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<document id="3193FDF763876BF0A322459D0AEA64CE" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13829FFF22B7DFBFFFD03F9AD" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium okuense Grebennikov 2009" docType="treatment" docVersion="7" lastPageNumber="61" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="60" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8DECC17B445973777FBF1E247DE961E6">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
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<mods:namePart id="FE1FE024C7C0A541F438FF7FFFA84A40">Darby, Michael</mods:namePart>
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<mods:title id="0F50037C180F75ADCCDD4984A94D06BB">European Journal of Taxonomy</mods:title>
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<mods:date id="456D5DE4FA7DA8BD6199C1E2C65D3E34">2020</mods:date>
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<treatment id="039987A13829FFF22B7DFBFFFD03F9AD" ID-DOI="http://doi.org/10.5281/zenodo.3795550" ID-GBIF-Taxon="163877844" ID-Zenodo-Dep="3795550" LSID="urn:lsid:plazi:treatment:039987A13829FFF22B7DFBFFFD03F9AD" httpUri="http://treatment.plazi.org/id/039987A13829FFF22B7DFBFFFD03F9AD" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<subSubSection id="C32A653C3829FFF52B7DFBFFFB9CFBB3" box="[564,1024,1041,1068]" pageId="59" pageNumber="60" type="nomenclature">
<paragraph id="8B8F36B73829FFF52B7DFBFFFB9CFBB3" blockId="59.[564,1024,1041,1102]" box="[564,1024,1041,1068]" pageId="59" pageNumber="60">
<heading id="D0C781DB3829FFF52B7DFBFFFB9CFBB3" box="[564,1024,1041,1068]" centered="true" fontSize="11" level="2" pageId="59" pageNumber="60" reason="2">
<taxonomicName id="4C304D343829FFF52B7DFBFFFB9CFBB3" ID-CoL="VK8K" authority="Grebennikov, 2009" authorityName="Grebennikov" authorityYear="2009" box="[564,1024,1041,1068]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="okuense">
<emphasis id="B944EAA53829FFF52B7DFBFFFC8BFBB4" bold="true" box="[564,791,1041,1068]" italics="true" pageId="59" pageNumber="60">Cissidium okuense</emphasis>
<bibRefCitation id="EFA14B463829FFF52A56FBFFFB9CFBB3" author="Grebennikov" box="[799,1024,1041,1067]" pageId="59" pageNumber="60" pagination="113 - 136" refId="ref59510" refString="Grebennikov 2009. Discheramocephalini, a new pantropical tribe of featherwing beetles (Coleoptera: Ptiliidae): description of new taxa and phylogenetic analysis. Systematic Entomology 34: 113 - 136. https: // doi. org / 10.1111 / j. 1365 - 3113.2008.00444. x" type="journal article" year="2009">Grebennikov, 2009</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3829FFF52BB9FBDAFCDFFBD6" box="[752,835,1076,1102]" pageId="59" pageNumber="60" type="description">
<paragraph id="8B8F36B73829FFF52BB9FBDAFCDFFBD6" blockId="59.[564,1024,1041,1102]" box="[752,835,1076,1102]" pageId="59" pageNumber="60">
<figureCitation id="130B2A323829FFF52BB9FBDAFCDFFBD6" box="[752,835,1076,1102]" captionStart="Fig" captionStartId="59.[189,232,1962,1988]" captionTargetBox="[190,1398,1246,1915]" captionTargetId="figure@59.[189,1398,1245,1926]" captionTargetPageId="59" captionText="Fig. 38. Cissidium okuense Grebennikov, 2009. A. Habitus. B. Pronotum, C. Mesoventrite (fore shortened by camera position, SEM images provided to me by V. Grebennikov)" figureDoi="http://doi.org/10.5281/zenodo.3777360" httpUri="https://zenodo.org/record/3777360/files/figure.png" pageId="59" pageNumber="60">Fig. 38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3829FFF229F4FB9BFD03F9AD" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="materials_examined">
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<heading id="D0C781DB3829FFF529F4FB9BFE36FB17" bold="true" box="[189,426,1141,1167]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="B944EAA53829FFF529F4FB9BFE36FB17" bold="true" box="[189,426,1141,1167]" pageId="59" pageNumber="60">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73829FFF529F4FB4AFC07FB26" blockId="59.[189,923,1188,1215]" box="[189,923,1188,1215]" pageId="59" pageNumber="60">
SEMs of a
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and photographs of a slide of the
<typeStatus id="548B88153829FFF52A7DFB4AFC07FB26" box="[820,923,1188,1214]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
</paragraph>
<caption id="DF4F663F3829FFF529F4F844FBFFF871" ID-DOI="http://doi.org/10.5281/zenodo.3777360" ID-Zenodo-Dep="3777360" httpUri="https://zenodo.org/record/3777360/files/figure.png" pageId="59" pageNumber="60" startId="59.[189,232,1962,1988]" targetBox="[190,1398,1246,1915]" targetPageId="59">
<paragraph id="8B8F36B73829FFF529F4F844FBFFF871" blockId="59.[189,1398,1962,2025]" pageId="59" pageNumber="60">
<emphasis id="B944EAA53829FFF529F4F844FEBCF85C" bold="true" box="[189,288,1962,1988]" pageId="59" pageNumber="60">Fig. 38.</emphasis>
<taxonomicName id="4C304D343829FFF52867F844FC95F85D" authority="Grebennikov, 2009" authorityName="Grebennikov" authorityYear="2009" box="[302,777,1962,1989]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="okuense">
<emphasis id="B944EAA53829FFF52867F844FD8CF85C" box="[302,528,1962,1988]" italics="true" pageId="59" pageNumber="60">Cissidium okuense</emphasis>
<bibRefCitation id="EFA14B463829FFF52B56F844FC95F85D" author="Grebennikov" box="[543,777,1962,1989]" pageId="59" pageNumber="60" pagination="113 - 136" refId="ref59510" refString="Grebennikov 2009. Discheramocephalini, a new pantropical tribe of featherwing beetles (Coleoptera: Ptiliidae): description of new taxa and phylogenetic analysis. Systematic Entomology 34: 113 - 136. https: // doi. org / 10.1111 / j. 1365 - 3113.2008.00444. x" type="journal article" year="2009">Grebennikov, 2009</bibRefCitation>
</taxonomicName>
.
<emphasis id="B944EAA53829FFF52A55F844FCADF85C" bold="true" box="[796,817,1962,1988]" pageId="59" pageNumber="60">A</emphasis>
. Habitus.
<emphasis id="B944EAA53829FFF52AFEF845FC57F85D" bold="true" box="[951,971,1963,1989]" pageId="59" pageNumber="60">B</emphasis>
. Pronotum,
<emphasis id="B944EAA53829FFF52D23F844FBE3F85C" bold="true" box="[1130,1151,1962,1988]" pageId="59" pageNumber="60">C</emphasis>
. Mesoventrite (fore shortened by camera position, SEM images provided to me by V. Grebennikov)
</paragraph>
</caption>
<paragraph id="8B8F36B7382EFFF229F4FEE5FDD2FED3" blockId="60.[189,1398,265,332]" pageId="60" pageNumber="61">
<materialsCitation id="3B583CEA382EFFF229F4FEE5FDD2FED3" ID-GBIF-Occurrence="2609493527" collectingDate="2006-05-06" collectorName="V. Grebennikov" country="Cameroon" county="Lake Oku" elevation="2200" latitude="6.2036333" location="North West province" longLatPrecision="1" longitude="10.459933" municipality="Mt" pageId="60" pageNumber="61" specimenCount="1">
<collectingCountry id="F3277627382EFFF229F4FEE5FEF9FEBD" box="[189,357,267,293]" name="Cameroon" pageId="60" pageNumber="61">CAMEROON</collectingCountry>
•
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,
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and
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;
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,
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;
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a.s.l.
</elevation>
;
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<collectingDate id="EFCAE99F382EFFF229F4FEDFFED2FED3" box="[189,334,305,332]" pageId="60" pageNumber="61" value="2006-05-06">6 May 2006</collectingDate>
</date>
;
<collectorName id="26C55361382EFFF22813FEDCFD8FFED3" box="[346,531,305,332]" pageId="60" pageNumber="61">V. Grebennikov</collectorName>
coll.
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</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FE9BFD8DFE17" blockId="60.[189,529,373,399]" box="[189,529,373,399]" pageId="60" pageNumber="61">
<heading id="D0C781DB382EFFF229F4FE9BFD8DFE17" bold="true" box="[189,529,373,399]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="B944EAA5382EFFF229F4FE9BFD8DFE17" bold="true" box="[189,529,373,399]" pageId="60" pageNumber="61">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FE48FCD7FDA9" blockId="60.[189,1399,422,562]" pageId="60" pageNumber="61">
The following amplifies the description of
<bibRefCitation id="EFA14B46382EFFF22BF8FE48FC3CFE58" author="Grebennikov" box="[689,928,422,448]" pageId="60" pageNumber="61" pagination="113 - 136" refId="ref59510" refString="Grebennikov 2009. Discheramocephalini, a new pantropical tribe of featherwing beetles (Coleoptera: Ptiliidae): description of new taxa and phylogenetic analysis. Systematic Entomology 34: 113 - 136. https: // doi. org / 10.1111 / j. 1365 - 3113.2008.00444. x" type="journal article" year="2009">Grebennikov (2009)</bibRefCitation>
to conform with the other entries in this paper. It is based on the
<typeStatus id="548B8815382EFFF22890FE22FDA2FE7E" box="[473,574,460,486]" pageId="60" pageNumber="61" type="paratype">paratype</typeStatus>
illustrated in 20 SEM images and a single photograph of a slide given to the author by Grebennikov. This specimen was subsequently destroyed in the SEM, but was also used as the basis for Grebennikov’s short textual description.
</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FDB2FD36FDEE" blockId="60.[189,682,603,630]" box="[189,682,603,630]" pageId="60" pageNumber="61">
SIZE. Habitus (
<figureCitation id="130B2A32382EFFF2282EFDB5FE48FDEE" box="[359,468,603,630]" captionStart="Fig" captionStartId="59.[189,232,1962,1988]" captionTargetBox="[190,1398,1246,1915]" captionTargetId="figure@59.[189,1398,1245,1926]" captionTargetPageId="59" captionText="Fig. 38. Cissidium okuense Grebennikov, 2009. A. Habitus. B. Pronotum, C. Mesoventrite (fore shortened by camera position, SEM images provided to me by V. Grebennikov)" figureDoi="http://doi.org/10.5281/zenodo.3777360" httpUri="https://zenodo.org/record/3777360/files/figure.png" pageId="60" pageNumber="61">Fig. 38A</figureCitation>
), length
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.
</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FD4EFE4EFD21" blockId="60.[189,466,671,698]" box="[189,466,671,698]" pageId="60" pageNumber="61">COLOUR. Yellow brown.</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FD0AFB70FCBB" blockId="60.[189,1399,739,804]" pageId="60" pageNumber="61">
HEAD. With a shallow median depression between the eyes; width across eyes
<quantity id="4CC89B52382EFFF22D13FD0DFB5BFD66" box="[1114,1223,739,766]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="60" pageNumber="61" unit="mm" value="0.21">0.21 mm</quantity>
; antennomeres III–XI length
<quantity id="4CC89B52382EFFF22829FCE7FE50FCBC" box="[352,460,777,804]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="60" pageNumber="61" unit="mm" value="0.39">0.39 mm</quantity>
, III–IX length
<quantity id="4CC89B52382EFFF22B33FCE7FD79FCBC" box="[634,741,777,804]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="60" pageNumber="61" unit="mm" value="0.2">0.20 mm</quantity>
, X–XI length
<quantity id="4CC89B52382EFFF22AC5FCE7FC6BFCBC" box="[908,1015,777,804]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="60" pageNumber="61" unit="mm" value="0.19">0.19 mm</quantity>
; mentum as
<figureCitation id="130B2A32382EFFF22DC2FCE7FB70FCBB" box="[1163,1260,777,803]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="60" pageNumber="61">Fig. 4A.</figureCitation>
</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FCA0FD73FC66" blockId="60.[189,1399,845,1022]" pageId="60" pageNumber="61">
PRONOTUM. Length
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, width
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, sparsely pubescent, without foveolae but with a row of ± ten foveae along the posterior margin in two groups of five, the outermost two in each group smaller than the others; sides with rounded medial angles, straight to obtuse anterior angles, almost parallelsided to hind angles, bordered, the border not continuing along the posterior margin which has a sinuous emargination opposite the scutellum (
<figureCitation id="130B2A32382EFFF22B3EFC0AFD7FFC66" box="[631,739,996,1022]" captionStart="Fig" captionStartId="59.[189,232,1962,1988]" captionTargetBox="[190,1398,1246,1915]" captionTargetId="figure@59.[189,1398,1245,1926]" captionTargetPageId="59" captionText="Fig. 38. Cissidium okuense Grebennikov, 2009. A. Habitus. B. Pronotum, C. Mesoventrite (fore shortened by camera position, SEM images provided to me by V. Grebennikov)" figureDoi="http://doi.org/10.5281/zenodo.3777360" httpUri="https://zenodo.org/record/3777360/files/figure.png" pageId="60" pageNumber="61">Fig. 38B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FBC7FEABFBF0" blockId="60.[189,1399,1064,1128]" pageId="60" pageNumber="61">
ELYTRA. Length
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, width
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, sparsely pubescent and foveolate contrasting with the pronotum.
</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FB7CFE1DFADC" blockId="60.[189,1399,1170,1348]" pageId="60" pageNumber="61">
MESOVENTRITE. Collar with a short, strongly tapering median extension widening posteriorly; mid-keel wide, parallel-sided, the posterior angles with narrow carinae to the mesocoxal anterior borders; keel with a single medial seta, slightly sinuate before termination at interruption point of the mesocoxal borders; mesoventral lateral margins serrate posteriorly; humeri toothed (
<figureCitation id="130B2A32382EFFF22D66FAEDFB06FA85" box="[1071,1178,1283,1309]" captionStart="Fig" captionStartId="59.[189,232,1962,1988]" captionTargetBox="[190,1398,1246,1915]" captionTargetId="figure@59.[189,1398,1245,1926]" captionTargetPageId="59" captionText="Fig. 38. Cissidium okuense Grebennikov, 2009. A. Habitus. B. Pronotum, C. Mesoventrite (fore shortened by camera position, SEM images provided to me by V. Grebennikov)" figureDoi="http://doi.org/10.5281/zenodo.3777360" httpUri="https://zenodo.org/record/3777360/files/figure.png" pageId="60" pageNumber="61">Fig. 38C</figureCitation>
fore-shortened by camera position)
</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FA83FEEEFA35" blockId="60.[189,1398,1389,1453]" pageId="60" pageNumber="61">
METAVENTRITE. Glabrous, length
<quantity id="4CC89B52382EFFF22B7BFA83FD07FA1F" box="[562,667,1389,1415]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="60" pageNumber="61" unit="mm" value="0.21">0.21 mm</quantity>
, disc simple, width across spines
<quantity id="4CC89B52382EFFF22D51FA83FB1EFA1F" box="[1048,1154,1389,1415]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="60" pageNumber="61" unit="mm" value="0.12">0.12 mm</quantity>
; mesocoxal posterior borders serrate.
</paragraph>
<paragraph id="8B8F36B7382EFFF229F4FA39FE15FA69" blockId="60.[189,393,1495,1521]" box="[189,393,1495,1521]" pageId="60" pageNumber="61">WINGS. Apterous.</paragraph>
<paragraph id="8B8F36B7382EFFF229F4F9F5FD03F9AD" blockId="60.[189,671,1563,1589]" box="[189,671,1563,1589]" pageId="60" pageNumber="61">GENITALIA. Female spermatheca globular.</paragraph>
</subSubSection>
</treatment>
</document>
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