EA94FF49FB1F" box="[197,228,1186,1210]" name="American Samoa" pageId="8" pageNumber="9">As</collectingCountry>
|
||
reported in other
|
||
<taxonomicName id="4C0DF09D1D5EFFF6B3D3EA97FD81FB1C" box="[435,556,1185,1209]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
|
||
<emphasis id="B979570C1D5EFFF6B3D3EA97FD81FB1C" box="[435,556,1185,1209]" italics="true" pageId="8" pageNumber="9">Gonatodes</emphasis>
|
||
</taxonomicName>
|
||
species (see
|
||
<bibRefCitation id="EF9CF6EF1D5EFFF6B0A4EA97FC3EFB1C" author="Cole" box="[708,915,1185,1209]" pageId="8" pageNumber="9" refString="Cole, C. J. & Kok, P. J. R. (2006) A new species of gekkonid lizard (Sphaerodactylinae: Gonatodes) from Guyana, South America. American Museum Novitates, 3524, 1 - 13." type="journal article" year="2006">Cole & Kok 2006</bibRefCitation>
|
||
) individuals of
|
||
<taxonomicName id="4C0DF09D1D5EFFF6B628EA97FAB0FB1C" box="[1096,1309,1185,1209]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="timidus">
|
||
<emphasis id="B979570C1D5EFFF6B628EA97FAB0FB1C" box="[1096,1309,1185,1209]" italics="true" pageId="8" pageNumber="9">Gonatodes timidus</emphasis>
|
||
</taxonomicName>
|
||
easily lose part of their skin as an antipredator strategy (regional intergumentary loss
|
||
<emphasis id="B979570C1D5EFFF6B667EAF0FBE8FB78" box="[1031,1093,1222,1245]" italics="true" pageId="8" pageNumber="9">sensu</emphasis>
|
||
<bibRefCitation id="EF9CF6EF1D5EFFF6B631EAF0FA8EFB78" author="Bauer" box="[1105,1315,1221,1245]" pageId="8" pageNumber="9" refString="Bauer, A. M., Russell, A. P. & Shadwick, R. E. (1989) Mechanical properties and morphological correlates of fragile skin in gekkonid lizards. The Journal of Experimental Biology, 145, 79 - 102." type="journal article" year="1989">
|
||
Bauer
|
||
<emphasis id="B979570C1D5EFFF6B6FFEAF0FB71FB78" box="[1183,1244,1221,1245]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
1989
|
||
</bibRefCitation>
|
||
), and also promptly self-amputate their tail (caudal autotomy), which makes exceedingly difficult to secure undamaged specimens.
|
||
</paragraph>
|
||
<paragraph id="8BB28B1E1D5EFFF6B2A5EB07FCEAFAC8" blockId="8.[151,1437,151,1389]" pageId="8" pageNumber="9">
|
||
The new species was found sympatric, but never syntopic, with
|
||
<taxonomicName id="4C0DF09D1D5EFFF6B1C5EB07FB5EFAEC" box="[933,1267,1329,1353]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="alexandermendesi">
|
||
<emphasis id="B979570C1D5EFFF6B1C5EB07FB5EFAEC" box="[933,1267,1329,1353]" italics="true" pageId="8" pageNumber="9">Gonatodes alexandermendesi</emphasis>
|
||
</taxonomicName>
|
||
, which is here reported for the first time from the Iwokrama Forest Reserve.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |