338 lines
47 KiB
XML
338 lines
47 KiB
XML
<document id="23B71FA7CE130464D8D9876855DB6133" ID="10.11646/zootaxa.4250.5.5SLASH10746" ID-CLB-Dataset="33274" ID-DOI="10.11646/zootaxa.4250.5.5" ID-GBIF-Dataset="3e3123a6-d497-4ef4-afa0-7cf0ef7f9afa" ID-ISSN="1175-5326" ID-Zenodo-Dep="495550" ID-ZooBank="5448198D-7CF2-47CE-A3CF-E7760EC5DDB6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491551433022" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Lemaitre, Rafael" docDate="2017" docId="03B88780FF91EC4360A1320DFA70B5D8" docLanguage="en" docName="zootaxa.4250.5.5SLASH10746.pdf" docOrigin="Zootaxa 4250 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geosesarma vicentense Rathbun 1914" docType="treatment" docVersion="7" lastPageNumber="476" masterDocId="FF81FFF8FF90EC426036332FFFF0B23C" masterDocTitle="On the taxonomy of two poorly known species of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) from the Philippines" masterLastPageNumber="483" masterPageNumber="475" pageNumber="476" updateTime="1698438789559" updateUser="ExternalLinkService">
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<mods:mods id="485E73BEE7BBE732064E660F514E8265" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="64A44D9648385872F110D3C51C47232E">
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<mods:title id="F98838EB806DA4EA10F12F66D3DFE43C">On the taxonomy of two poorly known species of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) from the Philippines</mods:title>
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<mods:name id="579AC7804868C6120600225604F3A2FF" type="personal">
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<mods:namePart id="8D7C58D22AB663A457A02AB4F77246C8">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="21E2BA15AAB1F5AAC1CE9945F5A53861">Lemaitre, Rafael</mods:namePart>
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<mods:title id="D4A141484271BF9F18CA9DD1248D0376">Zootaxa</mods:title>
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<mods:part id="70948621BE345E1C3A495652F60CAA06">
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<mods:date id="543FAC4F108FB5096153AE48122229CD">2017</mods:date>
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<mods:number id="B685767E4386AFF4DBD072943AA33214">4250</mods:number>
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<mods:detail id="41663F7EBE74295721CA0F103C2AF2AD" type="issue">
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<mods:number id="4A900C9255393D47A0E69F4A58A303C5">5</mods:number>
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<mods:identifier id="810EA388E74DBEEFC9ED9FD1B0AE14C4" type="CLB-Dataset">33274</mods:identifier>
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<mods:identifier id="6C8CFE797A653CB21A7B8CBB01D73441" type="DOI">10.11646/zootaxa.4250.5.5</mods:identifier>
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<mods:identifier id="194A9E577D9AAFBB635A5CC1474E80FC" type="GBIF-Dataset">3e3123a6-d497-4ef4-afa0-7cf0ef7f9afa</mods:identifier>
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<treatment id="03B88780FF91EC4360A1320DFA70B5D8" ID-DOI="http://doi.org/10.5281/zenodo.6028323" ID-GBIF-Taxon="128344778" ID-Zenodo-Dep="6028323" LSID="urn:lsid:plazi:treatment:03B88780FF91EC4360A1320DFA70B5D8" httpUri="http://treatment.plazi.org/id/03B88780FF91EC4360A1320DFA70B5D8" lastPageNumber="476" pageId="1" pageNumber="476">
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<subSubSection id="C30B651DFF91EC4360A1320DFEB8B362" pageId="1" pageNumber="476" type="nomenclature">
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<paragraph id="8BAE3696FF91EC4360A1320DFD72B300" blockId="1.[151,642,290,350]" box="[151,642,290,316]" pageId="1" pageNumber="476">
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<heading id="D0E681FAFF91EC4360A1320DFD72B300" bold="true" box="[151,642,290,316]" fontSize="11" level="1" pageId="1" pageNumber="476" reason="1">
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<taxonomicName id="4C114D15FF91EC4360A1320DFD72B300" ID-CoL="6KB9Q" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[151,642,290,316]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense">
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<emphasis id="B965EA84FF91EC4360A1320DFD72B300" bold="true" box="[151,642,290,316]" pageId="1" pageNumber="476">
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<emphasis id="B965EA84FF91EC4360A1320DFE5FB300" bold="true" box="[151,431,290,316]" italics="true" pageId="1" pageNumber="476">Geosesarma vicentense</emphasis>
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(
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<bibRefCitation id="EF804B67FF91EC436188320DFD8AB300" author="Rathbun" box="[446,634,290,316]" pageId="1" pageNumber="476" refString="Rathbun, M. J. (1914) New species of crabs of the families Grapsidae and Ocypodidae. Proceedings of the United States National Museum, 47, 69 - 85." type="journal article" year="1914">Rathbun, 1914</bibRefCitation>
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||
)
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||
</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BAE3696FF91EC4360A1326AFEB8B362" blockId="1.[151,642,290,350]" box="[151,328,325,350]" pageId="1" pageNumber="476">
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(
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<figureCitation id="132A2A13FF91EC4360A9326AFF1CB362" box="[159,236,325,350]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Figs. 1</figureCitation>
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,
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<figureCitation id="132A2A13FF91EC4360CC326AFEB1B362" box="[250,321,325,350]" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="1" pageNumber="476">5A–D</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30B651DFF91EC4360A132A1FD27B3FE" pageId="1" pageNumber="476" type="reference_group">
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||
<paragraph id="8BAE3696FF91EC4360A132A1FD65B39F" blockId="1.[151,936,397,511]" box="[151,661,397,419]" pageId="1" pageNumber="476">
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||
<treatmentCitationGroup id="AB0111B8FF91EC4360A132A1FD65B39F" box="[151,661,397,419]" pageId="1" pageNumber="476">
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<taxonomicName id="4C114D15FF91EC4360A132A1FD61B39F" ID-CoL="4WYWG" authority="Rathbun, 1914: 74" authorityName="Rathbun" authorityPageNumber="74" authorityYear="1914" box="[151,657,397,419]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense" subGenus="Sesarma">
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||
<emphasis id="B965EA84FF91EC4360A132A1FF1EB39F" box="[151,238,398,419]" italics="true" pageId="1" pageNumber="476">Sesarma</emphasis>
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||
(
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||
<emphasis id="B965EA84FF91EC4360CB32A1FEA4B39F" box="[253,340,398,419]" italics="true" pageId="1" pageNumber="476">Sesarma</emphasis>
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||
)
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||
<emphasis id="B965EA84FF91EC43615532A1FE3DB39F" box="[355,461,398,419]" italics="true" pageId="1" pageNumber="476">vicentense</emphasis>
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<treatmentCitation id="0AB01087FF91EC4361E432A2FD61B39F" author="Rathbun" box="[466,657,397,419]" page="74" pageId="1" pageNumber="476" year="1914">
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||
<bibRefCitation id="EF804B67FF91EC4361E432A2FD9BB39F" author="Rathbun" box="[466,619,397,419]" pageId="1" pageNumber="476" refString="Rathbun, M. J. (1914) New species of crabs of the families Grapsidae and Ocypodidae. Proceedings of the United States National Museum, 47, 69 - 85." type="journal article" year="1914">Rathbun, 1914</bibRefCitation>
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||
: 74
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||
</treatmentCitation>
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||
</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BAE3696FF91EC4360A13283FD27B3FE" blockId="1.[151,936,397,511]" box="[151,727,428,450]" pageId="1" pageNumber="476">
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||
<treatmentCitationGroup id="AB0111B8FF91EC4360A13283FD27B3FE" box="[151,727,428,450]" pageId="1" pageNumber="476">
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||
<taxonomicName id="4C114D15FF91EC4360A13283FE09B3FE" box="[151,505,428,450]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentensis" subGenus="Geosesarma">
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||
<emphasis id="B965EA84FF91EC4360A13283FF1EB3FD" box="[151,238,428,449]" italics="true" pageId="1" pageNumber="476">Sesarma</emphasis>
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||
(
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||
<emphasis id="B965EA84FF91EC4360CB3283FE8BB3FD" box="[253,379,428,449]" italics="true" pageId="1" pageNumber="476">Geosesarma</emphasis>
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||
)
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||
<emphasis id="B965EA84FF91EC4361BC3282FE09B3FE" box="[394,505,429,450]" italics="true" pageId="1" pageNumber="476">vicentensis</emphasis>
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||
</taxonomicName>
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||
—
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<treatmentCitation id="0AB01087FF91EC4362253283FD23B3FE" author="Serene" box="[531,723,428,450]" page="106" pageId="1" pageNumber="476" year="1968">
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||
<bibRefCitation id="EF804B67FF91EC4362253283FD51B3FE" author="Serene" box="[531,673,428,450]" pageId="1" pageNumber="476" refString="Serene, R. (1968 b) The Brachyura of the Indo-West Pacific Region. In: Prodromus for a Check List of the (non-planctonic) Marine Fauna of South East Asia. Singapore National Academy of Sciences, Special Publication No. 1, Fauna IIICc 3, pp. 33 - 112." type="journal article" year="1968" yearSuffix="b">Serène 1968b</bibRefCitation>
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||
: 106
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30B651DFF91EC4360A132E4FD6FB3C3" pageId="1" pageNumber="476" type="description">
|
||
<paragraph id="8BAE3696FF91EC4360A132E4FD6FB3C3" blockId="1.[151,936,397,511]" pageId="1" pageNumber="476">
|
||
<treatmentCitationGroup id="AB0111B8FF91EC4360A132E4FD6FB3C3" pageId="1" pageNumber="476">
|
||
<taxonomicName id="4C114D15FF91EC4360A132E4FE7BB3DC" box="[151,395,459,480]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentensis">
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||
<emphasis id="B965EA84FF91EC4360A132E4FE7BB3DC" box="[151,395,459,480]" italics="true" pageId="1" pageNumber="476">Geosesarma vicentensis</emphasis>
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||
</taxonomicName>
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||
—
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||
<treatmentCitation id="0AB01087FF91EC43619232E5FD6DB3DC" author="Serene" box="[420,669,458,480]" page="402" pageId="1" pageNumber="476" year="1970">
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||
<bibRefCitation id="EF804B67FF91EC43619232E5FD9DB3DC" author="Serene" box="[420,621,458,480]" pageId="1" pageNumber="476" refString="Serene, R. & Soh, C. L. (1970) New Indo-Pacific genera allied to Sesarma Say, 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4), 387 - 416." type="journal article" year="1970">Serène & Soh 1970</bibRefCitation>
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||
: 402
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||
</treatmentCitation>
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||
(list); Ng 1988: 119 (list).
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||
<taxonomicName id="4C114D15FF91EC4360A132C5FE76B3C3" box="[151,390,490,511]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense">
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||
<emphasis id="B965EA84FF91EC4360A132C5FE76B3C3" box="[151,390,490,511]" italics="true" pageId="1" pageNumber="476">Geosesarma vicentense</emphasis>
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</taxonomicName>
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||
—Ng
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||
<emphasis id="B965EA84FF91EC4361F232C5FE07B3C3" box="[452,503,489,511]" italics="true" pageId="1" pageNumber="476">et al.</emphasis>
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||
2008: 221 (list).
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C30B651DFF91EC4360A13103FBBEB08C" pageId="1" pageNumber="476" type="materials_examined">
|
||
<paragraph id="8BAE3696FF91EC4360A13103FC00B054" blockId="1.[151,1437,556,2021]" pageId="1" pageNumber="476">
|
||
<materialsCitation id="3B793CCBFF91EC4360A13103FC1CB054" ID-GBIF-Occurrence="1455709146" collectingDate="1908-11-15" collectionCode="USNM" county="northern Luzon" latitude="18.516666" location="Port San Vicente" longLatPrecision="925" longitude="122.13334" municipality="Palaui Island" pageId="1" pageNumber="476" specimenCode="USNM 45758" specimenCount="2" typeStatus="holotype">
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||
<emphasis id="B965EA84FF91EC4360A13103FECEB079" bold="true" box="[151,318,556,581]" pageId="1" pageNumber="476">
|
||
<typeStatus id="54AA8834FF91EC4360A13103FF22B079" box="[151,210,556,581]" pageId="1" pageNumber="476">Type</typeStatus>
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||
material
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||
</emphasis>
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||
.
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||
<typeStatus id="54AA8834FF91EC43617B3103FE4AB079" box="[333,442,556,581]" pageId="1" pageNumber="476" type="holotype">Holotype</typeStatus>
|
||
: male (9.2 ×
|
||
<quantity id="4CE99B73FF91EC4362783103FD58B078" box="[590,680,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="476" unit="mm" value="8.5">8.5 mm</quantity>
|
||
),
|
||
<location id="8ECE604DFF91EC43628A3102FC8AB079" LSID="urn:lsid:plazi:treatment:03B88780FF91EC4360A1320DFA70B5D8:8ECE604DFF91EC43628A3102FC8AB079" box="[700,890,556,581]" county="northern Luzon" latitude="18.516666" longLatPrecision="925" longitude="122.13334" municipality="Palaui Island" name="Port San Vicente" pageId="1" pageNumber="476">Port San Vicente</location>
|
||
(Luzon side),
|
||
<collectingMunicipality id="6BCAACECFF91EC4364293103FB43B079" box="[1055,1203,556,581]" pageId="1" pageNumber="476">Palaui Island</collectingMunicipality>
|
||
, off
|
||
<collectingCounty id="62CF4E1AFF91EC4364D13103FA68B078" box="[1255,1432,556,581]" pageId="1" pageNumber="476">northern Luzon</collectingCounty>
|
||
,
|
||
<geoCoordinate id="EE255051FF91EC4360A13160FF05B054" box="[151,245,591,616]" direction="north" orientation="latitude" pageId="1" pageNumber="476" precision="925" value="18.516666">18°31′N</geoCoordinate>
|
||
<geoCoordinate id="EE255051FF91EC4360CD3160FE70B054" box="[251,384,591,616]" direction="east" orientation="longitude" pageId="1" pageNumber="476" precision="15" value="122.13334">122°7′60″E</geoCoordinate>
|
||
, along beach, coll.
|
||
<collectingDate id="EFEBE9BEFF91EC43626E3160FCC0B054" box="[600,816,591,616]" pageId="1" pageNumber="476" value="1908-11-15">
|
||
<date id="FFAF1056FF91EC43626E3160FCC0B054" box="[600,816,591,616]" pageId="1" pageNumber="476" value="1908-11-15">15 November 1908</date>
|
||
</collectingDate>
|
||
(
|
||
<specimenCode id="DBB79EEDFF91EC436309317FFC14B054" box="[831,996,591,616]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="476">
|
||
<collectionCode id="ED00AE53FF91EC436309317FFC62B054" box="[831,914,592,616]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="476">USNM</collectionCode>
|
||
45758
|
||
</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAE3696FF91EC4360F1315BFBBEB08C" blockId="1.[151,1437,556,2021]" pageId="1" pageNumber="476">
|
||
<emphasis id="B965EA84FF91EC4360F1315BFE22B0B1" bold="true" box="[199,466,628,653]" pageId="1" pageNumber="476">Comparative material</emphasis>
|
||
.
|
||
<taxonomicName id="4C114D15FF91EC4361D4315BFC48B0B0" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[482,952,628,652]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="maculatum">
|
||
<emphasis id="B965EA84FF91EC4361D4315BFD04B0B0" box="[482,756,628,652]" italics="true" pageId="1" pageNumber="476">Geosesarma maculatum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EF804B67FF91EC436319315AFC40B0B0" author="Man" box="[815,944,628,652]" pageId="1" pageNumber="476" refString="Man, J. G. de (1892) Decapoden des Indischen Archipels. Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, 2, 265 - 527." type="journal article" year="1892">Man, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
: 1 male (12.4 ×
|
||
<quantity id="4CE99B73FF91EC43644E315BFB11B0B0" box="[1144,1249,628,652]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="1" pageNumber="476" unit="mm" value="11.9">11.9 mm</quantity>
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||
), station MALU 47, Halmahera, coll. MALU Expedition, L. Deharveng,
|
||
<date id="FFAF1056FF91EC43633A31B8FC8AB08C" box="[780,890,663,688]" pageId="1" pageNumber="476" value="1988-07">July 1988</date>
|
||
(ZRC 2017.0113).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B651DFF91EC4360F13193FF1AB694" pageId="1" pageNumber="476" type="diagnosis">
|
||
<paragraph id="8BAE3696FF91EC4360F13193FF1AB694" blockId="1.[151,1437,556,2021]" pageId="1" pageNumber="476">
|
||
<emphasis id="B965EA84FF91EC4360F13193FECAB0E9" bold="true" box="[199,314,700,725]" pageId="1" pageNumber="476">Diagnosis</emphasis>
|
||
. Carapace almost quadrate, wider than long, width to length ratio 1.1, lateral margins subparallel (
|
||
<figureCitation id="132A2A13FF91EC4360A931F0FF11B0C4" box="[159,225,735,760]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A, B); dorsal surface with regions weakly demarcated, anterior regions densely covered with low, small rounded, flattened granules (
|
||
<figureCitation id="132A2A13FF91EC4361E0302BFDE4B120" box="[470,532,772,797]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A, B); front deflexed, frontal lobes broad with subtruncate margins, separated by shallow broad concavity; postfrontal, postorbital cristae moderately low, distinct (
|
||
<figureCitation id="132A2A13FF91EC43640C3008FB8AB17C" box="[1082,1146,807,832]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A, B, E); external orbital tooth acutely triangular, directed anteriorly, outer margin almost straight, first epibranchial tooth low, second epibranchial tooth barely demarcated (
|
||
<figureCitation id="132A2A13FF91EC43626E3040FD6BB1B4" box="[600,667,879,904]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A, B). Merus of third maxilliped subovate, subequal in length to ischium; exopod slender, with long flagellum (
|
||
<figureCitation id="132A2A13FF91EC43629A30BBFCF8B191" box="[684,776,916,941]" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="1" pageNumber="476">Fig. 5A</figureCitation>
|
||
). Outer surface of palm of adult male with numerous low granules, shallow pits; inner surface granular but without distinct transverse ridge; dorsal margin of dactylus with 3–5 low conical granules on proximal part, tips not chitinous, distal half uneven but without distinct granules (
|
||
<figureCitation id="132A2A13FF91EC43655930F3FF51B624" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
F–H). Ambulatory legs each with relatively stout merus having sharp subdistal spine on dorsal margin and lateral, mesial surfaces gently rugose (
|
||
<figureCitation id="132A2A13FF91EC436238370BFDA4B600" box="[526,596,1060,1085]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A); propodus without brush-like setae on ventral margin. Male pleon triangular, relatively wide; somite 6 with convex lateral margins; telson semicircular (
|
||
<figureCitation id="132A2A13FF91EC4364493768FB32B65C" box="[1151,1218,1095,1120]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
C, D). G1 slender, distal chitinous part elongated, almost straight, directed upwards, subspatuliform, tip dilated, medially clefted (
|
||
<figureCitation id="132A2A13FF91EC4365593743FF2EB694" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="1" pageNumber="476">Fig. 5B–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B651DFF91EC4360F1379BFF4CB544" pageId="1" pageNumber="476" type="discussion">
|
||
<paragraph id="8BAE3696FF91EC4360F1379BFCF6B798" blockId="1.[151,1437,556,2021]" pageId="1" pageNumber="476">
|
||
<emphasis id="B965EA84FF91EC4360F1379BFEC4B6F1" bold="true" box="[199,308,1204,1229]" pageId="1" pageNumber="476">Remarks</emphasis>
|
||
. The taxonomic position of
|
||
<taxonomicName id="4C114D15FF91EC43624A379BFC76B6F0" box="[636,902,1204,1228]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense">
|
||
<emphasis id="B965EA84FF91EC43624A379BFC76B6F0" box="[636,902,1204,1228]" italics="true" pageId="1" pageNumber="476">Geosesarma vicentense</emphasis>
|
||
</taxonomicName>
|
||
has been uncertain, with Ng &
|
||
<bibRefCitation id="EF804B67FF91EC4364CF379BFA68B6F0" author="Takeda" box="[1273,1432,1204,1229]" pageId="1" pageNumber="476" refString="Ng, P. K. L. & Takeda, M. (1992) A new freshwater crab of the genus Geosesarma De Man, 1892 from the Philippines (Crustacea, Brachyura, Grapsidae). Proceedings of the Japanese Society of Systematic Zoology, 47, 29 - 32." type="journal article" year="1992">Takeda (1992</bibRefCitation>
|
||
: 81) questioning its placement in
|
||
<taxonomicName id="4C114D15FF91EC43622F37F7FD55B6D3" box="[537,677,1240,1263]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B965EA84FF91EC43622F37F7FD55B6D3" box="[537,677,1240,1263]" italics="true" pageId="1" pageNumber="476">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
, especially since the only known specimen was obtained from a beach.
|
||
<taxonomicName id="4C114D15FF91EC4360DD37D3FE87B72F" box="[235,375,1276,1299]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B965EA84FF91EC4360DD37D3FE87B72F" box="[235,375,1276,1299]" italics="true" pageId="1" pageNumber="476">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
species are otherwise found further inland in freshwater habitats (Ng 1988).
|
||
<taxonomicName id="4C114D15FF91EC43652637D3FEFBB70B" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense">
|
||
<emphasis id="B965EA84FF91EC43652637D3FEFBB70B" italics="true" pageId="1" pageNumber="476">Geosesarma vicentense</emphasis>
|
||
</taxonomicName>
|
||
, however, belongs to the same group of species as
|
||
<taxonomicName id="4C114D15FF91EC436372360FFB6BB704" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[836,1179,1312,1336]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="maculatum">
|
||
<emphasis id="B965EA84FF91EC436372360FFC2FB704" box="[836,991,1312,1336]" italics="true" pageId="1" pageNumber="476">G. maculatum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EF804B67FF91EC436420360EFB63B704" author="Man" box="[1046,1171,1312,1336]" pageId="1" pageNumber="476" refString="Man, J. G. de (1892) Decapoden des Indischen Archipels. Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, 2, 265 - 527." type="journal article" year="1892">Man, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C114D15FF91EC436490360FFF17B761" authority="Serene, 1968a" authorityName="Serene" authorityYear="1968" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="ternatense">
|
||
<emphasis id="B965EA84FF91EC436490360FFAC9B704" box="[1190,1337,1312,1336]" italics="true" pageId="1" pageNumber="476">G. ternatense</emphasis>
|
||
(
|
||
<bibRefCitation id="EF804B67FF91EC436571360FFF2FB761" author="Serene" pageId="1" pageNumber="476" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, 39 (5), 1084 - 1095." type="journal article" year="1968" yearSuffix="a">Serène, 1968a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C114D15FF91EC436114366BFD30B760" authority="Ng, Liu & Schubart, 2004" authorityName="Ng, Liu & Schubart" authorityYear="2004" box="[290,704,1348,1373]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="hednon">
|
||
<emphasis id="B965EA84FF91EC436114366BFE63B760" box="[290,403,1348,1372]" italics="true" pageId="1" pageNumber="476">G. hednon</emphasis>
|
||
Ng,
|
||
<bibRefCitation id="EF804B67FF91EC4361FF366BFD30B760" author="Liu" box="[457,704,1348,1373]" pageId="1" pageNumber="476" refString="Ng, P. K. L., Liu, H. - C. & Schubart, C. D. (2004) Geosesarma hednon, a new species of terrestrial crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Taiwan and the Philippines. Raffles Bulletin of Zoology, 52 (1), 239 - 249." type="journal article" year="2004">Liu & Schubart, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
, in which the G1 is slender, straight, with the chitinous distal part subspatuliform and directed upwards. Most of the species in this group have small eggs and almost certainly have planktotrophic larvae that must be released into the sea.
|
||
</paragraph>
|
||
<paragraph id="8BAE3696FF91EC4360F13680FF4CB544" blockId="1.[151,1437,556,2021]" pageId="1" pageNumber="476">
|
||
Compared to
|
||
<taxonomicName id="4C114D15FF91EC436150369FFDF4B7F4" box="[358,516,1456,1480]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="maculatum">
|
||
<emphasis id="B965EA84FF91EC436150369FFDF4B7F4" box="[358,516,1456,1480]" italics="true" pageId="1" pageNumber="476">G. maculatum</emphasis>
|
||
</taxonomicName>
|
||
, the frontal orbital lobes of
|
||
<taxonomicName id="4C114D15FF91EC436367369FFC18B7FB" box="[849,1000,1456,1479]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense">
|
||
<emphasis id="B965EA84FF91EC436367369FFC18B7FB" box="[849,1000,1456,1479]" italics="true" pageId="1" pageNumber="476">G. vicentense</emphasis>
|
||
</taxonomicName>
|
||
are more convex (
|
||
<figureCitation id="132A2A13FF91EC4364F03680FAF8B7F4" box="[1222,1288,1455,1480]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A, B) (lobes truncate in
|
||
<taxonomicName id="4C114D15FF91EC43612236FBFE5EB7D0" box="[276,430,1492,1516]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="maculatum">
|
||
<emphasis id="B965EA84FF91EC43612236FBFE5EB7D0" box="[276,430,1492,1516]" italics="true" pageId="1" pageNumber="476">G. maculatum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="132A2A13FF91EC43618B36FBFE0EB7D0" box="[445,510,1492,1517]" captionStart="FIGURE 2" captionStartId="3.[151,250,1277,1299]" captionTargetBox="[180,1408,187,1255]" captionTargetId="figure@3.[180,1408,187,1255]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma maculatum (De Man, 1892), male (12.4 × 11.9 mm) (ZRC 2017.0113), Halmahera. A, overall habitus; B, dorsal view of carapace; C, ventral view of cephalothorax showing thoracic sternum and pleon." httpUri="https://zenodo.org/record/495552/files/figure.png" pageId="1" pageNumber="476">Fig. 2</figureCitation>
|
||
A, B); the external orbital tooth is relatively less prominent and separated from rest of margin by a shallow cleft (
|
||
<figureCitation id="132A2A13FF91EC4361D936D8FDC0B42C" box="[495,560,1527,1552]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A, B) (tooth more prominent and separated from margin by deep cleft in
|
||
<taxonomicName id="4C114D15FF91EC4365B236D7FEE3B408" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="maculatum">
|
||
<emphasis id="B965EA84FF91EC4365B236D7FEE3B408" italics="true" pageId="1" pageNumber="476">G. maculatum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="132A2A13FF91EC4361153533FE95B408" box="[291,357,1564,1589]" captionStart="FIGURE 2" captionStartId="3.[151,250,1277,1299]" captionTargetBox="[180,1408,187,1255]" captionTargetId="figure@3.[180,1408,187,1255]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma maculatum (De Man, 1892), male (12.4 × 11.9 mm) (ZRC 2017.0113), Halmahera. A, overall habitus; B, dorsal view of carapace; C, ventral view of cephalothorax showing thoracic sternum and pleon." httpUri="https://zenodo.org/record/495552/files/figure.png" pageId="1" pageNumber="476">Fig. 2</figureCitation>
|
||
A, B); the ambulatory meri are relatively slender (
|
||
<figureCitation id="132A2A13FF91EC4363AB3533FC2CB408" box="[925,988,1564,1589]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A) (meri broader in
|
||
<taxonomicName id="4C114D15FF91EC4364F23533FA90B408" box="[1220,1376,1564,1588]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="maculatum">
|
||
<emphasis id="B965EA84FF91EC4364F23533FA90B408" box="[1220,1376,1564,1588]" italics="true" pageId="1" pageNumber="476">G. maculatum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="132A2A13FF91EC4365463533FF56B464" captionStart="FIGURE 2" captionStartId="3.[151,250,1277,1299]" captionTargetBox="[180,1408,187,1255]" captionTargetId="figure@3.[180,1408,187,1255]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma maculatum (De Man, 1892), male (12.4 × 11.9 mm) (ZRC 2017.0113), Halmahera. A, overall habitus; B, dorsal view of carapace; C, ventral view of cephalothorax showing thoracic sternum and pleon." httpUri="https://zenodo.org/record/495552/files/figure.png" pageId="1" pageNumber="476">Fig. 2</figureCitation>
|
||
A); and the male pleonal somite 6 is proportionately broader (
|
||
<figureCitation id="132A2A13FF91EC43634F3510FC4AB464" box="[889,954,1599,1624]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
D) (somite 6 proportionately longer in
|
||
<taxonomicName id="4C114D15FF91EC4365B2356FFEE3B440" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="maculatum">
|
||
<emphasis id="B965EA84FF91EC4365B2356FFEE3B440" italics="true" pageId="1" pageNumber="476">G. maculatum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="132A2A13FF91EC436115354BFE95B440" box="[291,357,1636,1661]" captionStart="FIGURE 2" captionStartId="3.[151,250,1277,1299]" captionTargetBox="[180,1408,187,1255]" captionTargetId="figure@3.[180,1408,187,1255]" captionTargetPageId="3" captionText="FIGURE 2. Geosesarma maculatum (De Man, 1892), male (12.4 × 11.9 mm) (ZRC 2017.0113), Halmahera. A, overall habitus; B, dorsal view of carapace; C, ventral view of cephalothorax showing thoracic sternum and pleon." httpUri="https://zenodo.org/record/495552/files/figure.png" pageId="1" pageNumber="476">Fig. 2</figureCitation>
|
||
C) (see also Ng
|
||
<emphasis id="B965EA84FF91EC43622A354AFDA4B440" box="[540,596,1636,1660]" italics="true" pageId="1" pageNumber="476">et al.</emphasis>
|
||
2004: figs. 1A, B, 2A).
|
||
<taxonomicName id="4C114D15FF91EC43635E354BFB81B440" box="[872,1137,1636,1660]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="ternatense">
|
||
<emphasis id="B965EA84FF91EC43635E354BFB81B440" box="[872,1137,1636,1660]" italics="true" pageId="1" pageNumber="476">Geosesarma ternatense</emphasis>
|
||
</taxonomicName>
|
||
is distinct in that the male pleonal somite 6 is proportionately very broad with strongly convex lateral margins and the telson is relatively wider (Ng
|
||
<emphasis id="B965EA84FF91EC4361273582FEBAB4F8" box="[273,330,1708,1732]" italics="true" pageId="1" pageNumber="476">et al.</emphasis>
|
||
2004 fig. 8B) (somite 6 and telson less wide in
|
||
<taxonomicName id="4C114D15FF91EC43635E3583FC0CB4F8" box="[872,1020,1708,1732]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense">
|
||
<emphasis id="B965EA84FF91EC43635E3583FC0CB4F8" box="[872,1020,1708,1732]" italics="true" pageId="1" pageNumber="476">G. vicentense</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="132A2A13FF91EC43643A3583FBBAB4F8" box="[1036,1098,1708,1733]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
D).
|
||
<taxonomicName id="4C114D15FF91EC43644C3583FA72B4F8" box="[1146,1410,1708,1732]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense">
|
||
<emphasis id="B965EA84FF91EC43644C3583FA72B4F8" box="[1146,1410,1708,1732]" italics="true" pageId="1" pageNumber="476">Geosesarma vicentense</emphasis>
|
||
</taxonomicName>
|
||
is closest to
|
||
<taxonomicName id="4C114D15FF91EC43613935FFFE76B4D4" box="[271,390,1744,1768]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="hednon">
|
||
<emphasis id="B965EA84FF91EC43613935FFFE76B4D4" box="[271,390,1744,1768]" italics="true" pageId="1" pageNumber="476">G. hednon</emphasis>
|
||
</taxonomicName>
|
||
in general form, but can be distinguished by its external orbital tooth being distinct and separated from the rest of the margin by a small cleft (
|
||
<figureCitation id="132A2A13FF91EC4362C535DBFCC1B530" box="[755,817,1780,1805]" captionStart="FIGURE 1" captionStartId="2.[151,250,1693,1715]" captionTargetBox="[151,1436,183,1668]" captionTargetId="figure@2.[151,1436,183,1668]" captionTargetPageId="2" captionText="FIGURE 1. Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758). A, overall habitus; B, dorsal view of carapace; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of left chelae at different angles; H, dorsal view of left chela." httpUri="https://zenodo.org/record/495551/files/figure.png" pageId="1" pageNumber="476">Fig. 1</figureCitation>
|
||
A, B) (tooth subparallel to the rest of the lateral margin and separated by a fissure in
|
||
<taxonomicName id="4C114D15FF91EC4361EF3437FDBAB50C" box="[473,586,1816,1840]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="hednon">
|
||
<emphasis id="B965EA84FF91EC4361EF3437FDBAB50C" box="[473,586,1816,1840]" italics="true" pageId="1" pageNumber="476">G. hednon</emphasis>
|
||
</taxonomicName>
|
||
; cf. Ng
|
||
<emphasis id="B965EA84FF91EC4362923436FD2CB50C" box="[676,732,1816,1840]" italics="true" pageId="1" pageNumber="476">et al.</emphasis>
|
||
2004: figs. 9B, 11B) and the slender chitinous part of the G1 is relatively shorter (
|
||
<figureCitation id="132A2A13FF91EC4361503413FE2FB569" box="[358,479,1852,1877]" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="1" pageNumber="476">Fig. 5B–D</figureCitation>
|
||
) (chitinous part relatively longer in
|
||
<taxonomicName id="4C114D15FF91EC43634F3413FC1BB568" box="[889,1003,1852,1876]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="hednon">
|
||
<emphasis id="B965EA84FF91EC43634F3413FC1BB568" box="[889,1003,1852,1876]" italics="true" pageId="1" pageNumber="476">G. hednon</emphasis>
|
||
</taxonomicName>
|
||
; cf. Ng
|
||
<emphasis id="B965EA84FF91EC43647C3412FB73B568" box="[1098,1155,1852,1876]" italics="true" pageId="1" pageNumber="476">et al.</emphasis>
|
||
2004: figs. 13C–G,14C– H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B651DFF91EC4360F134ABFA70B5D8" pageId="1" pageNumber="476" type="reference_group">
|
||
<paragraph id="8BAE3696FF91EC4360F134ABFA70B5D8" blockId="1.[151,1437,556,2021]" pageId="1" pageNumber="476">
|
||
<taxonomicName id="4C114D15FF91EC4360F134ABFD63B5A0" ID-CoL="3FS9V" authority="Ng & Takeda, 1992" authorityName="Ng & Takeda" authorityYear="1992" box="[199,659,1924,1949]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="protos">
|
||
<emphasis id="B965EA84FF91EC4360F134ABFE53B5A0" box="[199,419,1924,1948]" italics="true" pageId="1" pageNumber="476">Geosesarma protos</emphasis>
|
||
Ng &
|
||
<bibRefCitation id="EF804B67FF91EC4361CE34ABFD63B5A0" author="Takeda" box="[504,659,1924,1949]" pageId="1" pageNumber="476" refString="Ng, P. K. L. & Takeda, M. (1992) A new freshwater crab of the genus Geosesarma De Man, 1892 from the Philippines (Crustacea, Brachyura, Grapsidae). Proceedings of the Japanese Society of Systematic Zoology, 47, 29 - 32." type="journal article" year="1992">Takeda, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
, also described from Luzon, is very different from
|
||
<taxonomicName id="4C114D15FF91EC4364D034ABFA8DB5A0" ID-CoL="6KB9Q" box="[1254,1405,1924,1948]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="vicentense">
|
||
<emphasis id="B965EA84FF91EC4364D034ABFA8DB5A0" box="[1254,1405,1924,1948]" italics="true" pageId="1" pageNumber="476">G. vicentense</emphasis>
|
||
</taxonomicName>
|
||
in possessing an external orbital tooth that is prominent and directed obliquely laterally, a third maxilliped in which the exopod has no flagellum and the G1 is stout with a short chitinous part (Ng &
|
||
<bibRefCitation id="EF804B67FF91EC43641934E3FB32B5D8" author="Takeda" box="[1071,1218,1996,2021]" pageId="1" pageNumber="476" refString="Ng, P. K. L. & Takeda, M. (1992) A new freshwater crab of the genus Geosesarma De Man, 1892 from the Philippines (Crustacea, Brachyura, Grapsidae). Proceedings of the Japanese Society of Systematic Zoology, 47, 29 - 32." type="journal article" year="1992">Takeda 1992</bibRefCitation>
|
||
: fig. 1A, B, G–I)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |