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<document id="185293B89D7323C6D33A5F6E2B0B3EFA" ID-DOI="10.5281/zenodo.205995" ID-GBIF-Dataset="556343c6-a289-4d17-841e-ef4666ca7406" ID-ISSN="1175-5326" ID-Zenodo-Dep="205995" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460274268212" checkinUser="plazi" docAuthor="Meyer, Harry A." docDate="2011" docId="0399878AFFD6FF8758B6C3BCFC2E3B27" docLanguage="en" docName="zt02812p040.pdf" docOrigin="Zootaxa 2812" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Doryphoribius tessellatus Meyer, 2011, sp. n." docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="FFA0FFF2FFD5FF815821C179FFD2387B" masterDocTitle="Tardigrada of Grand Cayman, West Indies, with descriptions of two new species of eutardigrade, Doryphoribius tessellatus (Hypsibiidae) and Macrobiotus caymanensis (Macrobiotidae)" masterLastPageNumber="40" masterPageNumber="28" pageNumber="31" updateTime="1698247824567" updateUser="plazi">
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<mods:title id="E3246C4ACFD1B3EA7AB31707B45454C0">Tardigrada of Grand Cayman, West Indies, with descriptions of two new species of eutardigrade, Doryphoribius tessellatus (Hypsibiidae) and Macrobiotus caymanensis (Macrobiotidae)</mods:title>
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<mods:namePart id="B06F801CE34BE39F42FDC347C4B67168">Meyer, Harry A.</mods:namePart>
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<mods:date id="8077288A87DF0859007DF49BB68F44AA">2011</mods:date>
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<treatment id="0399878AFFD6FF8758B6C3BCFC2E3B27" ID-DOI="http://doi.org/10.5281/zenodo.5684273" ID-GBIF-Taxon="119395751" ID-Zenodo-Dep="5684273" LSID="urn:lsid:plazi:treatment:0399878AFFD6FF8758B6C3BCFC2E3B27" httpUri="http://treatment.plazi.org/id/0399878AFFD6FF8758B6C3BCFC2E3B27" lastPageId="6" lastPageNumber="34" pageId="3" pageNumber="31">
<subSubSection id="C32A6517FFD6FF8258B6C3BCFECC3B7A" pageId="3" pageNumber="31" type="nomenclature">
<paragraph id="8B8F369CFFD6FF8258B6C3BCFDD03AA6" blockId="3.[151,514,709,769]" box="[151,514,709,734]" pageId="3" pageNumber="31">
<heading id="D0C781F0FFD6FF8258B6C3BCFDD03AA6" bold="true" box="[151,514,709,734]" fontSize="10" level="2" pageId="3" pageNumber="31" reason="2">
<emphasis id="B944EA8EFFD6FF8258B6C3BCFDD03AA6" bold="true" box="[151,514,709,734]" pageId="3" pageNumber="31">
<taxonomicName id="4C304D1FFFD6FF8258B6C3BCFE6F3AA5" box="[151,445,709,734]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="31" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD6FF8258B6C3BCFE6F3AA5" bold="true" box="[151,445,709,734]" italics="true" pageId="3" pageNumber="31">Doryphoribius tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD6FF8259E2C3BCFDD03AA6" box="[451,514,709,733]" pageId="3" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F369CFFD6FF8258B6C390FECC3B7A" blockId="3.[151,514,709,769]" box="[151,286,745,769]" pageId="3" pageNumber="31">
(
<figureCitation id="130B2A19FFD6FF8258BEC390FEC43B7A" box="[159,278,745,769]" captionStart-0="FIGURE 2 3" captionStart-1="FIGURES 4 7" captionStart-2="FIGURES 8 11" captionStartId-0="3.[151,249,1975,1998]" captionStartId-1="4.[151,264,1194,1217]" captionStartId-2="5.[151,264,1661,1684]" captionTargetBox-0="[172,1424,1105,1937]" captionTargetBox-1="[207,1375,193,1163]" captionTargetBox-2="[168,1417,456,1627]" captionTargetId-0="figure@3.[151,1436,1074,1954]" captionTargetId-1="figure@4.[196,1392,182,1173]" captionTargetId-2="figure@5.[151,1436,445,1641]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2 3. Doryphoribius tessellatus sp. n. (2) Habitus, phase contrast microscopy. Scale bar: 100 μm. (3) Schematic pattern of gibbosities." captionText-1="FIGURES 4 7. Doryphoribius tessellatus sp. n. (4) Anterior gibbosities. (5) Posterior gibbosities. (6) Reticular design, middorsal region. (7) Reticular design, anterior region of the body. Scale bars: 4 6, 10 μm; 7, 5 μm." captionText-2="FIGURES 8 11. Doryphoribius tessellatus sp. n. (8) Buccopharyngeal apparatus. (9) Buccal cavity armature, dorsal view. (10) Claw, leg I. (11) Claw, leg IV. Scale bars: 8, 20 μm; 9, 5 μm; 10 11, 10 μm." httpUri-0="https://zenodo.org/record/205997/files/figure.png" httpUri-1="https://zenodo.org/record/205998/files/figure.png" httpUri-2="https://zenodo.org/record/205999/files/figure.png" pageId="3" pageNumber="31">Figs. 211</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFD6FF8258B6C248FD7A3BEA" pageId="3" pageNumber="31" type="diagnosis">
<paragraph id="8B8F369CFFD6FF8258B6C248FD7A3BEA" blockId="3.[151,1436,817,1021]" pageId="3" pageNumber="31">
<emphasis id="B944EA8EFFD6FF8258B6C248FEC33B31" bold="true" box="[151,273,817,842]" pageId="3" pageNumber="31">Diagnosis.</emphasis>
A
<taxonomicName id="4C304D1FFFD6FF825914C248FE083B32" box="[309,474,817,841]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="31" phylum="Tardigrada" rank="genus">
<emphasis id="B944EA8EFFD6FF825914C248FE083B32" box="[309,474,817,841]" italics="true" pageId="3" pageNumber="31">Doryphoribius</emphasis>
</taxonomicName>
of the
<taxonomicName id="4C304D1FFFD6FF825A0AC248FD5A3B32" box="[555,648,817,841]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="31" phylum="Tardigrada" rank="species" species="evelinae">
<emphasis id="B944EA8EFFD6FF825A0AC248FD5A3B32" box="[555,648,817,841]" italics="true" pageId="3" pageNumber="31">evelinae</emphasis>
</taxonomicName>
-group, with two macroplacoids, buccal cavity armature comprised of six large, round dorsal teeth in two rows, dorsolateral gibbosities in sequence III:4:2:2 and a cuticle covered by shallow depressions forming a reticular design.
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFD6FF8258E4C2E4FD5B3B86" pageId="3" pageNumber="31" type="materials_examined">
<paragraph id="8B8F369CFFD6FF8258E4C2E4FD5B3B86" blockId="3.[151,1436,817,1021]" pageId="3" pageNumber="31">
<emphasis id="B944EA8EFFD6FF8258E4C2E4FE613BCD" bold="true" box="[197,435,925,950]" pageId="3" pageNumber="31">Material examined.</emphasis>
<typeStatus id="548B883EFFD6FF82599EC2E4FDF83BCE" box="[447,554,925,949]" pageId="3" pageNumber="31" type="holotype">Holotype</typeStatus>
, one
<typeStatus id="548B883EFFD6FF825A4EC2E7FD023BCD" box="[623,720,926,950]" pageId="3" pageNumber="31" type="paratype">paratype</typeStatus>
and one contracted juvenile specimen: sample 4c (
<tableCitation id="C6B20327FFD6FF825D13C2E4FA583BCE" box="[1330,1418,925,949]" captionStart="TABLE 1" captionStartId="1.[151,235,506,527]" captionTargetBox="[151,1436,592,1919]" captionTargetPageId="1" captionText="TABLE 1. Samples containing terrestrial tardigrades from Grand Cayman, Cayman Islands, West Indies. Letters after site num-" httpUri="http://table.plazi.org/id/DF4F6614FFD4FF8058B6C083FA4E3A6B" pageId="3" pageNumber="31" tableUuid="DF4F6614FFD4FF8058B6C083FA4E3A6B">Table 1</tableCitation>
),
<taxonomicName id="4C304D1FFFD6FF8258B6C2B8FE933BA2" authority="Reserve" authorityName="Reserve" box="[151,321,961,985]" class="Entognatha" family="Paronellidae" genus="Salina" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="31" phylum="Arthropoda" rank="genus">Salina Reserve</taxonomicName>
, East
<collectingCountry id="F327760CFFD6FF8259A4C2B8FE603BA2" box="[389,434,961,985]" name="India" pageId="3" pageNumber="31">End</collectingCountry>
District, Grand
<collectingCountry id="F327760CFFD6FF825A4BC2B8FD183BA2" box="[618,714,961,985]" name="Cayman Islands" pageId="3" pageNumber="31">Cayman</collectingCountry>
,
<collectingCountry id="F327760CFFD6FF825AF6C2B8FC5F3BA2" box="[727,909,961,985]" name="Cayman Islands" pageId="3" pageNumber="31">Cayman Islands</collectingCountry>
, West Indies, 1921ʹ0 0.3ʺ N and 8105ʹ48.7ʺ W,
<quantity id="4CC89B79FFD6FF8258B6C29CFF023B86" box="[151,208,997,1021]" metricMagnitude="0" metricUnit="m" metricValue="9.1" pageId="3" pageNumber="31" unit="m" value="9.1">9.1m</quantity>
asl, dry tropical forest, crustose lichen.
</paragraph>
</subSubSection>
<caption id="DF4F6614FFD6FF8258B6C6CEFE873F97" httpUri="https://zenodo.org/record/205997/files/figure.png" pageId="3" pageNumber="31" targetBox="[172,1424,1105,1937]" targetPageId="3">
<paragraph id="8B8F369CFFD6FF8258B6C6CEFE873F97" blockId="3.[151,1435,1975,2028]" pageId="3" pageNumber="31">
<emphasis id="B944EA8EFFD6FF8258B6C6CEFEFF3FB6" bold="true" box="[151,301,1975,1998]" pageId="3" pageNumber="31">FIGURE 23.</emphasis>
<taxonomicName id="4C304D1FFFD6FF825912C6C1FDEB3FB5" box="[307,569,1976,1998]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="31" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD6FF825912C6C1FDEB3FB5" box="[307,569,1976,1998]" italics="true" pageId="3" pageNumber="31">Doryphoribius tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD6FF825A1EC6C1FDAB3FB6" box="[575,633,1976,1997]" pageId="3" pageNumber="31" rank="species">
<emphasis id="B944EA8EFFD6FF825A1EC6C1FDAB3FB6" bold="true" box="[575,633,1976,1997]" pageId="3" pageNumber="31">sp. n.</emphasis>
</taxonomicNameLabel>
(2) Habitus, phase contrast microscopy. Scale bar: 100 μm. (3) Schematic pattern of gibbosities.
</paragraph>
</caption>
<caption id="DF4F6614FFD1FF8558B6C5D3FBB13CA4" httpUri="https://zenodo.org/record/205998/files/figure.png" pageId="4" pageNumber="32" targetBox="[207,1375,193,1163]" targetPageId="4">
<paragraph id="8B8F369CFFD1FF8558B6C5D3FBB13CA4" blockId="4.[151,1436,1194,1248]" pageId="4" pageNumber="32">
<emphasis id="B944EA8EFFD1FF8558B6C5D3FEEE3CBB" bold="true" box="[151,316,1194,1217]" pageId="4" pageNumber="32">FIGURES 47.</emphasis>
<taxonomicName id="4C304D1FFFD1FF855962C5D3FD9B3CBB" box="[323,585,1194,1216]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="32" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD1FF855962C5D3FD9B3CBB" box="[323,585,1194,1216]" italics="true" pageId="4" pageNumber="32">Doryphoribius tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD1FF855A71C5D2FD583CBB" box="[592,650,1195,1216]" pageId="4" pageNumber="32" rank="species">
<emphasis id="B944EA8EFFD1FF855A71C5D2FD583CBB" bold="true" box="[592,650,1195,1216]" pageId="4" pageNumber="32">sp. n.</emphasis>
</taxonomicNameLabel>
(4) Anterior gibbosities. (5) Posterior gibbosities. (6) Reticular design, middorsal region. (7) Reticular design, anterior region of the body. Scale bars: 46, 10 μm; 7, 5 μm.
</paragraph>
</caption>
<subSubSection id="C32A6517FFD1FF8458E4C47FFC1939FC" lastPageId="5" lastPageNumber="33" pageId="4" pageNumber="32" type="description">
<paragraph id="8B8F369CFFD1FF8558E4C47FFD913E45" blockId="4.[151,1437,1286,2031]" pageId="4" pageNumber="32">
<emphasis id="B944EA8EFFD1FF8558E4C47FFE843D64" bold="true" box="[197,342,1286,1311]" pageId="4" pageNumber="32">Description.</emphasis>
Holotype—Body length 448.9 μm. Sex undetermined. Body yellow in live animal, eyes absent (
<figureCitation id="130B2A19FFD1FF8558BEC453FF373D39" box="[159,229,1322,1346]" captionStart="FIGURE 2 3" captionStartId="3.[151,249,1975,1998]" captionTargetBox="[172,1424,1105,1937]" captionTargetId="figure@3.[151,1436,1074,1954]" captionTargetPageId="3" captionText="FIGURE 2 3. Doryphoribius tessellatus sp. n. (2) Habitus, phase contrast microscopy. Scale bar: 100 μm. (3) Schematic pattern of gibbosities." httpUri="https://zenodo.org/record/205997/files/figure.png" pageId="4" pageNumber="32">Fig. 2</figureCitation>
). Dorsal cuticle with 16 transverse undulations. Eight dorsolateral gibbosities in three rows, sequence III:4- 2-2 (
<figureCitation id="130B2A19FFD1FF8558EDC437FEC33D1D" box="[204,273,1358,1382]" captionStart="FIGURE 2 3" captionStartId="3.[151,249,1975,1998]" captionTargetBox="[172,1424,1105,1937]" captionTargetId="figure@3.[151,1436,1074,1954]" captionTargetPageId="3" captionText="FIGURE 2 3. Doryphoribius tessellatus sp. n. (2) Habitus, phase contrast microscopy. Scale bar: 100 μm. (3) Schematic pattern of gibbosities." httpUri="https://zenodo.org/record/205997/files/figure.png" pageId="4" pageNumber="32">Fig. 3</figureCitation>
). Anterior row in second undulation with four low gibbosities in two partially fused pairs (
<figureCitation id="130B2A19FFD1FF855D37C437FA8E3D1D" box="[1302,1372,1358,1382]" captionStart="FIGURES 4 7" captionStartId="4.[151,264,1194,1217]" captionTargetBox="[207,1375,193,1163]" captionTargetId="figure@4.[196,1392,182,1173]" captionTargetPageId="4" captionText="FIGURES 4 7. Doryphoribius tessellatus sp. n. (4) Anterior gibbosities. (5) Posterior gibbosities. (6) Reticular design, middorsal region. (7) Reticular design, anterior region of the body. Scale bars: 4 6, 10 μm; 7, 5 μm." httpUri="https://zenodo.org/record/205998/files/figure.png" pageId="4" pageNumber="32">Fig. 4</figureCitation>
); two caudal rows, each with two prominent gibbosities (
<figureCitation id="130B2A19FFD1FF855AC5C40BFCF83DF1" box="[740,810,1394,1418]" captionStart="FIGURES 4 7" captionStartId="4.[151,264,1194,1217]" captionTargetBox="[207,1375,193,1163]" captionTargetId="figure@4.[196,1392,182,1173]" captionTargetPageId="4" captionText="FIGURES 4 7. Doryphoribius tessellatus sp. n. (4) Anterior gibbosities. (5) Posterior gibbosities. (6) Reticular design, middorsal region. (7) Reticular design, anterior region of the body. Scale bars: 4 6, 10 μm; 7, 5 μm." httpUri="https://zenodo.org/record/205998/files/figure.png" pageId="4" pageNumber="32">Fig. 5</figureCitation>
). Dorsal cuticle, including undulations and gibbosities covered by shallow depressions forming a reticular design. Reticular mesh in caudal gibbosities and posterior ten undulations composed mostly of large depressions, 2.64.5 μm wide in gibbosities, 5.58.1 μm wide in undulations (
<figureCitation id="130B2A19FFD1FF8558BEC4A7FF373D8D" box="[159,229,1502,1526]" captionStart="FIGURES 4 7" captionStartId="4.[151,264,1194,1217]" captionTargetBox="[207,1375,193,1163]" captionTargetId="figure@4.[196,1392,182,1173]" captionTargetPageId="4" captionText="FIGURES 4 7. Doryphoribius tessellatus sp. n. (4) Anterior gibbosities. (5) Posterior gibbosities. (6) Reticular design, middorsal region. (7) Reticular design, anterior region of the body. Scale bars: 4 6, 10 μm; 7, 5 μm." httpUri="https://zenodo.org/record/205998/files/figure.png" pageId="4" pageNumber="32">Fig. 6</figureCitation>
); in anterior gibbosities and anterior five undulations composed of large depressions (2.25.5 μm wide in gibbosities, 3.77.6 μm wide in undulations) intermixed with areas filled by very small depressions (&lt;1.0 μm wide) (
<figureCitation id="130B2A19FFD1FF8558C7C75FFEF83E45" box="[230,298,1574,1598]" captionStart="FIGURES 4 7" captionStartId="4.[151,264,1194,1217]" captionTargetBox="[207,1375,193,1163]" captionTargetId="figure@4.[196,1392,182,1173]" captionTargetPageId="4" captionText="FIGURES 4 7. Doryphoribius tessellatus sp. n. (4) Anterior gibbosities. (5) Posterior gibbosities. (6) Reticular design, middorsal region. (7) Reticular design, anterior region of the body. Scale bars: 4 6, 10 μm; 7, 5 μm." httpUri="https://zenodo.org/record/205998/files/figure.png" pageId="4" pageNumber="32">Fig. 7</figureCitation>
). Ventral cuticle smooth.
</paragraph>
<paragraph id="8B8F369CFFD1FF8558E4C733FAF23F41" blockId="4.[151,1437,1286,2031]" pageId="4" pageNumber="32">
Buccopharyngeal apparatus of
<taxonomicName id="4C304D1FFFD1FF855A08C732FD1C3E18" box="[553,718,1611,1635]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="32" phylum="Tardigrada" rank="genus">
<emphasis id="B944EA8EFFD1FF855A08C732FD1C3E18" box="[553,718,1611,1635]" italics="true" pageId="4" pageNumber="32">Doryphoribius</emphasis>
</taxonomicName>
<typeStatus id="548B883EFFD1FF855AF9C732FCDA3E18" box="[728,776,1611,1635]" pageId="4" pageNumber="32">type</typeStatus>
(
<figureCitation id="130B2A19FFD1FF855B3BC733FCB33E19" box="[794,865,1610,1634]" captionStart="FIGURES 8 11" captionStartId="5.[151,264,1661,1684]" captionTargetBox="[168,1417,456,1627]" captionTargetId="figure@5.[151,1436,445,1641]" captionTargetPageId="5" captionText="FIGURES 8 11. Doryphoribius tessellatus sp. n. (8) Buccopharyngeal apparatus. (9) Buccal cavity armature, dorsal view. (10) Claw, leg I. (11) Claw, leg IV. Scale bars: 8, 20 μm; 9, 5 μm; 10 11, 10 μm." httpUri="https://zenodo.org/record/205999/files/figure.png" pageId="4" pageNumber="32">Fig. 8</figureCitation>
). Peribuccal lamellae and papulae absent. Buccal cavity armature composed of six large round teeth on dorsal wall in two rows of three (
<figureCitation id="130B2A19FFD1FF855C49C717FB7F3EFD" box="[1128,1197,1646,1670]" captionStart="FIGURES 8 11" captionStartId="5.[151,264,1661,1684]" captionTargetBox="[168,1417,456,1627]" captionTargetId="figure@5.[151,1436,445,1641]" captionTargetPageId="5" captionText="FIGURES 8 11. Doryphoribius tessellatus sp. n. (8) Buccopharyngeal apparatus. (9) Buccal cavity armature, dorsal view. (10) Claw, leg I. (11) Claw, leg IV. Scale bars: 8, 20 μm; 9, 5 μm; 10 11, 10 μm." httpUri="https://zenodo.org/record/205999/files/figure.png" pageId="4" pageNumber="32">Fig. 9</figureCitation>
). No other teeth visible with light microscopy. Mouth antero-ventral. Buccal tube 43.6 μm long and 4.7 μm wide [
<emphasis id="B944EA8EFFD1FF855CE3C7EAFB213ED0" box="[1218,1267,1683,1707]" italics="true" pageId="4" pageNumber="32">10.8</emphasis>
], with anterior bend. Triangular pharyngeal apophyses present at end of buccal tube. Ventral lamina 24.3 μm [
<emphasis id="B944EA8EFFD1FF855CF7C7CEFB283EB5" box="[1238,1274,1719,1742]" italics="true" pageId="4" pageNumber="32">55.</emphasis>
7] long. Stylet supports inserted on buccal tube at 32.6 μm [
<emphasis id="B944EA8EFFD1FF855A82C7A2FD143E88" box="[675,710,1755,1779]" italics="true" pageId="4" pageNumber="32">74.</emphasis>
8]. Pharyngeal bulb oval with two macroplacoids decreasing in length. Microplacoid and septulum absent. First macroplacoid 8.7 μm [
<emphasis id="B944EA8EFFD1FF855B93C787FC313F6D" box="[946,995,1790,1814]" italics="true" pageId="4" pageNumber="32">20.0</emphasis>
] long with central constriction. Second macroplacoid 5.0 μm [
<emphasis id="B944EA8EFFD1FF8559B4C65AFE103F40" box="[405,450,1827,1851]" italics="true" pageId="4" pageNumber="32">11.5</emphasis>
] long with subterminal constriction. Macroplacoid row 14.9 μm [
<emphasis id="B944EA8EFFD1FF855C84C65AFB043F40" box="[1189,1238,1827,1851]" italics="true" pageId="4" pageNumber="32">34.2</emphasis>
] long.
</paragraph>
<paragraph id="8B8F369CFFD1FF8558E4C63FFA403F94" blockId="4.[151,1437,1286,2031]" pageId="4" pageNumber="32">
Gibbosities and other cuticular thickenings absent on legs. Claws stout, of
<taxonomicName id="4C304D1FFFD1FF855C3FC63FFB783F25" box="[1054,1194,1862,1886]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="32" phylum="Tardigrada" rank="genus">
<emphasis id="B944EA8EFFD1FF855C3FC63FFB783F25" box="[1054,1194,1862,1886]" italics="true" pageId="4" pageNumber="32">Isohypsibius</emphasis>
</taxonomicName>
-
<typeStatus id="548B883EFFD1FF855C90C63EFB373F24" box="[1201,1253,1863,1887]" pageId="4" pageNumber="32">type</typeStatus>
, with small and smooth lunules (
<figureCitation id="130B2A19FFD1FF855971C613FE0B3FF9" box="[336,473,1898,1922]" captionStart="FIGURES 8 11" captionStartId="5.[151,264,1661,1684]" captionTargetBox="[168,1417,456,1627]" captionTargetId="figure@5.[151,1436,445,1641]" captionTargetPageId="5" captionText="FIGURES 8 11. Doryphoribius tessellatus sp. n. (8) Buccopharyngeal apparatus. (9) Buccal cavity armature, dorsal view. (10) Claw, leg I. (11) Claw, leg IV. Scale bars: 8, 20 μm; 9, 5 μm; 10 11, 10 μm." httpUri="https://zenodo.org/record/205999/files/figure.png" pageId="4" pageNumber="32">Figs. 1011</figureCitation>
). Claw lengths: external claw, leg I basal claw 8.8 μm [
<emphasis id="B944EA8EFFD1FF855C65C612FBA73FF8" box="[1092,1141,1899,1923]" italics="true" pageId="4" pageNumber="32">20.2</emphasis>
], primary branch 15.1 μm [
<emphasis id="B944EA8EFFD1FF855881C6F7FF033FDD" box="[160,209,1934,1958]" italics="true" pageId="4" pageNumber="32">34.6</emphasis>
], secondary branch 11.7 μm [
<emphasis id="B944EA8EFFD1FF855A01C6F7FD833FDD" box="[544,593,1934,1958]" italics="true" pageId="4" pageNumber="32">26.8</emphasis>
]; internal claw, leg I, bc 8.0 μm [
<emphasis id="B944EA8EFFD1FF855BE9C6F7FC2B3FDD" box="[968,1017,1934,1958]" italics="true" pageId="4" pageNumber="32">18.3</emphasis>
], pb 8.4 μm [
<emphasis id="B944EA8EFFD1FF855CB3C6F7FB113FDD" box="[1170,1219,1934,1958]" italics="true" pageId="4" pageNumber="32">19.3</emphasis>
], sb 8.1 μm [
<emphasis id="B944EA8EFFD1FF855D7BC6F7FA593FDD" box="[1370,1419,1934,1958]" italics="true" pageId="4" pageNumber="32">18.6</emphasis>
]; external claw, leg II bc 9.8 μm [
<emphasis id="B944EA8EFFD1FF855A2EC6CAFD923FB0" box="[527,576,1971,1995]" italics="true" pageId="4" pageNumber="32">22.5</emphasis>
], pb 14.7 μm [
<emphasis id="B944EA8EFFD1FF855AD2C6CAFCF63FB0" box="[755,804,1971,1995]" italics="true" pageId="4" pageNumber="32">33.7</emphasis>
], sb 13.3 μm [
<emphasis id="B944EA8EFFD1FF855BF2C6CAFBD63FB0" box="[979,1028,1971,1995]" italics="true" pageId="4" pageNumber="32">30.5</emphasis>
]; external claw, leg III bc 10.0 μm [
<emphasis id="B944EA8EFFD1FF855881C6AEFF033F94" box="[160,209,2007,2031]" italics="true" pageId="4" pageNumber="32">22.9</emphasis>
], pb 17.0 μm [
<emphasis id="B944EA8EFFD1FF855959C6AEFE7B3F94" box="[376,425,2007,2031]" italics="true" pageId="4" pageNumber="32">39.0</emphasis>
], sb 12.0 μm [
<emphasis id="B944EA8EFFD1FF855A6FC6AEFDAD3F94" box="[590,639,2007,2031]" italics="true" pageId="4" pageNumber="32">27.5</emphasis>
]. Orientation of other claws in
<typeStatus id="548B883EFFD1FF855BFEC6AEFB933F94" box="[991,1089,2007,2031]" pageId="4" pageNumber="32" type="holotype">holotype</typeStatus>
unsuitable for measurements.
</paragraph>
<paragraph id="8B8F369CFFD0FF8458E4C1EEFBD6388C" blockId="5.[151,1436,151,391]" pageId="5" pageNumber="33">
Paratype—Body length 358.7 μm. Buccal tube 41.0 μm long and 4.4 μm [
<emphasis id="B944EA8EFFD0FF845C2AC1E1FBEE38CB" box="[1035,1084,152,176]" italics="true" pageId="5" pageNumber="33">10.7</emphasis>
] wide. Ventral lamina 21.8 μm [
<emphasis id="B944EA8EFFD0FF845881C1C5FF0338AF" box="[160,209,188,212]" italics="true" pageId="5" pageNumber="33">53.2</emphasis>
] long. Stylet supports inserted on buccal tube at 28.7 μm [70.0]. First macroplacoid 6.9 μm [
<emphasis id="B944EA8EFFD0FF845CCDC1C5FACF38AF" box="[1260,1309,188,212]" italics="true" pageId="5" pageNumber="33">16.8</emphasis>
] long, second macroplacoid 4.9 μm [
<emphasis id="B944EA8EFFD0FF8459E7C199FE253883" box="[454,503,224,248]" italics="true" pageId="5" pageNumber="33">12.0</emphasis>
] long. Macroplacoid row 12.8 μm [
<emphasis id="B944EA8EFFD0FF845BA8C199FC683883" box="[905,954,224,248]" italics="true" pageId="5" pageNumber="33">31.2</emphasis>
] long.
</paragraph>
<paragraph id="8B8F369CFFD0FF8458E4C07DFC1939FC" blockId="5.[151,1436,151,391]" pageId="5" pageNumber="33">
Claw lengths: Internal claw, leg II bc 6.8 μm [
<emphasis id="B944EA8EFFD0FF845AEDC07DFD2F3967" box="[716,765,260,284]" italics="true" pageId="5" pageNumber="33">16.6</emphasis>
], pb 11.0 μm [
<emphasis id="B944EA8EFFD0FF845B84C07DFC043967" box="[933,982,260,284]" italics="true" pageId="5" pageNumber="33">26.8</emphasis>
], sb 8.2 μm [
<emphasis id="B944EA8EFFD0FF845C4FC07DFB4D3967" box="[1134,1183,260,284]" italics="true" pageId="5" pageNumber="33">20.0</emphasis>
]; internal claw, leg III bc 6.4 μm [
<emphasis id="B944EA8EFFD0FF845937C051FE95393B" box="[278,327,296,320]" italics="true" pageId="5" pageNumber="33">15.6</emphasis>
], pb 11.8 μm [
<emphasis id="B944EA8EFFD0FF8459CAC051FDCE393B" box="[491,540,296,320]" italics="true" pageId="5" pageNumber="33">28.8</emphasis>
], sb 9.5 μm [
<emphasis id="B944EA8EFFD0FF845A91C051FD33393B" box="[688,737,296,320]" italics="true" pageId="5" pageNumber="33">23.2</emphasis>
]; external claw, leg IV bc 4.3 μm [
<emphasis id="B944EA8EFFD0FF845C44C051FB44393B" box="[1125,1174,296,320]" italics="true" pageId="5" pageNumber="33">10.5</emphasis>
], pb 12.3 μm [
<emphasis id="B944EA8EFFD0FF845D1AC051FABE393B" box="[1339,1388,296,320]" italics="true" pageId="5" pageNumber="33">30.0</emphasis>
], sb 7.2 μm [
<emphasis id="B944EA8EFFD0FF8458D8C035FEF8391F" box="[249,298,332,356]" italics="true" pageId="5" pageNumber="33">17.6</emphasis>
]; internal claw, leg IV bc 6.1 μm [
<emphasis id="B944EA8EFFD0FF845A99C035FD3B391F" box="[696,745,332,356]" italics="true" pageId="5" pageNumber="33">14.9</emphasis>
], pb 14.4 μm [
<emphasis id="B944EA8EFFD0FF845BB7C035FC15391F" box="[918,967,332,356]" italics="true" pageId="5" pageNumber="33">35.1</emphasis>
], sb 10.0 μm [
<emphasis id="B944EA8EFFD0FF845C50C035FB70391F" box="[1137,1186,332,356]" italics="true" pageId="5" pageNumber="33">24.4</emphasis>
]. Orientation of other claws unsuitable for measurements. Other characteristics as for
<typeStatus id="548B883EFFD0FF845B40C016FC1539FC" box="[865,967,367,391]" pageId="5" pageNumber="33" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF4F6614FFD0FF8458B6C704FC5D3EC9" httpUri="https://zenodo.org/record/205999/files/figure.png" pageId="5" pageNumber="33" targetBox="[168,1417,456,1627]" targetPageId="5">
<paragraph id="8B8F369CFFD0FF8458B6C704FC5D3EC9" blockId="5.[151,1436,1661,1715]" pageId="5" pageNumber="33">
<emphasis id="B944EA8EFFD0FF8458B6C704FE973EE8" bold="true" box="[151,325,1661,1684]" pageId="5" pageNumber="33">FIGURES 811.</emphasis>
<taxonomicName id="4C304D1FFFD0FF84596BC707FD9D3EEF" box="[330,591,1662,1684]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="33" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD0FF84596BC707FD9D3EEF" box="[330,591,1662,1684]" italics="true" pageId="5" pageNumber="33">Doryphoribius tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD0FF845A75C707FD5F3EE8" box="[596,653,1662,1683]" pageId="5" pageNumber="33" rank="species">
<emphasis id="B944EA8EFFD0FF845A75C707FD5F3EE8" bold="true" box="[596,653,1662,1683]" pageId="5" pageNumber="33">sp. n.</emphasis>
</taxonomicNameLabel>
(8) Buccopharyngeal apparatus. (9) Buccal cavity armature, dorsal view. (10) Claw, leg I. (11) Claw, leg IV. Scale bars: 8, 20 μm; 9, 5 μm; 1011, 10μm.
</paragraph>
</caption>
<subSubSection id="C32A6517FFD0FF8458E4C799FD5C3F67" pageId="5" pageNumber="33" type="discussion">
<paragraph id="8B8F369CFFD0FF8458E4C799FD5C3F67" blockId="5.[151,1436,1759,2036]" pageId="5" pageNumber="33">
<emphasis id="B944EA8EFFD0FF8458E4C799FEEB3E83" bold="true" box="[197,313,1760,1784]" pageId="5" pageNumber="33">Remarks.</emphasis>
No eggs were found. On the contracted juvenile specimen the anterior row of gibbosities was more prominent than in the
<typeStatus id="548B883EFFD0FF8459AFC67AFE223F60" box="[398,496,1795,1819]" pageId="5" pageNumber="33" type="holotype">holotype</typeStatus>
and
<typeStatus id="548B883EFFD0FF845A07C67DFD583F67" box="[550,650,1796,1820]" pageId="5" pageNumber="33" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFD0FF8458E4C651FD903F18" pageId="5" pageNumber="33" type="etymology">
<paragraph id="8B8F369CFFD0FF8458E4C651FD903F18" blockId="5.[151,1436,1759,2036]" pageId="5" pageNumber="33">
<emphasis id="B944EA8EFFD0FF8458E4C651FE993F3B" bold="true" box="[197,331,1832,1856]" pageId="5" pageNumber="33">Etymology.</emphasis>
The specific name
<taxonomicName id="4C304D1FFFD0FF845A08C65EFD4C3F44" box="[553,670,1831,1855]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="33" phylum="Tardigrada" rank="species" species="tessellatus">
<emphasis id="B944EA8EFFD0FF845A08C65EFD4C3F44" box="[553,670,1831,1855]" italics="true" pageId="5" pageNumber="33">tessellatus</emphasis>
</taxonomicName>
is a Latin masculine adjective referring to the mosaic pattern of the reticular design in the anterior cuticle.
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFD0FF8758E4C609FC2E3B27" lastPageId="6" lastPageNumber="34" pageId="5" pageNumber="33" type="materials_examined">
<paragraph id="8B8F369CFFD0FF8458E4C609FAFD3FD0" blockId="5.[151,1436,1759,2036]" pageId="5" pageNumber="33">
<emphasis id="B944EA8EFFD0FF8458E4C609FE5D3FF3" bold="true" box="[197,399,1903,1928]" pageId="5" pageNumber="33">
<typeStatus id="548B883EFFD0FF8458E4C609FED23FF3" box="[197,256,1904,1928]" pageId="5" pageNumber="33">Type</typeStatus>
depository.
</emphasis>
The
<typeStatus id="548B883EFFD0FF8459F2C616FDE73FFC" box="[467,565,1903,1927]" pageId="5" pageNumber="33" type="holotype">holotype</typeStatus>
and
<typeStatus id="548B883EFFD0FF845A54C609FD073FF3" box="[629,725,1904,1928]" pageId="5" pageNumber="33" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED21AE59FFD0FF845AC8C616FCEB3FFC" box="[745,825,1903,1927]" pageId="5" pageNumber="33">SMLA</collectionCode>
9400) are deposited in the W.A.K. Seale Museum, Department of Biology and Health Science, McNeese State University in Lake Charles, Louisiana,
<collectingCountry id="F327760CFFD0FF845CD1C6EAFAFE3FD0" box="[1264,1324,1939,1963]" name="United States of America" pageId="5" pageNumber="33">USA</collectingCountry>
.
</paragraph>
<paragraph id="8B8F369CFFD0FF8758E4C6CEFCB43944" blockId="5.[151,1436,1759,2036]" lastBlockId="6.[151,1436,151,861]" lastPageId="6" lastPageNumber="34" pageId="5" pageNumber="33">
<emphasis id="B944EA8EFFD0FF8458E4C6CEFE193FAB" bold="true" box="[197,459,1975,2000]" pageId="5" pageNumber="33">Differential diagnosis.</emphasis>
<bibRefCitation id="EFA14B6DFFD0FF8459F3C6CEFC983FB4" author="Michalczyk" box="[466,842,1975,1999]" pageId="5" pageNumber="33" refString="Michalczyk, L. &amp; Kaczmarek, L. (2010) Description of Doryphoribius dawkinsi, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from the Costa Rican highlands, with the key to the genus Doryphoribius. Zootaxa, 2393, 46 - 58." type="journal article" year="2010">Michalczyk &amp; Kaczmarek (2010)</bibRefCitation>
divide the species of
<taxonomicName id="4C304D1FFFD0FF845C1DC6CEFB333FB4" box="[1084,1249,1975,1999]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="33" phylum="Tardigrada" rank="genus">
<emphasis id="B944EA8EFFD0FF845C1DC6CEFB333FB4" box="[1084,1249,1975,1999]" italics="true" pageId="5" pageNumber="33">Doryphoribius</emphasis>
</taxonomicName>
into four groups based on the number of macroplacoids and the presence or absence of cuticular gibbosities.
<taxonomicName id="4C304D1FFFD0FF875CB2C6A5FF6A38CB" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" lastPageId="6" lastPageNumber="34" order="Parachela" pageId="5" pageNumber="33" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD0FF875CB2C6A5FF6A38CB" italics="true" lastPageId="6" lastPageNumber="34" pageId="5" pageNumber="33">Doryphoribius tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD3FF8758E1C1E1FED038CB" box="[192,258,152,176]" pageId="6" pageNumber="34" rank="species">
<emphasis id="B944EA8EFFD3FF8758E1C1E1FED038CB" bold="true" box="[192,258,152,176]" pageId="6" pageNumber="34">sp. n.</emphasis>
</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C304D1FFFD3FF875995C1E1FDC338CB" box="[436,529,152,176]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="evelinae">
<emphasis id="B944EA8EFFD3FF875995C1E1FDC338CB" box="[436,529,152,176]" italics="true" pageId="6" pageNumber="34">evelinae</emphasis>
</taxonomicName>
-group, characterised by two macroplacoids and a cuticle with gibbosities. This group includes nine species:
<taxonomicName id="4C304D1FFFD3FF8759FFC1C4FC0D38AF" authority="Michalczyk &amp; Kaczmarek, 2010" authorityName="Michalczyk &amp; Kaczmarek" authorityYear="2010" box="[478,991,188,212]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="dawkinsi">
<emphasis id="B944EA8EFFD3FF8759FFC1C4FDB738AF" box="[478,613,188,212]" italics="true" pageId="6" pageNumber="34">D. dawkinsi</emphasis>
<bibRefCitation id="EFA14B6DFFD3FF875A4DC1C5FC0D38AF" author="Michalczyk" box="[620,991,188,212]" pageId="6" pageNumber="34" refString="Michalczyk, L. &amp; Kaczmarek, L. (2010) Description of Doryphoribius dawkinsi, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from the Costa Rican highlands, with the key to the genus Doryphoribius. Zootaxa, 2393, 46 - 58." type="journal article" year="2010">Michalczyk &amp; Kaczmarek, 2010</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF875BCAC1C4FAFD38AF" authority="Ito, 1995" authorityName="Ito" authorityYear="1995" box="[1003,1327,188,212]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="dupliglobulatus">
<emphasis id="B944EA8EFFD3FF875BCAC1C4FB6C38AF" box="[1003,1214,188,212]" italics="true" pageId="6" pageNumber="34">D. dupliglobulatus</emphasis>
<bibRefCitation id="EFA14B6DFFD3FF875CE7C1C4FAFD38AF" author="Ito" box="[1222,1327,188,212]" pageId="6" pageNumber="34" refString="Ito, M. (1995) Taxonomic study of the Eutardigrada from the northern slope of Mt. Fuji, central Japan. III. Families Macrobiotidae and Milnesiidae. Proceedings of the Japanese Society of Systematic Zoology, 53, 18 - 39." type="journal article" year="1995">Ito, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF875D1AC1C4FEA9388C" authority="Marcus, 1928" authorityName="Marcus" authorityYear="1928" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="evelinae">
<emphasis id="B944EA8EFFD3FF875D1AC1C4FF123883" italics="true" pageId="6" pageNumber="34">D. evelinae</emphasis>
(
<bibRefCitation id="EFA14B6DFFD3FF8758F0C1A6FEA0388C" author="Marcus" box="[209,370,223,247]" pageId="6" pageNumber="34" refString="Marcus, E. (1928) Spinnentiere oder Arachoides. IV. Bartirchen (Tardigrada). In: Dahl, F. (Ed.), Die Tierreich Deutschlands und der angrenzenden Meeresteile Jena. Gustav Fischer, Jena, pp. 1 - 230." type="book chapter" year="1928">Marcus, 1928</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF8759A8C198FD4D388C" authority="Iharos, 1966" authorityName="Iharos" authorityYear="1966" box="[393,671,223,248]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="flavus">
<emphasis id="B944EA8EFFD3FF8759A8C198FE233883" box="[393,497,224,248]" italics="true" pageId="6" pageNumber="34">D. flavus</emphasis>
(
<bibRefCitation id="EFA14B6DFFD3FF875A22C1A6FD4A388C" author="Iharos" box="[515,664,223,247]" pageId="6" pageNumber="34" refString="Iharos, G. (1966) Neue Tardigraden-arten aus Ungarn: Neuere Beitrage zur Kenntnis der Tardigraden-Fauna Ungarns, VI. Acta Zoologica Academiae Scientarum Hungaricae, 12, 111 - 122." type="journal article" year="1966">Iharos, 1966</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF875A8CC198FB19388C" authority="Wang, Wang &amp; Li, 2007" authorityName="Wang, Wang &amp; Li" authorityYear="2007" box="[685,1227,223,248]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="huangguoshuensis">
<emphasis id="B944EA8EFFD3FF875A8CC198FC733883" box="[685,929,224,248]" italics="true" pageId="6" pageNumber="34">D. huangguoshuensis</emphasis>
<bibRefCitation id="EFA14B6DFFD3FF875B8DC199FB19388C" author="Wang" box="[940,1227,223,248]" pageId="6" pageNumber="34" refString="Wang, H., Wang, L. &amp; Li, X. (2007) Tardigrades from Guizhou Province (China) with description of a new species of genus Doryphoribius (Tardigrada: Hypsibiidae). Zootaxa, 1626, 59 - 67." type="journal article" year="2007">Wang, Wang &amp; Li, 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF875CFBC198FEAD3967" authority="Binda &amp; Pilato 1995" authorityName="Binda &amp; Pilato" authorityYear="1995" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="maranguensis">
<emphasis id="B944EA8EFFD3FF875CFBC198FA4E3883" box="[1242,1436,225,248]" italics="true" pageId="6" pageNumber="34">D. maranguensis</emphasis>
<bibRefCitation id="EFA14B6DFFD3FF8758B6C07DFEAD3967" author="Binda" box="[151,383,260,284]" pageId="6" pageNumber="34" refString="Binda, M. G. &amp; Pilato, G. (1995) Some notes on African tardigrades with description of two new species. Tropical Zoology, 8, 367 - 372." type="journal article" year="1995">Binda &amp; Pilato 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF8759ABC07CFCB73967" authority="Fontoura, Pilato &amp; Lisi, 2008" authorityName="Fontoura, Pilato &amp; Lisi" authorityYear="2008" box="[394,869,260,284]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="picoensis">
<emphasis id="B944EA8EFFD3FF8759ABC07CFDC63967" box="[394,532,261,284]" italics="true" pageId="6" pageNumber="34">D. picoensis</emphasis>
<bibRefCitation id="EFA14B6DFFD3FF875A3AC07CFCB73967" author="Fontoura" box="[539,869,260,284]" pageId="6" pageNumber="34" refString="Fontoura, P., Pilato, G. &amp; Lisi, O. (2008) New records of eutardigrades (Tardigrada) from Faial and Pico Islands, the Azores, with the description of two new species. Zootaxa, 1778, 37 - 47." type="journal article" year="2008">Fontoura, Pilato &amp; Lisi, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF875B51C07CFF003944" authority="Kaczmarek &amp; Michalczyk, 2004" authorityName="Kaczmarek &amp; Michalczyk" authorityYear="2004" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF875B51C07CFBB53967" box="[880,1127,260,284]" italics="true" pageId="6" pageNumber="34">D. quadrituberculatus</emphasis>
<bibRefCitation id="EFA14B6DFFD3FF875C4FC07DFF003944" author="Kaczmarek" pageId="6" pageNumber="34" refString="Kaczmarek, L. &amp; Michalczyk, L. (2004) First record of the genus Doryphoribius Pilato, 1969 from Costa Rica (Central America) and description of a new species Doryphoribius quadrituberculatus (Tardigrada: Hypsibiidae). Genus, 15, 447 - 453." type="journal article" year="2004">Kaczmarek &amp; Michalczyk, 2004</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C304D1FFFD3FF87592DC050FCB03944" authority="Horning, Schuster &amp; Grigarick, 1978" authorityName="Horning, Schuster &amp; Grigarick" authorityYear="1978" box="[268,866,295,320]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="zyxiglobus">
<emphasis id="B944EA8EFFD3FF87592DC050FE76393B" box="[268,420,296,320]" italics="true" pageId="6" pageNumber="34">D. zyxiglobus</emphasis>
(
<bibRefCitation id="EFA14B6DFFD3FF875992C05EFC883944" author="Horning" box="[435,858,295,319]" pageId="6" pageNumber="34" refString="Horning, D. S., Schuster, R. O. &amp; Grigarick, A. A. (1978) Tardigrada of New Zealand. New Zealand Journal of Zoology, 5, 185 - 280." type="journal article" year="1978">Horning, Schuster &amp; Grigarick, 1978</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8F369CFFD3FF8758E4C035FDA939B4" blockId="6.[151,1436,151,861]" pageId="6" pageNumber="34">
<taxonomicName id="4C304D1FFFD3FF8758E4C035FE34391F" box="[197,486,332,356]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD3FF8758E4C035FE34391F" box="[197,486,332,356]" italics="true" pageId="6" pageNumber="34">Doryphoribius tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD3FF8759CDC035FDFE391F" box="[492,556,332,356]" pageId="6" pageNumber="34" rank="species">
<emphasis id="B944EA8EFFD3FF8759CDC035FDFE391F" bold="true" box="[492,556,332,356]" pageId="6" pageNumber="34">sp. n.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C304D1FFFD3FF875AE1C034FCED391F" box="[704,831,332,356]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="evelinae">
<emphasis id="B944EA8EFFD3FF875AE1C034FCED391F" box="[704,831,332,356]" italics="true" pageId="6" pageNumber="34">D. evelinae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D1FFFD3FF875B57C034FC09391F" box="[886,987,332,356]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="flavus">
<emphasis id="B944EA8EFFD3FF875B57C034FC09391F" box="[886,987,332,356]" italics="true" pageId="6" pageNumber="34">D. flavus</emphasis>
</taxonomicName>
in having dorsal gibbosities but lacking eyes and gibbosities on the legs; from
<taxonomicName id="4C304D1FFFD3FF875A69C008FD1F39F3" box="[584,717,368,392]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="dawkinsi">
<emphasis id="B944EA8EFFD3FF875A69C008FD1F39F3" box="[584,717,368,392]" italics="true" pageId="6" pageNumber="34">D. dawkinsi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF875AFDC008FC7D39F3" box="[732,943,368,392]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="dupliglobulatus">
<emphasis id="B944EA8EFFD3FF875AFDC008FC7D39F3" box="[732,943,368,392]" italics="true" pageId="6" pageNumber="34">D. dupliglobulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C304D1FFFD3FF875B9CC008FB7D39F3" box="[957,1199,368,392]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="huangguoshuensis">
<emphasis id="B944EA8EFFD3FF875B9CC008FB7D39F3" box="[957,1199,368,392]" italics="true" pageId="6" pageNumber="34">D. huangguoshuensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D1FFFD3FF875CC7C008FAAD39F3" box="[1254,1407,368,392]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="zyxiglobus">
<emphasis id="B944EA8EFFD3FF875CC7C008FAAD39F3" box="[1254,1407,368,392]" italics="true" pageId="6" pageNumber="34">D. zyxiglobus</emphasis>
</taxonomicName>
in having fewer rows of gibbosities, and from
<taxonomicName id="4C304D1FFFD3FF875AAAC0ECFC9939D7" box="[651,843,405,428]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="maranguensis">
<emphasis id="B944EA8EFFD3FF875AAAC0ECFC9939D7" box="[651,843,405,428]" italics="true" pageId="6" pageNumber="34">D. maranguensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D1FFFD3FF875BA5C0ECFBC239D7" box="[900,1040,405,428]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="picoensis">
<emphasis id="B944EA8EFFD3FF875BA5C0ECFBC239D7" box="[900,1040,405,428]" italics="true" pageId="6" pageNumber="34">D. picoensis</emphasis>
</taxonomicName>
in having four evident caudal gibbosities and a cuticle with a reticular mesh.
</paragraph>
<paragraph id="8B8F369CFFD3FF8758E4C0A4FC2E3B27" blockId="6.[151,1436,151,861]" pageId="6" pageNumber="34">
In comparison to most species in the genus,
<taxonomicName id="4C304D1FFFD3FF875A93C0A5FC00398F" box="[690,978,476,500]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD3FF875A93C0A5FC00398F" box="[690,978,476,500]" italics="true" pageId="6" pageNumber="34">Doryphoribius tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD3FF875BF8C0A5FBCA398F" box="[985,1048,476,500]" pageId="6" pageNumber="34" rank="species">
<emphasis id="B944EA8EFFD3FF875BF8C0A5FBCA398F" bold="true" box="[985,1048,476,500]" pageId="6" pageNumber="34">sp. n.</emphasis>
</taxonomicNameLabel>
has relatively large macroplacoids and placoid row and a more posterior stylet support insertion point. While the new species shares with
<taxonomicName id="4C304D1FFFD3FF875D33C378FA493A63" box="[1298,1435,513,536]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="picoensis">
<emphasis id="B944EA8EFFD3FF875D33C378FA493A63" box="[1298,1435,513,536]" italics="true" pageId="6" pageNumber="34">D. picoensis</emphasis>
</taxonomicName>
the character of having smaller Claw IV than Claws IIII, it most closely resembles
<taxonomicName id="4C304D1FFFD3FF875C7DC35CFA873A47" box="[1116,1365,548,572]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF875C7DC35CFA873A47" box="[1116,1365,548,572]" italics="true" pageId="6" pageNumber="34">D. quadrituberculatus</emphasis>
</taxonomicName>
. Both the new species and
<taxonomicName id="4C304D1FFFD3FF87595EC330FDA23A1B" box="[383,624,584,608]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF87595EC330FDA23A1B" box="[383,624,584,608]" italics="true" pageId="6" pageNumber="34">D quadrituberculatus</emphasis>
</taxonomicName>
have six large, dorsal teeth and similar caudal gibbosities. The two species can be distinguished by several independent characters: (1) the two species differ in gibbosity sequence (III:4- 2-
<quantity id="4CC89B79FFD3FF87588CC3F6FF083ADC" box="[173,218,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="34" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C304D1FFFD3FF8758C3C3E9FDD13AD3" box="[226,515,656,680]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD3FF8758C3C3E9FDD13AD3" box="[226,515,656,680]" italics="true" pageId="6" pageNumber="34">Doryphoribius tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD3FF875A2AC3E9FD993ADC" box="[523,587,656,680]" pageId="6" pageNumber="34" rank="species">
<emphasis id="B944EA8EFFD3FF875A2AC3E9FD963AD3" bold="true" box="[523,580,656,680]" pageId="6" pageNumber="34">sp. n</emphasis>
.
</taxonomicNameLabel>
and II:2-
<quantity id="4CC89B79FFD3FF875A95C3F6FD333ADC" box="[692,737,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="34" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C304D1FFFD3FF875AC8C3E8FC333AD3" box="[745,993,656,680]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF875AC8C3E8FC333AD3" box="[745,993,656,680]" italics="true" pageId="6" pageNumber="34">D. quadrituberculatus</emphasis>
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) (2) the anterior reticular design of the new species (
<figureCitation id="130B2A19FFD3FF87590EC3CDFEA13AB7" box="[303,371,692,716]" captionStart="FIGURES 4 7" captionStartId="4.[151,264,1194,1217]" captionTargetBox="[207,1375,193,1163]" captionTargetId="figure@4.[196,1392,182,1173]" captionTargetPageId="4" captionText="FIGURES 4 7. Doryphoribius tessellatus sp. n. (4) Anterior gibbosities. (5) Posterior gibbosities. (6) Reticular design, middorsal region. (7) Reticular design, anterior region of the body. Scale bars: 4 6, 10 μm; 7, 5 μm." httpUri="https://zenodo.org/record/205998/files/figure.png" pageId="6" pageNumber="34">Fig. 7</figureCitation>
) differs from
<taxonomicName id="4C304D1FFFD3FF875A32C3CCFCDE3AB7" box="[531,780,692,716]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF875A32C3CCFCDE3AB7" box="[531,780,692,716]" italics="true" pageId="6" pageNumber="34">D. quadrituberculatus</emphasis>
</taxonomicName>
in consisting of large depressions interspersed with much smaller depressions (
<figureCitation id="130B2A19FFD3FF8759A6C3AEFE083A94" box="[391,474,727,751]" captionStart="FIGURE 12" captionStartId="6.[151,250,1951,1974]" captionTargetBox="[230,1341,923,1915]" captionTargetId="figure@6.[212,1376,887,1946]" captionTargetPageId="6" captionText="FIGURE 12. Doryphoribius quadrituberculatus. Reticular design of the anterior portion of the body. Scale bar: 10 μm." httpUri="https://zenodo.org/record/206000/files/figure.png" pageId="6" pageNumber="34">Fig. 12</figureCitation>
) (3) the new species lacks five ventral teeth found in
<taxonomicName id="4C304D1FFFD3FF875C6DC3A0FA953A8B" box="[1100,1351,728,752]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF875C6DC3A0FA953A8B" box="[1100,1351,728,752]" italics="true" pageId="6" pageNumber="34">D. quadrituberculatus</emphasis>
</taxonomicName>
(4) the macroplacoids of
<taxonomicName id="4C304D1FFFD3FF875943C384FE283B6F" box="[354,506,764,788]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="tessellatus" status="sp. nov.">
<emphasis id="B944EA8EFFD3FF875943C384FE283B6F" box="[354,506,764,788]" italics="true" pageId="6" pageNumber="34">D. tessellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD3FF875A23C385FD933B6F" box="[514,577,764,788]" pageId="6" pageNumber="34" rank="species">
<emphasis id="B944EA8EFFD3FF875A23C385FDE83B6F" bold="true" box="[514,570,764,788]" pageId="6" pageNumber="34">sp. n</emphasis>
.
</taxonomicNameLabel>
, especially the second, are longer than those of
<taxonomicName id="4C304D1FFFD3FF875C45C384FA8C3B6F" box="[1124,1374,764,788]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF875C45C384FA8C3B6F" box="[1124,1374,764,788]" italics="true" pageId="6" pageNumber="34">D. quadrituberculatus</emphasis>
</taxonomicName>
(second macroplacoid
<emphasis id="B944EA8EFFD3FF875948C258FEAD3B43" box="[361,383,801,824]" italics="true" pageId="6" pageNumber="34">pt</emphasis>
<quantity id="4CC89B79FFD3FF8759A6C266FE073B4C" box="[391,469,799,823]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" pageId="6" pageNumber="34" unit="in" value="11.5">11.5 in</quantity>
the new species and
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<taxonomicName id="4C304D1FFFD3FF875B2EC258FBDA3B43" box="[783,1032,800,824]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF875B2EC258FBDA3B43" box="[783,1032,800,824]" italics="true" pageId="6" pageNumber="34">D. quadrituberculatus</emphasis>
</taxonomicName>
in animals of comparable size) and (5) large eyes are present in
<taxonomicName id="4C304D1FFFD3FF8759F5C23CFD193B27" box="[468,715,836,860]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="34" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B944EA8EFFD3FF8759F5C23CFD193B27" box="[468,715,836,860]" italics="true" pageId="6" pageNumber="34">D. quadrituberculatus</emphasis>
</taxonomicName>
but not in the new species.
</paragraph>
</subSubSection>
</treatment>
</document>