162 lines
20 KiB
XML
162 lines
20 KiB
XML
<document id="750382A6F6F74630B1BC12AAA9D0AD64" ID-DOI="10.1111/j.1096-3642.2011.00738.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5442323" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630452876177" checkinUser="valdenar" docAuthor="Karaman, Ivo, Hammouti, Nasera, Pavićević, Dragan, Kiefer, Andreas, Horvatović, Mladen & Seitz, Alfred" docDate="2011" docId="03F75E0EFFD7FFAFFCE2F962FA82FBB5" docLanguage="en" docName="j.1096-3642.2011.00738.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00738.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Troglophilus brevicauda : US 1970" docType="treatment" docVersion="3" lastPageNumber="1048" masterDocId="FFCE2676FFDBFFA2FF98FFDBFFD3FFD9" masterDocTitle="The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans" masterLastPageNumber="1063" masterPageNumber="1035" pageNumber="1047" updateTime="1699275884152" updateUser="plazi">
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<mods:title id="D980E2EDFF94C03BAD748F2C8200A5D3">The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans</mods:title>
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<mods:namePart id="8FE6086ADAB88824CDD3C91DB1F06C45">Karaman, Ivo</mods:namePart>
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<mods:namePart id="771F499C6AE97C5F0F72BE2B075740FC">Hammouti, Nasera</mods:namePart>
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<mods:namePart id="AF2B5A69329FA6A1BF012C6056AB3345">Pavićević, Dragan</mods:namePart>
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<mods:namePart id="106DACCD9CF883B18C3A2FDA035C48D9">Kiefer, Andreas</mods:namePart>
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<mods:namePart id="01EBF6F972CD08114245D0897408E32E">Horvatović, Mladen</mods:namePart>
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<mods:namePart id="988ECE6B40BB41A7591AF4E787A991BF">Seitz, Alfred</mods:namePart>
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<mods:typeOfResource id="C2CB7877405DC7BCB8BEB01555796F47">text</mods:typeOfResource>
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<mods:title id="7F67D8062E401669DFE039B18BC7D155">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="32501EF8CB14AE1D318DC7D61600C64B">
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<mods:date id="F1BD2FA4EA44DAF2A7E4DCA00AFCFE7D">2011</mods:date>
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<mods:number id="28F20D31C5C3A00640542EA433B03850">2011-11-25</mods:number>
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<mods:detail id="BC01E7DC8A3A2056C9093CCDEB768E4B" type="volume">
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<mods:number id="FB2D29F5AD7C86DCA8D6C454FE8A33DE">163</mods:number>
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<mods:number id="C95195EB2DADA4925B25783410A28FF3">4</mods:number>
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<mods:start id="1CCF97B64D0B8286BBCFA14C8551FDD3">1035</mods:start>
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<mods:url id="F7AE4B12F4137CC134906F022314BE98">http://dx.doi.org/10.1111/j.1096-3642.2011.00738.x</mods:url>
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<mods:identifier id="D7CCE292C6FE1A51D627C429B3F5A05C" type="DOI">10.1111/j.1096-3642.2011.00738.x</mods:identifier>
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<mods:identifier id="E4065AB781C333A7AEBA0FF65C8F476B" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="F56C3F4C2A5642952BFA25500620BD10" type="Zenodo-Dep">5442323</mods:identifier>
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<treatment id="03F75E0EFFD7FFAFFCE2F962FA82FBB5" LSID="urn:lsid:plazi:treatment:03F75E0EFFD7FFAFFCE2F962FA82FBB5" httpUri="http://treatment.plazi.org/id/03F75E0EFFD7FFAFFCE2F962FA82FBB5" lastPageId="13" lastPageNumber="1048" pageId="12" pageNumber="1047">
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<subSubSection id="C344BC93FFD7FFAEFCE2F962FAEAF914" box="[890,1337,1720,1742]" pageId="12" pageNumber="1047" type="nomenclature">
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<paragraph id="8BE1EF18FFD7FFAEFCE2F962FAEAF914" blockId="12.[890,1337,1720,1742]" box="[890,1337,1720,1742]" pageId="12" pageNumber="1047">
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<heading id="D0A95874FFD7FFAEFCE2F962FAEAF914" box="[890,1337,1720,1742]" centered="true" fontSize="8" level="2" pageId="12" pageNumber="1047" reason="8">
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<taxonomicName id="4C5E949BFFD7FFAEFCE2F962FAEAF914" ID-CoL="5993T" authority=": US, 1970" authorityName=": US" authorityYear="1970" box="[890,1337,1720,1742]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="1047" phylum="Arthropoda" rank="species" species="brevicauda">
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<emphasis id="B92A330AFFD7FFAEFCE2F962FB6EF915" box="[890,1213,1721,1742]" italics="true" pageId="12" pageNumber="1047">TROGLOPHILUS BREVICAUDA</emphasis>
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:
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<bibRefCitation id="EFCF92E9FFD7FFAEFB56F962FAEAF914" author="Us P" box="[1230,1337,1720,1742]" pageId="12" pageNumber="1047" pagination="301 - 319" refId="ref11629" refString="Us P. 1970. Prispevek k poznavanju jamskih ortopterov Jugoslavije (Orthoptera - Tettigonoidea). Acta Carsologica SAZU 5: 301 - 319." type="journal article" year="1970">US, 1970</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C344BC93FFD7FFAFFCBEF904FA82FBB5" lastPageId="13" lastPageNumber="1048" pageId="12" pageNumber="1047" type="materials_examined">
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<paragraph id="8BE1EF18FFD7FFAEFCBEF904FB65F92C" blockId="12.[806,1206,1759,1781]" box="[806,1206,1759,1781]" pageId="12" pageNumber="1047">
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<emphasis id="B92A330AFFD7FFAEFCBEF904FBD5F92D" box="[806,1030,1759,1780]" italics="true" pageId="12" pageNumber="1047">Material examined:</emphasis>
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See Appendix.
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</paragraph>
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<paragraph id="8BE1EF18FFD7FFAFFCBEF8E0FED5FAA6" blockId="12.[806,1421,1850,1903]" lastBlockId="13.[164,779,1049,1407]" lastPageId="13" lastPageNumber="1048" pageId="12" pageNumber="1047">
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<emphasis id="B92A330AFFD7FFAEFCBEF8E0FC4EF889" box="[806,925,1851,1872]" italics="true" pageId="12" pageNumber="1047">Diagnosis:</emphasis>
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Medium to large sized species. Male body length ranges between
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<quantity id="4CA642FDFFD7FFAEFB05F882FAC4F8B7" box="[1181,1303,1881,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="12" pageNumber="1047" unit="mm" value="17.5" valueMax="21.0" valueMin="14.0">14–21 mm</quantity>
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; females, 14.5–
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<quantity id="4CA642FDFFD6FFAFFF79FBC2FEE0FBF7" box="[225,307,1049,1071]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="1048" unit="mm" value="22.0">22 mm</quantity>
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. Body colour dark brown to light brown marbled, rarely light testaceous. Male tenth tergite distal margin with wide, broadly rounded medial protrusion (
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<figureCitation id="1365F39DFFD6FFAFFE90FBAEFEB8FB52" box="[264,363,1141,1163]" captionStart="Figure 12" captionStartId="15.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-192@15.[323,1283,196,1286]" captionTargetPageId="15" captionText="Figure 12. A–C, Troglophilus cavicola: A, male 18 mm, tenth tergite, dorsal view (Bjeloševina, Montenegro); B, female 18.5 mm, tenth tergite, dorsal view (Bukovik, Macedonia); C, male 18 mm, phallus with titillator (arrow), lateral view (Bjeloševina, Montenegro). D–I ovipositor, lateral view: D, Troglophilus lazaropolensis, female 16.5 mm (Lazaropole, Macedonia); E, Troglophilus cavicola, female 18.5 mm (Bjeloševina, Montenegro); F, Troglophilus zorae sp. nov., paratype female 18 mm (Matka, Macedonia); G, Troglophilus brevicauda, female 15 mm (Olovo, Bosnia); H, Troglophilus ovuliformis, female 19.5 mm (D. Seoca, Montenegro); I, Troglophilus neglectus, female 15 mm (N. Breznica, Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5442347" httpUri="https://zenodo.org/record/5442347/files/figure.png" pageId="13" pageNumber="1048">Fig. 12A</figureCitation>
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), separated by narrow more or less deep incision in two lobes. Female tenth abdominal tergite with slightly shallow excavated margin (
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<figureCitation id="1365F39DFFD6FFAFFF34FB0BFEDCFB3E" box="[172,271,1232,1255]" captionStart="Figure 12" captionStartId="15.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-192@15.[323,1283,196,1286]" captionTargetPageId="15" captionText="Figure 12. A–C, Troglophilus cavicola: A, male 18 mm, tenth tergite, dorsal view (Bjeloševina, Montenegro); B, female 18.5 mm, tenth tergite, dorsal view (Bukovik, Macedonia); C, male 18 mm, phallus with titillator (arrow), lateral view (Bjeloševina, Montenegro). D–I ovipositor, lateral view: D, Troglophilus lazaropolensis, female 16.5 mm (Lazaropole, Macedonia); E, Troglophilus cavicola, female 18.5 mm (Bjeloševina, Montenegro); F, Troglophilus zorae sp. nov., paratype female 18 mm (Matka, Macedonia); G, Troglophilus brevicauda, female 15 mm (Olovo, Bosnia); H, Troglophilus ovuliformis, female 19.5 mm (D. Seoca, Montenegro); I, Troglophilus neglectus, female 15 mm (N. Breznica, Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5442347" httpUri="https://zenodo.org/record/5442347/files/figure.png" pageId="13" pageNumber="1048">Fig. 12B</figureCitation>
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). Titillator is well sclerotized, long and slender, arched, and acute at the apex (
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<figureCitation id="1365F39DFFD6FFAFFD03FB34FD2CFADC" box="[667,767,1263,1285]" captionStart="Figure 12" captionStartId="15.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-192@15.[323,1283,196,1286]" captionTargetPageId="15" captionText="Figure 12. A–C, Troglophilus cavicola: A, male 18 mm, tenth tergite, dorsal view (Bjeloševina, Montenegro); B, female 18.5 mm, tenth tergite, dorsal view (Bukovik, Macedonia); C, male 18 mm, phallus with titillator (arrow), lateral view (Bjeloševina, Montenegro). D–I ovipositor, lateral view: D, Troglophilus lazaropolensis, female 16.5 mm (Lazaropole, Macedonia); E, Troglophilus cavicola, female 18.5 mm (Bjeloševina, Montenegro); F, Troglophilus zorae sp. nov., paratype female 18 mm (Matka, Macedonia); G, Troglophilus brevicauda, female 15 mm (Olovo, Bosnia); H, Troglophilus ovuliformis, female 19.5 mm (D. Seoca, Montenegro); I, Troglophilus neglectus, female 15 mm (N. Breznica, Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5442347" httpUri="https://zenodo.org/record/5442347/files/figure.png" pageId="13" pageNumber="1048">Fig. 12C</figureCitation>
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). Ovipositor elongated and narrow, length ranges between 10.5–13.5 mm; the widest point is in its proximal third (
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<figureCitation id="1365F39DFFD6FFAFFEE9FA90FE00FAB8" box="[369,467,1355,1377]" captionStart="Figure 12" captionStartId="15.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-192@15.[323,1283,196,1286]" captionTargetPageId="15" captionText="Figure 12. A–C, Troglophilus cavicola: A, male 18 mm, tenth tergite, dorsal view (Bjeloševina, Montenegro); B, female 18.5 mm, tenth tergite, dorsal view (Bukovik, Macedonia); C, male 18 mm, phallus with titillator (arrow), lateral view (Bjeloševina, Montenegro). D–I ovipositor, lateral view: D, Troglophilus lazaropolensis, female 16.5 mm (Lazaropole, Macedonia); E, Troglophilus cavicola, female 18.5 mm (Bjeloševina, Montenegro); F, Troglophilus zorae sp. nov., paratype female 18 mm (Matka, Macedonia); G, Troglophilus brevicauda, female 15 mm (Olovo, Bosnia); H, Troglophilus ovuliformis, female 19.5 mm (D. Seoca, Montenegro); I, Troglophilus neglectus, female 15 mm (N. Breznica, Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5442347" httpUri="https://zenodo.org/record/5442347/files/figure.png" pageId="13" pageNumber="1048">Fig. 12E</figureCitation>
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), ventral margin almost straight.
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</paragraph>
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<caption id="DF21BF90FFD6FFAFFF3CFC6FFB2FFC3C" ID-DOI="http://doi.org/10.5281/zenodo.5442343" ID-Zenodo-Dep="5442343" httpUri="https://zenodo.org/record/5442343/files/figure.png" pageId="13" pageNumber="1048" startId="13.[164,243,948,967]" targetBox="[323,1283,196,919]" targetPageId="13">
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<paragraph id="8BE1EF18FFD6FFAFFF3CFC6FFB2FFC3C" blockId="13.[164,1441,948,997]" pageId="13" pageNumber="1048">
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<emphasis id="B92A330AFFD6FFAFFF3CFC6FFECAFC11" bold="true" box="[164,281,948,968]" pageId="13" pageNumber="1048">Figure 10.</emphasis>
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A–D,
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<taxonomicName id="4C5E949BFFD6FFAFFEFCFC6EFDADFC11" box="[356,638,949,968]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="1048" phylum="Arthropoda" rank="species" species="azaropolensis">
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<emphasis id="B92A330AFFD6FFAFFEFCFC6EFDADFC11" box="[356,638,949,968]" italics="true" pageId="13" pageNumber="1048">Troglophilus azaropolensis</emphasis>
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</taxonomicName>
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(Lazaropole, Macedonia): A–B, male 15 mm, tenth tergite, dorsal view; C, female 16.5 mm, tenth tergite, dorsal view; D, male 15 mm, phallus with titillator (arrow), dorsal view.
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</paragraph>
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</caption>
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<paragraph id="8BE1EF18FFD6FFAFFF3CFA72FEF5F96D" blockId="13.[164,779,1449,1716]" pageId="13" pageNumber="1048">
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<emphasis id="B92A330AFFD6FFAFFF3CFA72FEE6FA67" box="[164,309,1449,1470]" italics="true" pageId="13" pageNumber="1048">Distribution:</emphasis>
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This species was originally described from the small cave Schelmenloch, south of Vienna, and it is of Dinaric origin. It is distributed from Lombardy and Vienna in the north-west, across Dinarids and Šar-Pindus mountain system, to central Greece in the south-east (Oiti Mt. after
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<bibRefCitation id="EFCF92E9FFD6FFAFFD03F999FF08F9AE" author="Willemse F" pageId="13" pageNumber="1048" refId="ref11655" refString="Willemse F. 1984. Fauna Graeciae I. Catalogue of the Orthoptera of Greece. Athens: Hellenic Zoological Society." type="book" year="1984">Willemse, 1984</bibRefCitation>
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) (
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<figureCitation id="1365F39DFFD6FFAFFF6BF9BAFE90F9AE" box="[243,323,1633,1655]" captionStart="Figure 13" captionStartId="16.[144,223,1180,1199]" captionTargetBox="[144,1425,196,1151]" captionTargetId="figure-318@16.[144,1425,196,1151]" captionTargetPageId="16" captionText="Figure 13. The geographical distribution of Troglophilus cavicola (square – literature data). (ITA-Italy; A-Austria; SLO-Slovenia; CRO-Croatia; BIH-Bosnia and Herzegovina; SRB-Serbia; MNE-Montenegro; BUL-Bulgaria; ALB-Albania; MK-Macedonia; GRE-Greece)." figureDoi="http://doi.org/10.5281/zenodo.5442349" httpUri="https://zenodo.org/record/5442349/files/figure.png" pageId="13" pageNumber="1048">Fig. 13</figureCitation>
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). It is completely absent from southern and eastern
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<collectingCountry id="F349AF88FFD6FFAFFED3F95BFE44F94F" box="[331,407,1664,1686]" name="Serbia" pageId="13" pageNumber="1048">Serbia</collectingCountry>
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, from
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<collectingCountry id="F349AF88FFD6FFAFFE61F95BFD8CF94F" box="[505,607,1664,1686]" name="Bulgaria" pageId="13" pageNumber="1048">Bulgaria</collectingCountry>
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, and eastern
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<collectingCountry id="F349AF88FFD6FFAFFF3CF945FEF0F96D" box="[164,291,1694,1716]" name="Macedonia" pageId="13" pageNumber="1048">Macedonia</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BE1EF18FFD6FFAFFF3CF905FE8EF8B7" blockId="13.[164,779,1758,1902]" pageId="13" pageNumber="1048">
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<emphasis id="B92A330AFFD6FFAFFF3CF905FEDCF92A" box="[164,271,1758,1779]" italics="true" pageId="13" pageNumber="1048">Remarks:</emphasis>
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Across its distribution range, the species is sympatric with all other species of the genus
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<taxonomicName id="4C5E949BFFD6FFAFFD23F926FF3FF8E8" authorityName="Krauss" authorityYear="1879" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="1048" phylum="Arthropoda" rank="genus">
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<emphasis id="B92A330AFFD6FFAFFD23F926FF3FF8E8" italics="true" pageId="13" pageNumber="1048">Troglophilus</emphasis>
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</taxonomicName>
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. At some localities in western
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<collectingCountry id="F349AF88FFD6FFAFFD17F8C7FCD9F8EB" box="[655,778,1820,1842]" name="Macedonia" pageId="13" pageNumber="1048">Macedonia</collectingCountry>
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it occurs sympatrically with
|
||
<taxonomicName id="4C5E949BFFD6FFAFFD89F8E0FD1CF896" authorityName=": PAVICEVIC & KARAMAN" authorityYear="2001" box="[529,719,1850,1872]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="1048" phylum="Arthropoda" rank="species" species="lazaropolensis">
|
||
<emphasis id="B92A330AFFD6FFAFFD89F8E0FD1CF896" box="[529,719,1850,1872]" italics="true" pageId="13" pageNumber="1048">T. lazaropolensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5E949BFFD6FFAFFF3CF882FF2FF8B7" authority="Karaman & Hammouti & Pavićević & Kiefer & Horvatović & Seitz, 2011" authorityName="Karaman & Hammouti & Pavićević & Kiefer & Horvatović & Seitz" authorityYear="2011" box="[164,252,1881,1902]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="1048" phylum="Arthropoda" rank="species" species="zorae" status="sp. nov.">
|
||
<emphasis id="B92A330AFFD6FFAFFF3CF882FF2FF8B7" box="[164,252,1881,1902]" italics="true" pageId="13" pageNumber="1048">T. zorae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2198E71FFD6FFAFFE9DF882FE8EF8B7" box="[261,349,1881,1902]" pageId="13" pageNumber="1048" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BE1EF18FFD6FFAFFCCBFBC2FA82FBB5" blockId="13.[826,1441,1049,1132]" pageId="13" pageNumber="1048">
|
||
By examination of the
|
||
<typeStatus id="54E551BAFFD6FFAFFBE6FBC2FB0CFBF6" box="[1150,1247,1049,1071]" pageId="13" pageNumber="1048" type="holotype">holotype</typeStatus>
|
||
and specimens collected from the type locality, we concluded that
|
||
<taxonomicName id="4C5E949BFFD6FFAFFCA2FB8CFC6BFBB2" box="[826,952,1110,1132]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="1048" phylum="Arthropoda" rank="species" species="bukoviki">
|
||
<emphasis id="B92A330AFFD6FFAFFCA2FB8CFC6BFBB2" box="[826,952,1110,1132]" italics="true" pageId="13" pageNumber="1048">T. bukoviki</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="4C5E949BFFD6FFAFFB4AFB8CFA99FBB2" authorityName=": US" authorityYear="1970" box="[1234,1354,1110,1132]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="1048" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFD6FFAFFB4AFB8CFA99FBB2" box="[1234,1354,1110,1132]" italics="true" pageId="13" pageNumber="1048">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |