209 lines
18 KiB
XML
209 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.4052.3.1" ID-GBIF-Dataset="6293cc47-d833-487e-a6ab-c9853b48b1a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="240598" ID-ZooBank="775796D2-BEDF-459B-B937-FAC0EC457B92" checkinTime="1461350570995" checkinUser="plazi" docAuthor="László, Gyula M., Ronkay, Gábor & Ronkay, László" docDate="2015" docId="0399FA612851FFFBFF2086E1E5206F99" docLanguage="en" docName="zt04052p296.pdf" docOrigin="Zootaxa 4052 (2)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@92d1782" docStyleName="zootaxa.2013.journal_article" docTitle="Wittonola László, Ronkay & Ronkay, 2015, gen. n." docType="treatment" docVersion="9" lastPageNumber="279" masterDocId="FFA08219284AFFE7FFB78042E1256B3A" masterDocTitle="Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species" masterLastPageNumber="296" masterPageNumber="251" pageNumber="278" updateTime="1644950914869" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>László, Gyula M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ronkay, Gábor</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ronkay, László</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>4052</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>251</mods:start>
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<mods:end>296</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.4052.3.1</mods:identifier>
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<mods:identifier type="GBIF-Dataset">6293cc47-d833-487e-a6ab-c9853b48b1a4</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">240598</mods:identifier>
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<mods:identifier type="ZooBank">775796D2-BEDF-459B-B937-FAC0EC457B92</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6093973" ID-GBIF-Taxon="119646470" ID-Zenodo-Dep="6093973" LSID="urn:lsid:plazi:treatment:0399FA612851FFFBFF2086E1E5206F99" httpUri="http://treatment.plazi.org/id/0399FA612851FFFBFF2086E1E5206F99" lastPageId="28" lastPageNumber="279" pageId="27" pageNumber="278">
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<subSubSection pageId="27" pageNumber="278" type="nomenclature">
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<paragraph blockId="27.[151,546,1699,1760]" box="[151,349,1699,1725]" pageId="27" pageNumber="278">
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<heading bold="true" box="[151,349,1699,1725]" fontSize="10" level="2" pageId="27" pageNumber="278" reason="2">
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<emphasis bold="true" box="[151,349,1699,1725]" pageId="27" pageNumber="278">
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<taxonomicName box="[151,264,1699,1724]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[151,264,1699,1724]" italics="true" pageId="27" pageNumber="278">Wittonola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[270,349,1700,1725]" pageId="27" pageNumber="278" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="27.[151,546,1699,1760]" box="[151,546,1735,1760]" pageId="27" pageNumber="278">
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(Plate
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<number box="[223,239,1736,1760]" pageId="27" pageNumber="278" value="4.0">4</number>
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,
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<figureCitation box="[251,352,1735,1760]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 6" captionStartId-0="6.[151,264,2014,2036]" captionStartId-1="8.[151,264,2012,2034]" captionTargetBox-0="[186,1413,195,1976]" captionTargetBox-1="[196,1378,200,1984]" captionTargetId-0="figure@6.[158,1428,178,1992]" captionTargetId-1="figure@8.[158,1428,186,2000]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURES 1 – 3. Male genitalia of Meganola species." captionText-1="FIGURES 4 – 6. Genitalia of Meganola species." httpUri-0="https://zenodo.org/record/240599/files/figure.png" httpUri-1="https://zenodo.org/record/240600/files/figure.png" pageId="27" pageNumber="278">
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Figs
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<number box="[306,352,1735,1760]" pageId="27" pageNumber="278" value="4.5" valueMax="6.0" valueMin="3.0">3–6</number>
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</figureCitation>
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; gen. figs
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<number box="[467,538,1735,1760]" pageId="27" pageNumber="278" value="25.26">25, 26</number>
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)
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</paragraph>
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<paragraph blockId="27.[151,575,1808,1830]" box="[151,575,1808,1830]" pageId="27" pageNumber="278">
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Type-species:
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<taxonomicName box="[296,515,1808,1830]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="Latifasciata" status="sp. n.">
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<emphasis box="[296,515,1808,1830]" italics="true" pageId="27" pageNumber="278">Wittonola latifasciata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[521,575,1809,1830]" pageId="27" pageNumber="278" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="28" lastPageNumber="279" pageId="27" pageNumber="278" type="diagnosis">
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<paragraph blockId="27.[151,1436,1874,2007]" lastBlockId="28.[151,1437,151,1187]" lastPageId="28" lastPageNumber="279" pageId="27" pageNumber="278">
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<emphasis bold="true" box="[151,266,1874,1899]" pageId="27" pageNumber="278">Diagnosis</emphasis>
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. The external appearance of the type-species of the new genus is rather unique, resembling somewhat
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<taxonomicName authority="Swinhoe, 1904" authorityName="Swinhoe" authorityYear="1904" box="[151,540,1910,1935]" class="Insecta" family="Arctiidae" genus="Ctenane" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="Labuana">
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<emphasis box="[151,344,1910,1935]" italics="true" pageId="27" pageNumber="278">Ctenane labuana</emphasis>
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(Swinhoe,
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<number box="[475,532,1911,1935]" pageId="27" pageNumber="278" value="1904.0">1904</number>
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)
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</taxonomicName>
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(Plate
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<number box="[621,637,1911,1935]" pageId="27" pageNumber="278" value="4.0">4</number>
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,
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<figureCitation box="[650,752,1910,1935]" captionStart="FIGURES 7 – 9" captionStartId="10.[151,264,1986,2008]" captionTargetBox="[164,1412,267,1946]" captionTargetId="figure@10.[158,1428,148,1962]" captionTargetPageId="10" captionText="FIGURES 7 – 9. Genitalia of Meganola and Hampsonola species." httpUri="https://zenodo.org/record/240601/files/figure.png" pageId="27" pageNumber="278">
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Figs
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<number box="[707,752,1911,1935]" pageId="27" pageNumber="278" value="7.8">7, 8</number>
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</figureCitation>
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; gen. figs
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<number box="[871,943,1911,1935]" pageId="27" pageNumber="278" value="27.28">27, 28</number>
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) due to the presence of the similar, oblique dark forewing medial stripe and the postmedial line which is replaced by a wavy row of blackish dots. This similarity is only superficial, the genitalia of both sexes show the closer relationship of
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<taxonomicName box="[1128,1237,1982,2007]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[1128,1237,1982,2007]" italics="true" pageId="27" pageNumber="278">Wittonola</emphasis>
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</taxonomicName>
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not with
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<taxonomicName box="[1342,1435,1983,2006]" class="Insecta" family="Arctiidae" genus="Ctenane" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="genus">
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<emphasis box="[1342,1435,1983,2006]" italics="true" pageId="27" pageNumber="278">Ctenane</emphasis>
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</taxonomicName>
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but with the
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<taxonomicName box="[295,409,151,176]" class="Insecta" family="Nolidae" genus="Meganola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[295,409,151,176]" italics="true" pageId="28" pageNumber="279">Meganola</emphasis>
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</taxonomicName>
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lineage, according to the robust, long, evenly tapering, apically pointed uncus, the simple, relatively narrow, slightly arcuate valva with parallel margins and the harpe arising rather medially on the basal part of valva, however the very short, button-like harpe is unknown in
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<taxonomicName box="[956,1070,223,248]" class="Insecta" family="Nolidae" genus="Meganola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[956,1070,223,248]" italics="true" pageId="28" pageNumber="279">Meganola</emphasis>
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</taxonomicName>
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. The generic level separation of
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<taxonomicName box="[151,260,260,285]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[151,260,260,285]" italics="true" pageId="28" pageNumber="279">Wittonola</emphasis>
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</taxonomicName>
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from
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<taxonomicName box="[331,424,261,284]" class="Insecta" family="Arctiidae" genus="Ctenane" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[331,424,261,284]" italics="true" pageId="28" pageNumber="279">Ctenane</emphasis>
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</taxonomicName>
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is supported also by the hindwing venation as in
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<taxonomicName box="[990,1099,260,285]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[990,1099,260,285]" italics="true" pageId="28" pageNumber="279">Wittonola</emphasis>
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</taxonomicName>
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the vein M
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<number box="[1231,1239,275,289]" pageId="28" pageNumber="279" value="3.0">3</number>
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+CuA
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<number box="[1308,1316,275,289]" pageId="28" pageNumber="279" value="1.0">1</number>
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is entirely fused while in
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<taxonomicName box="[316,409,299,322]" class="Insecta" family="Arctiidae" genus="Ctenane" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[316,409,299,322]" italics="true" pageId="28" pageNumber="279">Ctenane</emphasis>
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</taxonomicName>
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they are variably long stalked.
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</paragraph>
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<paragraph blockId="28.[151,1437,151,1187]" pageId="28" pageNumber="279">
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The configuration of the female genitalia displays further distinctive characters between
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<taxonomicName box="[1272,1381,334,359]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[1272,1381,334,359]" italics="true" pageId="28" pageNumber="279">Wittonola</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[151,265,370,395]" class="Insecta" family="Nolidae" genus="Meganola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,265,370,395]" italics="true" pageId="28" pageNumber="279">Meganola</emphasis>
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</taxonomicName>
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.
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<taxonomicName box="[283,392,370,395]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[283,392,370,395]" italics="true" pageId="28" pageNumber="279">Wittonola</emphasis>
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</taxonomicName>
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has narrow, elongate papillae anales (this phenomenon is also known in certain speciesgroups of
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<taxonomicName box="[274,328,406,431]" class="Insecta" family="Nolidae" genus="Nola" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[274,328,406,431]" italics="true" pageId="28" pageNumber="279">Nola</emphasis>
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</taxonomicName>
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) with extraordinarily long apophyses; these characters are unknown in
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<taxonomicName box="[1175,1289,406,431]" class="Insecta" family="Nolidae" genus="Meganola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[1175,1289,406,431]" italics="true" pageId="28" pageNumber="279">Meganola</emphasis>
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</taxonomicName>
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. The single, thorn-like signum of
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<taxonomicName box="[394,503,442,467]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[394,503,442,467]" italics="true" pageId="28" pageNumber="279">Wittonola</emphasis>
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</taxonomicName>
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resembles rather to a signum of the
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<taxonomicName box="[926,1019,442,467]" class="Insecta" family="Nolidae" genus="Manoba" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[926,1019,442,467]" italics="true" pageId="28" pageNumber="279">Manoba</emphasis>
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</taxonomicName>
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species than to any other
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<taxonomicName box="[1322,1436,442,467]" class="Insecta" family="Nolidae" genus="Meganola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[1322,1436,442,467]" italics="true" pageId="28" pageNumber="279">Meganola</emphasis>
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</taxonomicName>
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taxa.
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</paragraph>
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<paragraph blockId="28.[151,1437,151,1187]" pageId="28" pageNumber="279">
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The genus
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<taxonomicName box="[321,430,514,539]" class="Insecta" family="Nolidae" genus="Wittonola" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis box="[321,430,514,539]" italics="true" pageId="28" pageNumber="279">Wittonola</emphasis>
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</taxonomicName>
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shares certain genitalia characters with
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<taxonomicName box="[876,990,514,539]" class="Insecta" family="Nolidae" genus="Meganola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[876,990,514,539]" italics="true" pageId="28" pageNumber="279">Meganola</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[1044,1137,514,539]" class="Insecta" family="Nolidae" genus="Manoba" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
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<emphasis box="[1044,1137,514,539]" italics="true" pageId="28" pageNumber="279">Manoba</emphasis>
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</taxonomicName>
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(Plate
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<number box="[1216,1232,514,539]" pageId="28" pageNumber="279" value="5.0">5</number>
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, figs
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<number box="[1293,1334,515,539]" pageId="28" pageNumber="279" value="1.5" valueMax="2.0" valueMin="1.0">1–2</number>
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and gen. figs
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<number box="[201,272,551,575]" pageId="28" pageNumber="279" value="29.5" valueMax="30.0" valueMin="29.0">29–30</number>
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) but, due to the above combination of the external and genital characters, it cannot be unified with any of them. Thus, this lineage is considered here as a distinct genus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="279" type="description">
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<paragraph blockId="28.[151,1437,151,1187]" pageId="28" pageNumber="279">
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<emphasis bold="true" box="[199,337,622,647]" pageId="28" pageNumber="279">Description</emphasis>
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. The only known species of the genus is rather small in size with the wingspan
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<quantity box="[1233,1354,622,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="28" pageNumber="279" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">11–15 mm</quantity>
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, length of forewing
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<quantity box="[298,398,658,683]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="28" pageNumber="279" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
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. Head and eyes relatively small; antennae bipectinate in male, filiform in female; frons, vertex, tegulae and thorax covered with pale brown scales, collar dark brown. Forewing elongate, apically rounded; ground colour pale brown with reddish sheen; costal margin dark brown from the base to the middle; median area with conspicuous, thick, oblique dark brown stripe; postmedial and terminal areas dark brown; all but one crosslines deleted, postmedial line present but replaced by a row of small dark brown dots. Hindwing pale brown, without discal spot and transverse lines.
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</paragraph>
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<paragraph blockId="28.[151,1437,151,1187]" pageId="28" pageNumber="279">Male genitalia. Uncus rather long, robust, evenly tapering, apically pointed, tegumen short, relatively broad, valva simple, relatively narrow, slightly arcuate, margins parallel, costal sclerotization rather narrow and weak. Harpe very short, button-like, arising in the middle of basal part of valva, vinculum well developed, pointed, Vshaped. Aedeagus simple, tubular, relatively short and thin, without apical carinal process; vesica with small, finger-shaped, finely scobinated section.</paragraph>
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<paragraph blockId="28.[151,1437,151,1187]" pageId="28" pageNumber="279">Female genitalia. Ovipositor relatively long, very narrow, apically expanded, papillae anales very long, thin, eighth tergite rather short, ostium bursae relatively broad, cup-shaped, ductus bursae medium-long, distal half strongly sclerotized, proximal half membraneous, corpus bursae elongate, distal half gradually broadening, proximal one ovoidal; signum bursae single, relatively large, acute, thorn-like.</paragraph>
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</subSubSection>
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</treatment>
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</document> |