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<document id="2E6D6A074C5A4A5DEE0F8404F17AA049" ID-CLB-Dataset="45364" ID-DOI="10.11646/zootaxa.3826.1.3" ID-GBIF-Dataset="6f4b1535-6c6b-4446-aa26-b1ba83ad9126" ID-ISSN="1175-5326" ID-Zenodo-Dep="228537" ID-ZooBank="53549D58-8F38-47B5-879F-0245E900C131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460444968431" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Métais, Grégoire, Bartolini, Annachiara, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H., Marivaux, Laurent &amp; Welcomme, Jean-Loup" docDate="2014" docId="03A087E00E4FFFB4FF7FFD97FCE1FB2C" docLanguage="en" docName="zt03826p138.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyria" docType="treatment" docVersion="9" lastPageNumber="107" masterDocId="FF99FF980E49FFB2FFE8FFEEFFDAFFAB" masterDocTitle="Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography" masterLastPageNumber="138" masterPageNumber="101" pageNumber="107" updateTime="1698333283847" updateUser="ExternalLinkService">
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<mods:title id="DEF5D42687038A877F1B39D8F52745EC">Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography</mods:title>
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<mods:namePart id="874A4F12CA883506AD645E5DB50E4743">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="B19A2F6A6B5C4F46DB1D543E520E1025">Bartolini, Annachiara</mods:namePart>
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<mods:namePart id="F622F66928B2A3A2D78BC74A80A5F199">Lashari, Rafiq A.</mods:namePart>
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<mods:namePart id="0D96CE3D11F5613A2D9075213C19C8B7">Brohi, Imdad A.</mods:namePart>
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<mods:namePart id="4DCF4CB23056EAF5477A899CEE01BA7A">Solangi, Sarfraz H.</mods:namePart>
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<treatment id="03A087E00E4FFFB4FF7FFD97FCE1FB2C" ID-DOI="http://doi.org/10.5281/zenodo.6137800" ID-GBIF-Taxon="119509275" ID-Zenodo-Dep="6137800" LSID="urn:lsid:plazi:treatment:03A087E00E4FFFB4FF7FFD97FCE1FB2C" httpUri="http://treatment.plazi.org/id/03A087E00E4FFFB4FF7FFD97FCE1FB2C" lastPageNumber="107" pageId="6" pageNumber="107">
<subSubSection id="C313657D0E4FFFB4FF7FFD97FF25FD1F" pageId="6" pageNumber="107" type="nomenclature">
<paragraph id="8BB636F60E4FFFB4FF7FFD97FED8FD38" blockId="6.[151,258,633,693]" box="[151,258,633,659]" pageId="6" pageNumber="107">
<heading id="D0FE819A0E4FFFB4FF7FFD97FED8FD38" bold="true" box="[151,258,633,659]" fontSize="11" level="1" pageId="6" pageNumber="107" reason="1">
<emphasis id="B97DEAE40E4FFFB4FF7FFD97FED8FD38" bold="true" box="[151,258,633,659]" pageId="6" pageNumber="107">
<taxonomicName id="4C094D750E4FFFB4FF7FFD97FF0CFD38" ID-CoL="7P83R" box="[151,214,633,659]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="107" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4FFFB4FF7FFD97FF0CFD38" bold="true" box="[151,214,633,659]" italics="true" pageId="6" pageNumber="107">Lyria</emphasis>
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sp.
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</heading>
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<paragraph id="8BB636F60E4FFFB4FF7FFD72FF25FD1F" blockId="6.[151,258,633,693]" box="[151,255,668,693]" pageId="6" pageNumber="107">
(
<figureCitation id="13322A730E4FFFB4FF77FD72FF2DFD1F" box="[159,247,668,693]" captionStart="FIGURE 4. A" captionStartId="8.[151,250,1824,1846]" captionTargetBox="[252,1335,195,1787]" captionTargetId="figure@8.[252,1335,195,1788]" captionTargetPageId="8" captionText="FIGURE 4. A. Lyria sp., CPAG. RAN. I. 1, stn 5; B H. Mitreola brohii sp. nov., B C (holotype, CPAG. RAN. I. 2, ventral (B) and dorsal (C) views, stn 4), D (paratype 1, CPAG RAN. I. 3, ventral view, stn 4), E F (paratype 2, CPAG. RAN. I. 4, ventral (E) and dorsal (F) views, stn 4), G H (paratype 3, CPAG. RAN. I. 5, juvenile, ventral view (G) and enlarged view of the spire (H), stn 4); I K. Lyriopsis cossmanni (Vredenburg, 1923), I J (CPAG. RAN. I. 6, ventral view (I) and enlarged view of the spire (J), stn 4), K (CPAG. RAN. I. 7, ventral view, stn 4); L. Lyrischapa haimei (dArchiac &amp; Haime, 1853), CPAG. RAN. I. 8, ventral view, stn 6; M N. Lyrischapa sismondai (dArchiac &amp; Haime, 1853), M (CPAG. RAN. I. 10, ventral view, stn 6), N (CPAG. RAN. I. 9, dorsal view, stn 3). Scale bars: B G, I, K N = 10 mm; A, H, J = 5 mm." httpUri="https://zenodo.org/record/228541/files/figure.png" pageId="6" pageNumber="107">Fig. 4A</figureCitation>
)
</paragraph>
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<subSubSection id="C313657D0E4FFFB4FF7FFD0AFCBBFCC3" pageId="6" pageNumber="107" type="description">
<paragraph id="8BB636F60E4FFFB4FF7FFD0AFCBBFCC3" blockId="6.[151,1436,740,1160]" pageId="6" pageNumber="107">
<emphasis id="B97DEAE40E4FFFB4FF7FFD0AFEF3FD56" bold="true" box="[151,297,740,765]" pageId="6" pageNumber="107">Description.</emphasis>
Fragment of a narrow spire, H 14.8, D 9.0 mm. Protoconch eroded, probably paucispiral. Teleoconch of four whorls, the last broken. Whorls high, weakly convex, delineated by a deep, linear suture. Axial sculpture of strong but narrowly rounded costae, almost corresponding from whorl to whorl. No spiral sculpture. Costae opisthocline, rather straight. Columella with at least three folds.
</paragraph>
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<subSubSection id="C313657D0E4FFFB4FF2FFC9AFC44FC26" box="[199,926,884,909]" pageId="6" pageNumber="107" type="materials_examined">
<paragraph id="8BB636F60E4FFFB4FF2FFC9AFC44FC26" blockId="6.[151,1436,740,1160]" box="[199,926,884,909]" pageId="6" pageNumber="107">
<emphasis id="B97DEAE40E4FFFB4FF2FFC9AFEECFC26" bold="true" box="[199,310,884,909]" pageId="6" pageNumber="107">Material.</emphasis>
1 spm (stn 5:
<collectionCode id="ED18AE330E4FFFB4FE3DFC9AFDFDFC27" box="[469,551,884,908]" pageId="6" pageNumber="107">CPAG</collectionCode>
.RAN.
<date id="FFB710360E4FFFB4FD87FC9AFD5EFC27" box="[623,644,884,908]" pageId="6" pageNumber="107">I.1</date>
, cast
<collectionCode id="ED18AE330E4FFFB4FD2AFC9AFCC7FC26" LSID="urn:lsid:biocol.org:col:34988" box="[706,797,884,909]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="107">MNHN</collectionCode>
.F.
<accessionNumber id="945AAB150E4FFFB4FCDCFC9AFC48FC26" box="[820,914,884,909]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A50339" pageId="6" pageNumber="107" type="EnaNcbi">A50339</accessionNumber>
).
</paragraph>
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<subSubSection id="C313657D0E4FFFB4FF2FFC79FCE1FB2C" pageId="6" pageNumber="107" type="discussion">
<paragraph id="8BB636F60E4FFFB4FF2FFC79FD3AFBCE" blockId="6.[151,1436,740,1160]" pageId="6" pageNumber="107">
<emphasis id="B97DEAE40E4FFFB4FF2FFC79FE95FC1B" bold="true" box="[199,335,919,944]" pageId="6" pageNumber="107">Comments.</emphasis>
Although the aperture of this specimen is lacking, the shape of its teleoconch whorls is similar to those found in other members of
<taxonomicName id="4C094D750E4FFFB4FDFDFC53FD8AFC7F" box="[533,592,957,980]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="107" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4FFFB4FDFDFC53FD8AFC7F" box="[533,592,957,980]" italics="true" pageId="6" pageNumber="107">Lyria</emphasis>
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, as for example in
<taxonomicName id="4C094D750E4FFFB4FCD8FC53FB02FC7E" authority="d'Orbigny, 1850" authorityName="d'Orbigny" authorityYear="1850" box="[816,1240,956,981]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="107" phylum="Mollusca" rank="species" species="subturgidula">
<emphasis id="B97DEAE40E4FFFB4FCD8FC53FBD8FC7E" box="[816,1026,956,981]" italics="true" pageId="6" pageNumber="107">Lyria subturgidula</emphasis>
(dOrbigny, 1850)
</taxonomicName>
from the Middle Eocene of the Paris Basin. By its high whorls and its straight costae, this specimen is easily distinguishable from
<taxonomicName id="4C094D750E4FFFB4FF7FFBEBFDFFFBB7" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" box="[151,549,1028,1053]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="107" phylum="Mollusca" rank="species" species="cossmanni">
<emphasis id="B97DEAE40E4FFFB4FF7FFBEBFE8BFBB7" box="[151,337,1029,1052]" italics="true" pageId="6" pageNumber="107">Lyria cossmanni</emphasis>
<bibRefCitation id="EF984B070E4FFFB4FEB2FBEAFDFFFBB7" author="Vredenburg" box="[346,549,1028,1053]" pageId="6" pageNumber="107" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg, 1923</bibRefCitation>
</taxonomicName>
(=?
<taxonomicName id="4C094D750E4FFFB4FDB3FBEBFC7BFBB7" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" box="[603,929,1028,1053]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="107" phylum="Mollusca" rank="species" species="feddeni">
<emphasis id="B97DEAE40E4FFFB4FDB3FBEBFD16FBB6" box="[603,716,1028,1053]" italics="true" pageId="6" pageNumber="107">L. feddeni</emphasis>
<bibRefCitation id="EF984B070E4FFFB4FD3DFBEAFC7BFBB7" author="Vredenburg" box="[725,929,1028,1053]" pageId="6" pageNumber="107" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg, 1923</bibRefCitation>
</taxonomicName>
) from the Lakhra Formation and from
<taxonomicName id="4C094D750E4FFFB4FA88FBEBFE71FBEB" authority="Cox, 1930" authorityName="Cox" authorityYear="1930" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="107" phylum="Mollusca" rank="species" species="samanaensis">
<emphasis id="B97DEAE40E4FFFB4FA88FBEBFEFCFBEB" italics="true" pageId="6" pageNumber="107">Lyria samanaensis</emphasis>
<bibRefCitation id="EF984B070E4FFFB4FEDAFBC6FE71FBEB" author="Cox" box="[306,427,1064,1088]" pageId="6" pageNumber="107" refString="Cox, L. R. (1930) The fossil fauna of the Samana Range and some neighbouring areas. Part 8. The mollusc of the Hangu Shales. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 15, 129 - 222." type="journal article" year="1930">Cox, 1930</bibRefCitation>
</taxonomicName>
from the Hangu Formation (Samana Range, North
<collectingCountry id="F31E76660E4FFFB4FBFCFBC9FBA2FBEB" box="[1044,1144,1063,1088]" name="Pakistan" pageId="6" pageNumber="107">Pakistan</collectingCountry>
). Both species belong to another genus,
<taxonomicName id="4C094D750E4FFFB4FEA9FBA3FE72FBCF" box="[321,424,1101,1124]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4FFFB4FEA9FBA3FE72FBCF" box="[321,424,1101,1124]" italics="true" pageId="6" pageNumber="107">Lyriopsis</emphasis>
</taxonomicName>
nov. gen., described below.
</paragraph>
<paragraph id="8BB636F60E4FFFB4FF2FFB81FCE1FB2C" blockId="6.[151,1436,740,1160]" box="[199,827,1135,1160]" pageId="6" pageNumber="107">
<emphasis id="B97DEAE40E4FFFB4FF2FFB81FE62FB23" bold="true" box="[199,440,1135,1160]" pageId="6" pageNumber="107">Stratigraphic range.</emphasis>
Lakhra Formation: Lakhra Dome.
</paragraph>
</subSubSection>
</treatment>
</document>