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<document id="9C29F4D0B0E16A9DF4990B9AF99CB138" ID-DOI="10.5281/zenodo.182366" ID-GBIF-Dataset="1d12401f-e456-4790-86dd-85dc3b935a6a" ID-ISSN="1175-5326" ID-Zenodo-Dep="182366" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459860504177" checkinUser="plazi" docAuthor="Liu, Xingyue, Hayashi, Fumio &amp; Yang, Ding" docDate="2008" docId="03992C27AE41E71FADA0FF62FE47F85B" docLanguage="en" docName="zt01782p060.pdf" docOrigin="Zootaxa 1782" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Protohermes sabahensis Liu, Hayashi &amp; Yang, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="FFA0545FAE43E71DAD37FFF5FC67FFB9" masterDocTitle="Species of the Protohermes sabahensis group (Megaloptera: Corydalidae) from eastern Malaysia" masterLastPageNumber="60" masterPageNumber="49" pageNumber="51" updateTime="1698225994607" updateUser="plazi">
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<mods:title id="A0A66CAA4158522AE8A3127FAD00A8A2">Species of the Protohermes sabahensis group (Megaloptera: Corydalidae) from eastern Malaysia</mods:title>
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<mods:namePart id="B12B6FB08F7F3132EDE0D563069EF335">Liu, Xingyue</mods:namePart>
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<mods:namePart id="35B1C84DE4DD295318221BBE426F8415">Hayashi, Fumio</mods:namePart>
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<mods:namePart id="D4ADE725FA984A5D08D97DC35A61FC76">Yang, Ding</mods:namePart>
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<mods:title id="12E1C6C9256B128470E24E872D760AA3">Zootaxa</mods:title>
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<mods:part id="53125B467156C857087AA2496DB89A67">
<mods:date id="BE35A8CDEDF63E0A408DF3880AA95AF9">2008</mods:date>
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<mods:number id="DA72CDCBC34CDBC3AF3A4D37DE0B0B7C">1782</mods:number>
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<treatment id="03992C27AE41E71FADA0FF62FE47F85B" ID-DOI="http://doi.org/10.5281/zenodo.6234897" ID-GBIF-Taxon="119368124" ID-Zenodo-Dep="6234897" LSID="urn:lsid:plazi:treatment:03992C27AE41E71FADA0FF62FE47F85B" httpUri="http://treatment.plazi.org/id/03992C27AE41E71FADA0FF62FE47F85B" lastPageNumber="51" pageId="2" pageNumber="51">
<subSubSection id="C32ACEBAAE41E71FADA0FF62FDEBFF6D" pageId="2" pageNumber="51" type="nomenclature">
<paragraph id="8B8F9D31AE41E71FADA0FF62FE46FF08" blockId="2.[151,545,151,212]" box="[151,545,151,177]" pageId="2" pageNumber="51">
<heading id="D0C72A5DAE41E71FADA0FF62FE46FF08" bold="true" box="[151,545,151,177]" fontSize="11" level="1" pageId="2" pageNumber="51" reason="1">
<emphasis id="B9444123AE41E71FADA0FF62FE46FF08" bold="true" box="[151,545,151,177]" pageId="2" pageNumber="51">
<taxonomicName id="4C30E6B2AE41E71FADA0FF62FDDAFF08" ID-CoL="4N5JF" box="[151,445,151,177]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
<emphasis id="B9444123AE41E71FADA0FF62FDDAFF08" bold="true" box="[151,445,151,177]" italics="true" pageId="2" pageNumber="51">Protohermes sabahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277FC58AE41E71FACF3FF62FE46FF08" box="[452,545,151,177]" pageId="2" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F9D31AE41E71FADA0FF4FFDEBFF6D" blockId="2.[151,545,151,212]" box="[151,396,186,212]" pageId="2" pageNumber="51">
(
<figureCitation id="130B81B4AE41E71FADA8FF4FFD77FF6D" box="[159,272,186,212]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1954,1978]" captionTargetBox="[242,1350,195,1929]" captionTargetId="figure@3.[242,1350,183,1929]" captionTargetPageId="3" captionText="FIGURES 1 3. Adults of Protohermes spp. 1. P. sabahensis sp. nov. (holotype), male; 2. P. sabahensis sp. nov. (paratype), female; 3. P. spectabilis sp. nov. (holotype), male. Scale lines: 5.0 mm." httpUri="https://zenodo.org/record/182367/files/figure.png" pageId="2" pageNumber="51">Figs. 12</figureCitation>
,
<figureCitation id="130B81B4AE41E71FAC2BFF4FFD3EFF6D" box="[284,345,186,212]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">714</figureCitation>
,
<figureCitation id="130B81B4AE41E71FAC52FF4FFDE3FF6D" box="[357,388,186,212]" captionStart="FIGURE 32" captionStartId="9.[151,255,1735,1759]" captionTargetBox="[452,1134,1114,1734]" captionTargetId="figure@9.[452,1136,1114,1761]" captionTargetPageId="9" captionText="FIGURE 32. Distribution map of the P. sabahensis species group. P. sabahensis sp. nov. (); P. spectabilis sp. nov. (); P. tortuosus sp. nov. ()." httpUri="https://zenodo.org/record/182372/files/figure.png" pageId="2" pageNumber="51">32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32ACEBAAE41E71FADA0FEFFF931FA0B" pageId="2" pageNumber="51" type="diagnosis">
<paragraph id="8B8F9D31AE41E71FADA0FEFFFF3BFE25" blockId="2.[151,1437,266,2018]" pageId="2" pageNumber="51">
<emphasis id="B9444123AE41E71FADA0FEFFFD70FE9D" bold="true" box="[151,279,266,292]" pageId="2" pageNumber="51">Diagnosis.</emphasis>
This species is characterized by the yellow head and pronotum possessing three pairs of narrow black markings. Additionally, the coloration of the longitudinal veins alternates black and yellow in the middle of the forewings. The male ninth sternum is widely incised and Vshaped posteriorly, and the distal portion of the male tenth tergum is strongly broadened laterally.
</paragraph>
<paragraph id="8B8F9D31AE41E71FADF1FE5FF94FFE7D" blockId="2.[151,1437,266,2018]" box="[198,1320,426,452]" pageId="2" pageNumber="51">
Male. Body length
<quantity id="4CC830D4AE41E71FAC9FFE5FFE40FE7D" box="[424,551,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="2" pageNumber="51" unit="mm" value="19.5" valueMax="23.0" valueMin="16.0">1623 mm</quantity>
(n=25); forewing length
<quantity id="4CC830D4AE41E71FAE78FE5FFFB7FE7D" box="[847,976,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="2" pageNumber="51" unit="mm" value="30.5" valueMax="33.0" valueMin="28.0">2833 mm</quantity>
, hindwing length
<quantity id="4CC830D4AE41E71FA995FE5FF942FE7D" box="[1186,1317,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="2" pageNumber="51" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">2630 mm</quantity>
.
</paragraph>
<paragraph id="8B8F9D31AE41E71FADF2FE27F948FDDD" blockId="2.[151,1437,266,2018]" pageId="2" pageNumber="51">
Head (
<figureCitation id="130B81B4AE41E71FAC23FE27FD3DFE55" box="[276,346,466,492]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">Fig. 7</figureCitation>
) yellow, vertex with pair of subquadrate or subtriangular black markings near posterolateral margins; postocular spine almost absent. Compound eyes brown to slightly pale green; ocelli yellow, medially margined black, posterior ocelli touching median ocellus. Antenna black with scape and pedicel yellow. Mouthparts yellow; mandibles with distal half black; maxillary and labial palpi somewhat darkened.
</paragraph>
<paragraph id="8B8F9D31AE41E71FADF2FD87FEF8FC4E" blockId="2.[151,1437,266,2018]" pageId="2" pageNumber="51">
Thorax yellow; pronotum (
<figureCitation id="130B81B4AE41E71FAF31FD87FE2AFD35" box="[518,589,626,652]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">Fig. 7</figureCitation>
) with two pairs of elongate black markings near lateral margins, posterior pair widely separated. Thoracic pilosity yellowish, much longer on meso and metathorax than that on prothorax. Legs yellow, with short, dense, yellowish setae; tarsal claws pale reddish brown. Wings (
<figureCitation id="130B81B4AE41E71FA87CFD37F9F6FD65" box="[1355,1425,706,732]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1954,1978]" captionTargetBox="[242,1350,195,1929]" captionTargetId="figure@3.[242,1350,183,1929]" captionTargetPageId="3" captionText="FIGURES 1 3. Adults of Protohermes spp. 1. P. sabahensis sp. nov. (holotype), male; 2. P. sabahensis sp. nov. (paratype), female; 3. P. spectabilis sp. nov. (holotype), male. Scale lines: 5.0 mm." httpUri="https://zenodo.org/record/182367/files/figure.png" pageId="2" pageNumber="51">Fig. 1</figureCitation>
) almost hyaline, with several whitish markings. Forewings slightly brownish, with several irregular markings on proximal half and a round marking at distal 1/3; markings on proximal half usually expanded and fused with each other, making proximal half of forewing mostly white. Hindwings mostly clear, except for anterior and distal veins, which are light brown. Veins in forewing mostly brownish, except for those in whitish markings, which are colorless, and in some proximal longitudinal veins that alternate black and yellow. Rs 9 to 10branched, last branch bifurcate; both wings with 710 crossveins between R1 and Rs; M1+2 mostly 6 branched, M3+4 2branched; 1A 3branched.
</paragraph>
<paragraph id="8B8F9D31AE41E71FADF2FBFDFEB2FAAB" blockId="2.[151,1437,266,2018]" pageId="2" pageNumber="51">
Abdomen yellow. Ninth tergum (
<figureCitation id="130B81B4AE41E71FAF7DFBFDFEE8FB9B" box="[586,655,1032,1058]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">Fig. 8</figureCitation>
) subtrapezoidal, with anterior margin nearly straight and posterior margin Vshaped. Ninth sternum (
<figureCitation id="130B81B4AE41E71FAF1CFBC5FE17FBF3" box="[555,624,1072,1098]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">Fig. 9</figureCitation>
) broad, slightly longer than ninth tergum; posterior margin with wide Vshaped incision, forming pair of broad, triangular posterior processes. Ninth gonostylus (
<figureCitation id="130B81B4AE41E71FA9FBFBADF978FBCB" box="[1228,1311,1112,1138]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">Fig. 11</figureCitation>
) slenderly unguiform, nearly as long as ninth tergum, with tip slightly curved. Tenth tergum (
<figureCitation id="130B81B4AE41E71FA969FB75F8B2FB23" box="[1118,1237,1152,1178]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">Figs 810</figureCitation>
) flattened, about 1.5 times as long as ninth tergum, twisted; distal half strongly broadened in lateral view, with inner portion bearing long hairlike setae. Tenth sternum (
<figureCitation id="130B81B4AE41E71FAF98FB25FF61FB53" box="[687,774,1232,1258]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">Fig. 12</figureCitation>
) arched with dorsomedial process expanded as a plate; lateral lobes digitiform, slightly curved medially.
</paragraph>
<paragraph id="8B8F9D31AE41E71FADF2FAD5F931FA0B" blockId="2.[151,1437,266,2018]" pageId="2" pageNumber="51">
Female. Body length
<quantity id="4CC830D4AE41E71FACF2FAD5FE22FA83" box="[453,581,1312,1338]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="3.0" metricValueMin="2.3" pageId="2" pageNumber="51" unit="mm" value="26.5" valueMax="30.0" valueMin="23.0">2330 mm</quantity>
(n=13); forewing length
<quantity id="4CC830D4AE41E71FAE46FAD5FF93FA83" box="[881,1012,1312,1338]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="2" pageNumber="51" unit="mm" value="39.0" valueMax="41.0" valueMin="37.0">3741 mm</quantity>
, hindwing length
<quantity id="4CC830D4AE41E71FA9FEFAD5F92BFA83" box="[1225,1356,1312,1338]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.8" metricValueMin="3.4" pageId="2" pageNumber="51" unit="mm" value="36.0" valueMax="38.0" valueMin="34.0">3438 mm</quantity>
. Color similar to male. Eighth sternum (
<figureCitation id="130B81B4AE41E71FAF20FABDFEC4FADB" box="[535,675,1352,1378]" captionStart="FIGURES 7 14" captionStartId="6.[151,269,1502,1526]" captionTargetBox="[174,1413,380,1453]" captionTargetId="figure@6.[151,1436,354,1478]" captionTargetPageId="6" captionText="FIGURES 7 14. Protohermes sabahensis sp. nov. 7. Male head and prothorax, dorsal view; 8. Male genitalia, dorsal view; 9. Male genitalia, ventral view; 10. Male genitalia, lateral view; 11. Male ninth gonostylus, ventral view; 12. Male tenth sternum, ventral view; 13. Female genitalia, lateral view; 14. Female eighth sternum, ventral view. Scale lines: 1.0 mm. G 9: ninth gonocoxite; Pdm: dorsomedial process; S 8: eighth sternum; S 9: ninth sternum; St 9: ninth gonostylus; T 9: ninth tergum; T 10: tenth tergum." httpUri="https://zenodo.org/record/182369/files/figure.png" pageId="2" pageNumber="51">Figs. 1314</figureCitation>
) subtriangular in lateral view, posterior margin slightly produced without any incision. Ninth gonocoxite subquadrate, with small gonostylus at tip. Tenth tergum short, with posterior margin medially incised, leaving a thick digitiform dorsal and a thin semicircular ventral lobe.
</paragraph>
</subSubSection>
<subSubSection id="C32ACEBAAE41E71FADF2FA35F8ADF94B" pageId="2" pageNumber="51" type="materials_examined">
<paragraph id="8B8F9D31AE41E71FADF2FA35FCBDF91B" blockId="2.[151,1437,266,2018]" pageId="2" pageNumber="51">
<emphasis id="B9444123AE41E71FADF2FA35FDE0FA63" bold="true" box="[197,391,1472,1498]" pageId="2" pageNumber="51">
<typeStatus id="548B2393AE41E71FADF2FA35FD67FA63" box="[197,256,1472,1498]" pageId="2" pageNumber="51">Type</typeStatus>
materials.
</emphasis>
<typeStatus id="548B2393AE41E71FACA3FA35FE66FA63" box="[404,513,1472,1498]" pageId="2" pageNumber="51" type="holotype">Holotype</typeStatus>
ɗ, Eastern
<collectingCountry id="F327DDA1AE41E71FAFA4FA35FF64FA63" box="[659,771,1472,1498]" name="Malaysia" pageId="2" pageNumber="51">Malaysia</collectingCountry>
: Sabah, near Kinabalu, 6°8N, 116°55E, collected on 8/
<date id="FF8EBBF1AE41E71FADA0FA1DFD7AF9BB" box="[151,285,1512,1538]" pageId="2" pageNumber="51" value="1994-03-12">12.III.1994</date>
, reared from larva and emerged on
<date id="FF8EBBF1AE41E71FAF89FA1DFF25F9BB" box="[702,834,1512,1538]" pageId="2" pageNumber="51" value="1994-04-21">21.IV.1994</date>
, F. Hayashi (
<collectionCode id="ED2105F4AE41E71FAEECFA1DF878F9BB" LSID="urn:lsid:biocol.org:col:33074" box="[987,1055,1512,1538]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="2" pageNumber="51">CAU</collectionCode>
).
<typeStatus id="548B2393AE41E71FA904FA1DF8CFF9BB" box="[1075,1192,1512,1538]" pageId="2" pageNumber="51" type="paratype">Paratypes</typeStatus>
, Eastern
<collectingCountry id="F327DDA1AE41E71FA825FA1DF9E5F9BB" box="[1298,1410,1512,1538]" name="Malaysia" pageId="2" pageNumber="51">Malaysia</collectingCountry>
: 1 Ψ, same data as
<typeStatus id="548B2393AE41E71FAC6DF9E5FDA3F993" box="[346,452,1552,1578]" pageId="2" pageNumber="51" type="holotype">holotype</typeStatus>
, collected on 8/
<date id="FF8EBBF1AE41E71FAFB7F9E5FF61F993" box="[640,774,1552,1578]" pageId="2" pageNumber="51" value="1994-03-12">12.III.1994</date>
, reared from larva and emerged on
<date id="FF8EBBF1AE41E71FA998F9E5F945F993" box="[1199,1314,1552,1578]" pageId="2" pageNumber="51" value="1995-04-08">8.IV.1995</date>
(
<collectionCode id="ED2105F4AE41E71FA803F9E5F91FF993" LSID="urn:lsid:biocol.org:col:33074" box="[1332,1400,1552,1578]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="2" pageNumber="51">CAU</collectionCode>
); 1 ɗ, Sabah, near Kinabalu,
<date id="FF8EBBF1AE41E71FACF6F9CDFE5DF9EB" box="[449,570,1592,1618]" pageId="2" pageNumber="51" value="1992-06-06">6.VI.1992</date>
, H. Karube (HC); 1 ɗ, Sabah, near Kinabalu,
<date id="FF8EBBF1AE41E71FA966F9CDF8AEF9EB" box="[1105,1225,1592,1618]" pageId="2" pageNumber="51" value="1994-03-09">9.III.1994</date>
, F. Hayashi (HC); 1 ɗ, Sabah, near Kinabalu,
<date id="FF8EBBF1AE41E71FACDFF995FE13F9C3" box="[488,628,1632,1658]" pageId="2" pageNumber="51" value="1994-08-05">5.VIII.1994</date>
, S. Suda (HC); 1 Ψ, Sabah, near Kinabalu,
<date id="FF8EBBF1AE41E71FA9BEF995F977F9C3" box="[1161,1296,1632,1658]" pageId="2" pageNumber="51" value="1989-03-24">24.III.1989</date>
, H. Karube (HC).
</paragraph>
<paragraph id="8B8F9D31AE41E71FADF2F945F8ADF94B" blockId="2.[151,1437,266,2018]" pageId="2" pageNumber="51">
<emphasis id="B9444123AE41E71FADF2F945FE4CF973" bold="true" box="[197,555,1712,1738]" pageId="2" pageNumber="51">Other specimens examined.</emphasis>
Eastern
<collectingCountry id="F327DDA1AE41E71FAF97F945FF77F973" box="[672,784,1712,1738]" name="Malaysia" pageId="2" pageNumber="51">Malaysia</collectingCountry>
: 22 ɗ 11 Ψ, Sabah, near Kinabalu, collected on 8/
<date id="FF8EBBF1AE41E71FADA0F92DFD7AF94B" box="[151,285,1752,1778]" pageId="2" pageNumber="51" value="1994-03-12">12.III.1994</date>
, reared from larvae and emerged from
<date id="FF8EBBF1AE41E71FAFD1F92DFF39F94B" box="[742,862,1752,1778]" pageId="2" pageNumber="51" value="1994-05-18">18.V.1994</date>
to
<date id="FF8EBBF1AE41E71FAEB3F92DFF90F94B" box="[900,1015,1752,1778]" pageId="2" pageNumber="51" value="1996-01-28">28.I.1996</date>
, F. Hayashi (HC).
</paragraph>
</subSubSection>
<subSubSection id="C32ACEBAAE41E71FADF2F8F5FEC6F8A3" box="[197,673,1792,1818]" pageId="2" pageNumber="51" type="distribution">
<paragraph id="8B8F9D31AE41E71FADF2F8F5FEC6F8A3" blockId="2.[151,1437,266,2018]" box="[197,673,1792,1818]" pageId="2" pageNumber="51">
<emphasis id="B9444123AE41E71FADF2F8F5FD01F8A3" bold="true" box="[197,358,1792,1818]" pageId="2" pageNumber="51">Distribution.</emphasis>
Eastern
<collectingCountry id="F327DDA1AE41E71FACFBF8F5FE50F8A3" box="[460,567,1792,1818]" name="Malaysia" pageId="2" pageNumber="51">Malaysia</collectingCountry>
(Sabah).
</paragraph>
</subSubSection>
<subSubSection id="C32ACEBAAE41E71FADF2F8DDF888F8FB" box="[197,1263,1832,1858]" pageId="2" pageNumber="51" type="etymology">
<paragraph id="8B8F9D31AE41E71FADF2F8DDF888F8FB" blockId="2.[151,1437,266,2018]" box="[197,1263,1832,1858]" pageId="2" pageNumber="51">
<emphasis id="B9444123AE41E71FADF2F8DDFD35F8FB" bold="true" box="[197,338,1832,1858]" pageId="2" pageNumber="51">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C30E6B2AE41E71FAF65F8DCFEB4F8FB" box="[594,723,1833,1858]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="B9444123AE41E71FAF65F8DCFEB4F8FB" box="[594,723,1833,1858]" italics="true" pageId="2" pageNumber="51">sabahensis</emphasis>
</taxonomicName>
refers to the
<typeStatus id="548B2393AE41E71FAE40F8DDFFCDF8FB" box="[887,938,1832,1858]" pageId="2" pageNumber="51">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C32ACEBAAE41E71FADF2F8A5FE47F85B" pageId="2" pageNumber="51" type="discussion">
<paragraph id="8B8F9D31AE41E71FADF2F8A5FE47F85B" blockId="2.[151,1437,266,2018]" pageId="2" pageNumber="51">
<emphasis id="B9444123AE41E71FADF2F8A5FD59F8D3" bold="true" box="[197,318,1872,1898]" pageId="2" pageNumber="51">Remarks.</emphasis>
The new species appears to be closely related to
<taxonomicName id="4C30E6B2AE41E71FAE4EF8A4F865F8D3" box="[889,1026,1873,1898]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="tortuosus" status="sp. nov.">
<emphasis id="B9444123AE41E71FAE4EF8A4F865F8D3" box="[889,1026,1873,1898]" italics="true" pageId="2" pageNumber="51">P. tortuosus</emphasis>
</taxonomicName>
<emphasis id="B9444123AE41E71FA93EF8A5F801F8D3" bold="true" box="[1033,1126,1872,1898]" pageId="2" pageNumber="51">
<taxonomicNameLabel id="A277FC58AE41E71FA93EF8A5F801F8D3" box="[1033,1126,1872,1898]" pageId="2" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having a similar yellow body coloration and its male tenth tergum distally broadened, but it can be easily separated from
<taxonomicName id="4C30E6B2AE41E71FA826F88CF9FBF82B" box="[1297,1436,1913,1938]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="tortuosus">
<emphasis id="B9444123AE41E71FA826F88CF9FBF82B" box="[1297,1436,1913,1938]" italics="true" pageId="2" pageNumber="51">P. tortuosus</emphasis>
</taxonomicName>
by the male ninth sternum with a Vshaped posterior incision. The posterior incision of the male ninth sternum of
<taxonomicName id="4C30E6B2AE41E71FADC5F83DFD1BF858" box="[242,380,1992,2017]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="tortuosus">
<emphasis id="B9444123AE41E71FADC5F83DFD1BF858" box="[242,380,1992,2017]" italics="true" pageId="2" pageNumber="51">P. tortuosus</emphasis>
</taxonomicName>
is suboblong.
</paragraph>
</subSubSection>
</treatment>
</document>