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<document id="6B240E77FBB71DBAE91B5197BB736931" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13855FF8629F4FAA9FAEBFA83" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium Motschulsky 1855" docType="key" docVersion="7" lastPageNumber="73" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="72" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="866B1B54FBD134E7D54E49C5D618C3A1">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
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<mods:namePart id="EF3C89757FB540C97CBC2B1BC47C6700">Darby, Michael</mods:namePart>
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<mods:title id="67E48B2A42C23C28C2B835AD63972351">European Journal of Taxonomy</mods:title>
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<mods:date id="C989209D75B6CBF8E4C5E891D5C49215">2020</mods:date>
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<treatment id="039987A13855FF8629F4FAA9FAEBFA83" ID-DOI="http://doi.org/10.5281/zenodo.3795612" ID-GBIF-Taxon="163877767" ID-Zenodo-Dep="3795612" LSID="urn:lsid:plazi:treatment:039987A13855FF8629F4FAA9FAEBFA83" httpUri="http://treatment.plazi.org/id/039987A13855FF8629F4FAA9FAEBFA83" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<subSubSection id="C32A653C3855FF8929F4FAA9FDCFFAF9" box="[189,595,1351,1377]" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="8B8F36B73855FF8929F4FAA9FDCFFAF9" blockId="71.[189,595,1351,1377]" box="[189,595,1351,1377]" pageId="71" pageNumber="72">
<heading id="D0C781DB3855FF8929F4FAA9FDCFFAF9" bold="true" box="[189,595,1351,1377]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
<emphasis id="B944EAA53855FF8929F4FAA9FDCFFAF9" bold="true" box="[189,595,1351,1377]" pageId="71" pageNumber="72">
Key to
<taxonomicName id="4C304D343855FF89285EFAA9FEF1FAF9" ID-CoL="8X77B" authority="Motschulsky, 1855" authorityName="Motschulsky" authorityYear="1855" box="[279,365,1351,1377]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">species</taxonomicName>
in species group 3
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3855FF8629F4FA9BFAEBFA83" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="key">
<key id="FFA161863855FF8629F4FA9BFAEBFA83" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<keyStep id="30C42E123855FF8929F4FA9BFAEBFA6F" pageId="71" pageNumber="72">
<paragraph id="8B8F36B73855FF8929F4FA9BFAEBFA4D" blockId="71.[189,1399,1397,1528]" pageId="71" pageNumber="72">
<keyLead id="30C195823855FF8929F4FA9BFAEBFA4D" pageId="71" pageNumber="72">
1. Pronotum with two linear depressions composed of united foveolae (note:
<taxonomicName id="4C304D343855FF892D28FA98FAEBFA17" ID-CoL="8XXQL" authority="Darby, 2019" authorityName="Darby" authorityYear="2019" box="[1121,1399,1397,1424]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="longum">
<emphasis id="B944EAA53855FF892D28FA98FB40FA17" box="[1121,1244,1397,1424]" italics="true" pageId="71" pageNumber="72">C. longum</emphasis>
<bibRefCitation id="EFA14B463855FF892DACFA9BFAEBFA17" author="Darby M." box="[1253,1399,1397,1423]" pageId="71" pageNumber="72" pagination="1 - 50" refId="ref59424" refString="Darby M. 2019. New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London. European Journal of Taxonomy 512: 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">Darby, 2019</bibRefCitation>
</taxonomicName>
shows traces on one side of the pronotum of a short depression but is not treated in this section of the key, see below)............................................................................................................................. 2
</keyLead>
</paragraph>
<paragraph id="8B8F36B73855FF892988FA30FAEBFA6F" blockId="71.[189,1399,1397,1528]" box="[193,1399,1501,1528]" pageId="71" pageNumber="72">
<keyLead id="30C195823855FF892988FA30FAEBFA6F" box="[193,1399,1501,1528]" pageId="71" pageNumber="72">– Pronotum without linear depressions................................................................................................. 6</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B8F36B73855FF8929F4F9F0FAEBF9C3" blockId="71.[189,1399,1566,1628]" pageId="71" pageNumber="72">
<keyStep id="30C42E123855FF8929F4F9F0FAEBF9C3" pageId="71" pageNumber="72">
<keyLead id="30C195823855FF8929F4F9F0FAEBF9C3" pageId="71" pageNumber="72">
2. Posterior corners of mid-keel rounded (
<figureCitation id="130B2A323855FF892BF0F9F0FCBEF9A0" box="[697,802,1566,1592]" captionStart="Fig" captionStartId="76.[189,232,1964,1990]" captionTargetBox="[202,1398,933,1920]" captionTargetId="figure@76.[189,1398,933,1920]" captionTargetPageId="76" captionText="Fig. 48. Cissidium dewhurst sp. nov. A. Habitus. B. Pronotum, × 530. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 565." figureDoi="http://doi.org/10.5281/zenodo.3777380" httpUri="https://zenodo.org/record/3777380/files/figure.png" pageId="71" pageNumber="72">Fig. 48C</figureCitation>
)..............................................
<taxonomicName id="4C304D343855FF892DCDF9F0FA89F9A1" ID-CoL="8XXPS" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1156,1301,1566,1593]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="dewhurst" status="sp. nov.">
<emphasis id="B944EAA53855FF892DCDF9F0FA89F9A1" bold="true" box="[1156,1301,1566,1593]" italics="true" pageId="71" pageNumber="72">C. dewhurst</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3855FF892C55F9F1FAEBF9A1" box="[1308,1399,1567,1593]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
– Posterior corners of mid-keel with projecting angles ........................................................................ 3
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73855FF8929F4F96CFAEBF958" blockId="71.[189,1399,1666,1728]" pageId="71" pageNumber="72">
<keyStep id="30C42E123855FF8929F4F96CFAEBF958" pageId="71" pageNumber="72">
<keyLead id="30C195823855FF8929F4F96CFAEBF958" pageId="71" pageNumber="72">3. Lateral margins of pronotum rounded ............................................................................................... 4 – Lateral margins of pronotum angulate............................................................................................... 5</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73855FF8929F4F908FAEBF8BB" blockId="71.[189,1399,1765,1827]" pageId="71" pageNumber="72">
<keyStep id="30C42E123855FF8929F4F908FAEBF8BB" pageId="71" pageNumber="72">
<keyLead id="30C195823855FF8929F4F908FAEBF8BB" pageId="71" pageNumber="72">
4. Larger more rounded species......................................................................
<taxonomicName id="4C304D343855FF892D01F908FAEBF898" ID-CoL="8XXQ6" authority="Darby, 2019" authorityName="Darby" authorityYear="2019" box="[1096,1399,1765,1792]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="globulum">
<emphasis id="B944EAA53855FF892D01F908FB43F898" bold="true" box="[1096,1247,1765,1792]" italics="true" pageId="71" pageNumber="72">C. globulum</emphasis>
<bibRefCitation id="EFA14B463855FF892DAFF908FAEBF898" author="Darby M." box="[1254,1399,1766,1792]" pageId="71" pageNumber="72" pagination="1 - 50" refId="ref59424" refString="Darby M. 2019. New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London. European Journal of Taxonomy 512: 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">Darby, 2019</bibRefCitation>
</taxonomicName>
– Smaller, narrower species ....................................................................................
<taxonomicName id="4C304D343855FF892DC5F8E7FA89F8BB" ID-CoL="8XXQZ" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1164,1301,1800,1827]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="murphyi" status="sp. nov.">
<emphasis id="B944EAA53855FF892DC5F8E7FA89F8BB" bold="true" box="[1164,1301,1800,1827]" italics="true" pageId="71" pageNumber="72">C. murphyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3855FF892C55F8E7FAEBF8BB" box="[1308,1399,1801,1827]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73855FF8929F4F8A7FAEBF81F" blockId="71.[189,1399,1865,1927]" pageId="71" pageNumber="72">
<keyStep id="30C42E123855FF8929F4F8A7FAEBF81F" pageId="71" pageNumber="72">
<keyLead id="30C195823855FF8929F4F8A7FAEBF81F" pageId="71" pageNumber="72">
5. Mesoventral mid-keel elongate (
<figureCitation id="130B2A323855FF892B2EF8A4FD4DF8FC" box="[615,721,1865,1892]" captionStart="Fig" captionStartId="87.[189,232,1964,1990]" captionTargetBox="[193,1398,1011,1919]" captionTargetId="figure@87.[189,1398,1005,1919]" captionTargetPageId="87" captionText="Fig. 56. Cissidium mindanao sp. nov. A. Habitus. B. Pronotum, × 430. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 660." figureDoi="http://doi.org/10.5281/zenodo.3777396" httpUri="https://zenodo.org/record/3777396/files/figure.png" pageId="71" pageNumber="72">Fig. 56C</figureCitation>
) ........................................................
<taxonomicName id="4C304D343855FF892D31F8A4FA89F8FC" ID-CoL="8XXQV" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1144,1301,1865,1892]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="mindanao" status="sp. nov.">
<emphasis id="B944EAA53855FF892D31F8A4FA89F8FC" bold="true" box="[1144,1301,1865,1892]" italics="true" pageId="71" pageNumber="72">C. mindanao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3855FF892C55F8A4FAEBF8FC" box="[1308,1399,1866,1892]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
– Mesoventral mid-keel square (
<figureCitation id="130B2A323855FF892B19F882FD25F81F" box="[592,697,1900,1927]" captionStart="Fig" captionStartId="86.[189,232,1964,1990]" captionTargetBox="[194,1398,1012,1922]" captionTargetId="figure@86.[189,1398,1012,1922]" captionTargetPageId="86" captionText="Fig. 55. Cissidium marshallae Darby, 2019. A. Habitus. B. Pronotum, × 640. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 750." figureDoi="http://doi.org/10.5281/zenodo.3777394" httpUri="https://zenodo.org/record/3777394/files/figure.png" pageId="71" pageNumber="72">Fig. 55C</figureCitation>
) ..................................................
<taxonomicName id="4C304D343855FF892D7CF882FAEBF81F" ID-CoL="8XXQR" authority="Darby, 2019" authorityName="Darby" authorityYear="2019" box="[1077,1399,1900,1927]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marshallae">
<emphasis id="B944EAA53855FF892D7CF882FB42F81F" bold="true" box="[1077,1246,1900,1927]" italics="true" pageId="71" pageNumber="72">C. marshallae</emphasis>
<bibRefCitation id="EFA14B463855FF892DAFF882FAEBF81F" author="Darby M." box="[1254,1399,1900,1927]" pageId="71" pageNumber="72" pagination="1 - 50" refId="ref59424" refString="Darby M. 2019. New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London. European Journal of Taxonomy 512: 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">Darby, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73855FF8629F4F843FAEAFED3" blockId="71.[189,1399,1965,2027]" lastBlockId="72.[189,1399,267,331]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<keyStep id="30C42E123855FF8629F4F843FAEAFED3" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<keyLead id="30C195823855FF8629F4F843FAEAFED3" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
6. Pronotal disc with only a trace of foveolae ..................................................
<taxonomicName id="4C304D343855FF892D11F843FA89F85F" ID-CoL="8XXP5" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1112,1301,1965,1991]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="amazonicum" status="sp. nov.">
<emphasis id="B944EAA53855FF892D11F843FA89F85F" bold="true" box="[1112,1301,1965,1991]" italics="true" pageId="71" pageNumber="72">C. amazonicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3855FF892C55F840FAEBF850" box="[1308,1399,1966,1992]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
– Pronotal foveolae covering the disc................................................................................................... 7 7. Lateral margins of pronotum angular ................................................................................................ 8 – Lateral margins of pronotum rounded ............................................................................................. 10
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FE9AFAEBFE2B" blockId="72.[189,1399,371,435]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FE9AFAEBFE2B" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FE9AFAEBFE2B" pageId="72" pageNumber="73">
8. Mesoventral lateral margins rectangular, hind angles acute..........................
<taxonomicName id="4C304D34385AFF862D16FE9AFAEBFE16" ID-CoL="8XXQL" authority="Darby, 2019" authorityName="Darby" authorityYear="2019" box="[1119,1399,371,398]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="longum">
<emphasis id="B944EAA5385AFF862D16FE9AFB42FE16" bold="true" box="[1119,1246,371,398]" italics="true" pageId="72" pageNumber="73">C. longum</emphasis>
<bibRefCitation id="EFA14B46385AFF862DAFFE9AFAEBFE16" author="Darby M." box="[1254,1399,372,398]" pageId="72" pageNumber="73" pagination="1 - 50" refId="ref59424" refString="Darby M. 2019. New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London. European Journal of Taxonomy 512: 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">Darby, 2019</bibRefCitation>
</taxonomicName>
– Mesoventral lateral margins not rectangular .................................................................................... 9
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FE32FAEBFD84" blockId="72.[189,1399,476,540]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FE32FAEBFD84" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FE32FAEBFD84" pageId="72" pageNumber="73">
9. Mid-keel very narrow at junction with collar.........................................
<taxonomicName id="4C304D34385AFF862D60FE32FAEBFE6F" ID-CoL="VK84" authority="Darby, 2015" authorityName="Darby" authorityYear="2015" box="[1065,1399,476,503]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="amboroense">
<emphasis id="B944EAA5385AFF862D60FE32FB43FE6F" bold="true" box="[1065,1247,476,503]" italics="true" pageId="72" pageNumber="73">C. amboroense</emphasis>
<bibRefCitation id="EFA14B46385AFF862DAFFE32FAEBFE6F" author="Darby M." box="[1254,1399,476,503]" pageId="72" pageNumber="73" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby, 2015</bibRefCitation>
</taxonomicName>
– Mid-keel wide at junction with collar..............................................................
<taxonomicName id="4C304D34385AFF862D24FDEFFA89FD84" ID-CoL="8XXRH" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1133,1301,513,540]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="puncticolle" status="sp. nov.">
<emphasis id="B944EAA5385AFF862D24FDEFFA89FD84" bold="true" box="[1133,1301,513,540]" italics="true" pageId="72" pageNumber="73">C. puncticolle</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE385AFF862C55FDECFAEBFD84" box="[1308,1399,514,540]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FDABFAEAFD1C" blockId="72.[189,1398,580,644]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FDABFAEAFD1C" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FDABFAEAFD1C" pageId="72" pageNumber="73">10. Elytra almost parallel-sided in posterior half....................................................................................11 – Elytra sinuate in posterior half ........................................................................................................ 12</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FD43FAEBFD74" blockId="72.[189,1399,684,748]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FD43FAEBFD74" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FD43FAEBFD74" pageId="72" pageNumber="73">
11. Pronotum densely and finely foveolate (
<figureCitation id="130B2A32385AFF862BF6FD43FCB4FD50" box="[703,808,685,712]" captionStart="Fig" captionStartId="84.[189,232,1965,1991]" captionTargetBox="[196,1397,1014,1922]" captionTargetId="figure@84.[189,1398,1014,1922]" captionTargetPageId="84" captionText="Fig. 54. Cissidium mahleri sp. nov. A. Habitus. B. Pronotum, × 630. C. Mesoventrite showing median process of collar, mid-keel and keel, × 545." figureDoi="http://doi.org/10.5281/zenodo.3777392" httpUri="https://zenodo.org/record/3777392/files/figure.png" pageId="72" pageNumber="73">Fig. 54B</figureCitation>
) ................................................
<taxonomicName id="4C304D34385AFF862DDDFD43FA89FD5F" ID-CoL="8XXQQ" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1172,1301,684,711]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="mahleri" status="sp. nov.">
<emphasis id="B944EAA5385AFF862DDDFD43FA89FD5F" bold="true" box="[1172,1301,684,711]" italics="true" pageId="72" pageNumber="73">C. mahleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE385AFF862C55FD43FAEBFD5F" box="[1308,1399,685,711]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
– Pronotum sparsely and more coarsely foveolate ...................................................
<taxonomicName id="4C304D34385AFF862DDFFD3CFA89FD74" ID-CoL="8XXS4" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1174,1301,722,748]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="werneri" status="sp. nov.">
<emphasis id="B944EAA5385AFF862DDFFD3CFA89FD74" bold="true" box="[1174,1301,722,748]" italics="true" pageId="72" pageNumber="73">C. werneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE385AFF862C55FD3CFAEBFD74" box="[1308,1399,722,748]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FCFBFAEAFCCC" blockId="72.[189,1398,789,853]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FCFBFAEAFCCC" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FCFBFAEAFCCC" pageId="72" pageNumber="73">12. Mid-keel not joining collar anteriorly.............................................................................................. 13 – Mid-keel joining collar .................................................................................................................... 14</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FC90FAEBFC25" blockId="72.[189,1399,893,957]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FC90FAEBFC25" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FC90FAEBFC25" pageId="72" pageNumber="73">
13. Smaller species length
<quantity id="4CC89B52385AFF8628B3FC93FD16FC00" box="[506,650,893,920]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="72" pageNumber="73" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
, elytra shorter in proportion to pronotum ..
<taxonomicName id="4C304D34385AFF862DC8FC90FAEBFC00" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1153,1399,893,920]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="B944EAA5385AFF862DC8FC90FB56FC00" bold="true" box="[1153,1226,894,920]" italics="true" pageId="72" pageNumber="73">C. itoi</emphasis>
<bibRefCitation id="EFA14B46385AFF862D9BFC93FAEBFC00" author="Sawada Y." box="[1234,1399,893,920]" pageId="72" pageNumber="73" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
– Larger species length
<quantity id="4CC89B52385AFF8628A5FC4DFDD5FC25" box="[492,585,931,957]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="72" pageNumber="73" unit="mm" value="0.8">0.8 mm</quantity>
, elytra longer in proportion to pronotum ........
<taxonomicName id="4C304D34385AFF862D2BFC4DFAEBFC25" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1122,1399,930,957]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B944EAA5385AFF862D2BFC4DFB56FC25" bold="true" box="[1122,1226,930,957]" italics="true" pageId="72" pageNumber="73">C. latum</emphasis>
<bibRefCitation id="EFA14B46385AFF862D9BFC4CFAEBFC25" author="Sawada Y." box="[1234,1399,930,957]" pageId="72" pageNumber="73" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FC08FAEAFBBE" blockId="72.[189,1399,997,1062]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FC08FAEAFBBE" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FC08FAEAFBBE" pageId="72" pageNumber="73">
14. Mid-keel almost parallel-sided, hind angles rounded.......................................
<taxonomicName id="4C304D34385AFF862D3CFC08FA89FB98" ID-CoL="8XXRD" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1141,1301,997,1024]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pilosellum" status="sp. nov.">
<emphasis id="B944EAA5385AFF862D3CFC08FA89FB98" bold="true" box="[1141,1301,997,1024]" italics="true" pageId="72" pageNumber="73">C. pilosellum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE385AFF862C55FC08FAEBFB98" box="[1308,1399,998,1024]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
– Mid-keel not parallel-sided.............................................................................................................. 15
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FBA0FAEAFB15" blockId="72.[189,1399,1102,1166]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FBA0FAEAFB15" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FBA0FAEAFB15" pageId="72" pageNumber="73">
15. Mid keel tapering from wide base by collar to keel..................................
<taxonomicName id="4C304D34385AFF862D09FBA0FAEBFBF0" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1088,1399,1102,1128]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="B944EAA5385AFF862D09FBA0FB56FBF0" bold="true" box="[1088,1226,1102,1128]" italics="true" pageId="72" pageNumber="73">C. nomurai</emphasis>
<bibRefCitation id="EFA14B46385AFF862D9BFBA0FAEBFBF0" author="Sawada Y." box="[1234,1399,1102,1128]" pageId="72" pageNumber="73" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
– Mid keel otherwise ............................................................................................................................. 16
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4FB58FAEBFA83" blockId="72.[189,1399,1206,1307]" pageId="72" pageNumber="73">
<keyStep id="30C42E12385AFF8629F4FB58FAEBFA83" pageId="72" pageNumber="73">
<keyLead id="30C19582385AFF8629F4FB58FAEBFA83" pageId="72" pageNumber="73">
16. Mid keel with projecting hind angles ........................................................
<taxonomicName id="4C304D34385AFF862D09FB59FAEBFB49" ID-CoL="8XXRB" authority="Darby, 2019" authorityName="Darby" authorityYear="2019" box="[1088,1399,1206,1233]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pauxillum">
<emphasis id="B944EAA5385AFF862D09FB59FB42FB49" bold="true" box="[1088,1246,1206,1233]" italics="true" pageId="72" pageNumber="73">C. pauxillum</emphasis>
<bibRefCitation id="EFA14B46385AFF862DAFFB59FAEBFB49" author="Darby M." box="[1254,1399,1207,1233]" pageId="72" pageNumber="73" pagination="1 - 50" refId="ref59424" refString="Darby M. 2019. New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London. European Journal of Taxonomy 512: 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">Darby, 2019</bibRefCitation>
</taxonomicName>
– Mid keel reduced to small fragments either side of medial extension of the collar ...........................................................................................................
<taxonomicName id="4C304D34385AFF862D0EFAEFFAEBFA83" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1095,1399,1280,1307]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="ikeuchii">
<emphasis id="B944EAA5385AFF862D0EFAEFFB57FA83" bold="true" box="[1095,1227,1280,1307]" italics="true" pageId="72" pageNumber="73">C. ikeuchii</emphasis>
<bibRefCitation id="EFA14B46385AFF862D9BFAEFFAEBFA83" author="Sawada Y." box="[1234,1399,1281,1307]" pageId="72" pageNumber="73" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
</key>
</subSubSection>
</treatment>
</document>
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