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<document id="379C460243794A67B1AC9377180CD3EE" ID-CLB-Dataset="22494" ID-DOI="10.11646/zootaxa.4765.1.1" ID-GBIF-Dataset="8af5a092-f062-48c7-8d7c-f96ae6282bfa" ID-ISSN="1175-5334" ID-Zenodo-Dep="3774174" ID-ZooBank="urn:lsid:zoobank.org:pub:C442D94C-0EB4-4509-B762-913707214819" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588094957734" checkinUser="carolina" docAuthor="Hodgson, Chris" docDate="2020" docId="03B2EA640AEB46942CFCFAADFA57D4B7" docLanguage="en" docName="zootaxa.4765.1.1.pdf.imf" docOrigin="Zootaxa 4765 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aclerdidae Cockerell 1905" docType="treatment" docVersion="6" lastPageNumber="199" masterDocId="FF8B921C0A2F46522C6BFF8EFFCBD747" masterDocTitle="A review of neococcid scale insects (Hemiptera: Sternorrhyncha: Coccomorpha) based on the morphology of the adult males" masterLastPageNumber="264" masterPageNumber="1" pageNumber="197" updateTime="1698837970293" updateUser="ExternalLinkService">
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<mods:title id="944363D3DB1DAAF328625ADAEAD933F4">A review of neococcid scale insects (Hemiptera: Sternorrhyncha: Coccomorpha) based on the morphology of the adult males</mods:title>
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<treatment id="03B2EA640AEB46942CFCFAADFA57D4B7" ID-DOI="http://doi.org/10.5281/zenodo.3796876" ID-GBIF-Taxon="163857737" ID-Zenodo-Dep="3796876" LSID="urn:lsid:plazi:treatment:03B2EA640AEB46942CFCFAADFA57D4B7" httpUri="http://treatment.plazi.org/id/03B2EA640AEB46942CFCFAADFA57D4B7" lastPageId="198" lastPageNumber="199" pageId="196" pageNumber="197">
<subSubSection id="C30108F90AEB46962CFCFAADFDEED27A" box="[151,549,1315,1341]" pageId="196" pageNumber="197" type="nomenclature">
<paragraph id="8BA45B720AEB46962CFCFAADFDEED27A" blockId="196.[151,549,1315,1341]" box="[151,549,1315,1341]" pageId="196" pageNumber="197">
<taxonomicName id="4C1B20F10AEB46962CFCFAADFDEED27A" ID-CoL="5XP" ID-ENA="249417" authority="Cockerell, 1905" authorityName="Cockerell" authorityYear="1905" box="[151,549,1315,1341]" class="Insecta" family="Aclerdidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">
<emphasis id="B96F87600AEB46962CFCFAADFDEED27A" bold="true" box="[151,549,1315,1341]" pageId="196" pageNumber="197">
ACLERDIDAE
<bibRefCitation id="EF8A26830AEB46962D0FFAADFDEED27A" author="Cockerell" box="[356,549,1315,1341]" firstAuthor="Cockerell" pageId="196" pageNumber="197" pagination="189 - 203" refId="ref199339" refString="Cockerell, T. D. A. (1905) Tables for the identification of Rocky Mountain Coccidae (Scale insects and mealybugs). Colorado University Studies, 2, 189 - 203." type="journal article" year="1905">Cockerell, 1905</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30108F90AEB46962CFCFAE9FE3AD2FB" pageId="196" pageNumber="197" type="reference_group">
<paragraph id="8BA45B720AEB46962CFCFAE9FC37D238" blockId="196.[151,1436,1382,1469]" box="[151,1020,1382,1407]" pageId="196" pageNumber="197">
<treatmentCitationGroup id="AB0B7C5C0AEB46962CFCFAE9FC37D238" box="[151,1020,1382,1407]" pageId="196" pageNumber="197">
<treatmentCitation id="0ABA7D630AEB46962CFCFAE9FE6FD238" box="[151,420,1382,1407]" pageId="196" pageNumber="197">
<taxonomicName id="4C1B20F10AEB46962CFCFAE9FE6FD238" ID-CoL="MZX" authority="Signoret 1874" authorityName="Signoret" authorityYear="1874" box="[151,420,1382,1407]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962CFCFAE9FF2DD238" box="[151,230,1383,1407]" italics="true" pageId="196" pageNumber="197">Aclerda</emphasis>
<bibRefCitation id="EF8A26830AEB46962C86FAE8FE6FD238" author="Signoret, V." box="[237,420,1382,1407]" pageId="196" pageNumber="197" pagination="87 - 106" refId="ref211088" refString="Signoret, V. (1874) Essai sur les cochenilles ou gallinsectes (Homopteres-Coccides), 12 e partie. Annales de la Societe Entomologique de France, Serie 5, 4, 87 - 106." type="journal article" year="1874">Signoret 1874, 96</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54A0E5D00AEB46962DC5FAE9FE2BD238" box="[430,480,1383,1407]" pageId="196" pageNumber="197">Type</typeStatus>
species:
<taxonomicName id="4C1B20F10AEB46962E56FAE9FCA2D238" ID-CoL="64JWP" authority="Signoret" authorityName="Signoret" authorityYear="1874" box="[573,873,1382,1407]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="B96F87600AEB46962E56FAE9FCC6D238" box="[573,781,1383,1407]" italics="true" pageId="196" pageNumber="197">Aclerda subterranea</emphasis>
Signoret
</taxonomicName>
by monotypy.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA45B720AEB46962CFCFA08FE3AD2FB" blockId="196.[151,1436,1382,1469]" pageId="196" pageNumber="197">
<treatmentCitationGroup id="AB0B7C5C0AEB46962CFCFA08FE3AD2FB" pageId="196" pageNumber="197">
<treatmentCitation id="0ABA7D630AEB46962CFCFA08FDD6D2D9" box="[151,541,1413,1438]" pageId="196" pageNumber="197">
<taxonomicName id="4C1B20F10AEB46962CFCFA08FDD6D2D9" authority="Cockerell 1896 b" authorityName="Cockerell" authorityYear="1896" box="[151,541,1413,1438]" class="Insecta" family="Aclerdidae" genus="Pseudolecanium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962CFCFA08FEF5D2D9" box="[151,318,1414,1438]" italics="true" pageId="196" pageNumber="197">Pseudolecanium</emphasis>
<bibRefCitation id="EF8A26830AEB46962D2CFA0BFDD6D2D9" author="Cockerell, T. D. A." box="[327,541,1413,1438]" pageId="196" pageNumber="197" pagination="18 - 21" refId="ref198916" refString="Cockerell, T. D. A. (1896 b) Preliminary diagnoses of new Coccidae. Psyche, Supplement 7, 18 - 21. https: // doi. org / 10.1155 / 1896 / 92937" type="journal article" year="1896">Cockerell 1896b, 19</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54A0E5D00AEB46962E42FA08FD90D2D9" box="[553,603,1414,1438]" pageId="196" pageNumber="197">Type</typeStatus>
species:
<taxonomicName id="4C1B20F10AEB46962ED6FA08FBFBD2D9" authority="Cockerell." authorityName="Cockerell." box="[701,1072,1413,1438]" class="Insecta" family="Aclerdidae" genus="Pseudolecanium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="tokionis">
<emphasis id="B96F87600AEB46962ED6FA08FC76D2D9" box="[701,957,1414,1438]" italics="true" pageId="196" pageNumber="197">Pseudolecanium tokionis</emphasis>
Cockerell.
</taxonomicName>
<typeStatus id="54A0E5D00AEB4696285CFA08FBA2D2D9" box="[1079,1129,1414,1438]" pageId="196" pageNumber="197">Type</typeStatus>
designation unknown. Junior synonym,
<treatmentCitation id="0ABA7D630AEB46962D5AFA2AFE26D2FB" box="[305,493,1444,1469]" pageId="196" pageNumber="197">
<bibRefCitation id="EF8A26830AEB46962D5AFA2AFE26D2FB" author="Fernald, M. E." box="[305,493,1444,1469]" pageId="196" pageNumber="197" pagination="1 - 360" refId="ref200607" refString="Fernald, M. E. (1903) A catalogue of the Coccidae of the world. Bulletin of the Hatch Experiment Station of the Massachusetts Agricultural College, 88, 1 - 360." type="journal article" year="1903">Fernald 1903, 210</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30108F90AEB46942CFCFA64FAE1D6CC" lastPageId="198" lastPageNumber="199" pageId="196" pageNumber="197" type="discussion">
<paragraph id="8BA45B720AEB46962CFCFA64FE6DD0BA" blockId="196.[151,1437,1514,2045]" pageId="196" pageNumber="197">
<emphasis id="B96F87600AEB46962CFCFA64FEF9D142" bold="true" box="[151,306,1514,1541]" pageId="196" pageNumber="197">Introduction.</emphasis>
<bibRefCitation id="EF8A26830AEB46962D53FA65FE29D142" author="Signoret" box="[312,482,1515,1541]" firstAuthor="Signoret" pageId="196" pageNumber="197" pagination="87 - 106" refId="ref211088" refString="Signoret, V. (1874) Essai sur les cochenilles ou gallinsectes (Homopteres-Coccides), 12 e partie. Annales de la Societe Entomologique de France, Serie 5, 4, 87 - 106." type="journal article" year="1874">Signoret (1874)</bibRefCitation>
introduced the genus
<taxonomicName id="4C1B20F10AEB46962EA4FA65FCEFD142" authorityName="Signoret" authorityYear="1874" box="[719,804,1515,1541]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962EA4FA65FCEFD142" box="[719,804,1515,1541]" italics="true" pageId="196" pageNumber="197">Aclerda</emphasis>
</taxonomicName>
for a group of scale insects that mainly inhabit grasses and exhibit rather specialised morphological characters. Signoret considered
<taxonomicName id="4C1B20F10AEB46962FD9F981FBCCD16E" authorityName="Signoret" authorityYear="1874" box="[946,1031,1551,1577]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962FD9F981FBCCD16E" box="[946,1031,1551,1577]" italics="true" pageId="196" pageNumber="197">Aclerda</emphasis>
</taxonomicName>
to be closely related to his soft scale genus
<taxonomicName id="4C1B20F10AEB46962C8BF9BAFE90D10A" box="[224,347,1588,1613]" class="Insecta" family="Coccidae" genus="Lecanopsis" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962C8BF9BAFE90D10A" box="[224,347,1588,1613]" italics="true" pageId="196" pageNumber="197">Lecanopsis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C1B20F10AEB46962D01F9BDFE0BD10A" authorityName="Signoret" authorityYear="1874" box="[362,448,1587,1613]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962D01F9BDFE0BD10A" box="[362,448,1587,1613]" italics="true" pageId="196" pageNumber="197">Aclerda</emphasis>
</taxonomicName>
was later placed in its own family,
<taxonomicName id="4C1B20F10AEB46962F25F9BDFC03D10A" authorityName="Cockerell" authorityYear="1905" box="[846,968,1587,1613]" class="Insecta" family="Aclerdidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">Aclerdidae</taxonomicName>
, by
<bibRefCitation id="EF8A26830AEB46962F93F9BDFB50D10A" author="Teague" box="[1016,1179,1586,1613]" firstAuthor="Teague" pageId="196" pageNumber="197" pagination="432 - 441" refId="ref212136" refString="Teague, M. M. (1925) A review of the genus Aclerda (Hemiptera; Coccidoidea). Annals of the Entomological Society of America, 18, 432 - 441. https: // doi. org / 10.1093 / aesa / 18.4.432" type="journal article" year="1925">Teague (1925)</bibRefCitation>
. Since then, the family was critically examined by
<bibRefCitation id="EF8A26830AEB46962DA3F9D9FD5DD136" author="McConnell" box="[456,662,1622,1649]" firstAuthor="McConnell" pageId="196" pageNumber="197" pagination="1 - 121" refId="ref208272" refString="McConnell, H. S. (1954) A classification of the coccid family Aclerdidae (Coccoidea, Homoptera). Bulletin of the Maryland Agriculture Experiment Station, 75, 1 - 121." type="journal article" year="1954">McConnell (1954)</bibRefCitation>
, who introduced 2 additional genera,
<taxonomicName id="4C1B20F10AEB46962852F9D9FA94D136" authority="McConnell" authorityName="McConnell" authorityYear="1954" box="[1081,1375,1623,1649]" class="Insecta" family="Aclerdidae" genus="Nipponaclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962852F9D9FB17D136" box="[1081,1244,1623,1649]" italics="true" pageId="196" pageNumber="197">Nipponaclerda</emphasis>
McConnell
</taxonomicName>
from
<collectingCountry id="F30C1BE20AEB46962CFCF9F5FF13D1D2" box="[151,216,1659,1685]" name="Japan" pageId="196" pageNumber="197">Japan</collectingCountry>
, clearly closely related to
<taxonomicName id="4C1B20F10AEB46962D9FF9F5FD82D1D2" authorityName="Signoret" authorityYear="1874" box="[500,585,1659,1685]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962D9FF9F5FD82D1D2" box="[500,585,1659,1685]" italics="true" pageId="196" pageNumber="197">Aclerda</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1B20F10AEB46962EEFF9F5FC62D1D2" authority="McConnell" authorityName="McConnell" authorityYear="1954" box="[644,937,1659,1685]" class="Insecta" family="Aclerdidae" genus="Rhodesaclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962EEFF9F5FCECD1D2" box="[644,807,1659,1685]" italics="true" pageId="196" pageNumber="197">Rhodesaclerda</emphasis>
McConnell
</taxonomicName>
which occurs on mistletoe in
<collectingCountry id="F30C1BE20AEB46962884F9F5FAA3D1D2" box="[1263,1384,1659,1685]" name="Zimbabwe" pageId="196" pageNumber="197">Zimbabwe</collectingCountry>
. The
<taxonomicName id="4C1B20F10AEB46962CFCF911FEC5D1FE" authorityName="Cockerell" authorityYear="1905" box="[151,270,1695,1721]" class="Insecta" family="Aclerdidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">Aclerdidae</taxonomicName>
is now considered to contain two subfamilies (
<bibRefCitation id="EF8A26830AEB46962F4FF911FBF0D1FE" author="Hodgson &amp; Millar" box="[804,1083,1695,1721]" firstAuthor="Hodgson" pageId="196" pageNumber="197" pagination="469 - 517" refId="ref204361" refString="Hodgson, C. J. &amp; Millar, I. M. (2002) A new subfamily, two new genera and three new species of Aclerdidae (Hemiptera: Coccoidea) from southern Africa, with a phylogenetic analysis of relationships. Systematic Entomology, 27 (4), 469 - 517. https: // doi. org / 10.1046 / j. 1365 - 3113.2002.00189. x" type="journal article" year="2002">Hodgson &amp; Millar 2002</bibRefCitation>
): Rhodesaclerdinae Hodgson &amp; Miller, with
<taxonomicName id="4C1B20F10AEB46962D4BF94DFE08D19A" authorityName="McConnell" authorityYear="1954" box="[288,451,1731,1757]" class="Insecta" family="Aclerdidae" genus="Rhodesaclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962D4BF94DFE08D19A" box="[288,451,1731,1757]" italics="true" pageId="196" pageNumber="197">Rhodesaclerda</emphasis>
</taxonomicName>
(3 spp.) and
<taxonomicName id="4C1B20F10AEB46962E3FF94DFD33D19A" authorityName="Hodgson &amp; Millar" authorityYear="2002" box="[596,760,1731,1757]" class="Insecta" family="Aclerdidae" genus="Kwazulaclerda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962E3FF94DFD33D19A" box="[596,760,1731,1757]" italics="true" pageId="196" pageNumber="197">Kwazulaclerda</emphasis>
</taxonomicName>
(
<specimenCount id="9D1D90FB0AEB46962F61F94DFCF5D19A" box="[778,830,1731,1757]" pageId="196" pageNumber="197" type="generic">1 sp.</specimenCount>
), both from southern Africa, and
<taxonomicName id="4C1B20F10AEB469628C6F94DFAEFD19A" box="[1197,1316,1731,1757]" pageId="196" pageNumber="197">Aclerdinae</taxonomicName>
Cockerell, including
<taxonomicName id="4C1B20F10AEB46962D6EF969FE91D046" authorityName="Signoret" authorityYear="1874" box="[261,346,1767,1793]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962D6EF969FE91D046" box="[261,346,1767,1793]" italics="true" pageId="196" pageNumber="197">Aclerda</emphasis>
</taxonomicName>
(52 spp.),
<taxonomicName id="4C1B20F10AEB46962DA4F969FDAFD046" authorityName="Hodgson &amp; Millar" authorityYear="2002" box="[463,612,1767,1793]" class="Insecta" family="Aclerdidae" genus="Lecanaclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962DA4F969FDAFD046" box="[463,612,1767,1793]" italics="true" pageId="196" pageNumber="197">Lecanaclerda</emphasis>
</taxonomicName>
(
<specimenCount id="9D1D90FB0AEB46962E1EF969FD6CD046" box="[629,679,1767,1793]" pageId="196" pageNumber="197" type="generic">1 sp.</specimenCount>
from
<collectingCountry id="F30C1BE20AEB46962E82F969FCBED045" box="[745,885,1767,1794]" name="South Africa" pageId="196" pageNumber="197">South Africa</collectingCountry>
) and
<taxonomicName id="4C1B20F10AEB46962FD9F969FB9ED046" authorityName="McConnell" authorityYear="1954" box="[946,1109,1767,1793]" class="Insecta" family="Aclerdidae" genus="Nipponaclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AEB46962FD9F969FB9ED046" box="[946,1109,1767,1793]" italics="true" pageId="196" pageNumber="197">Nipponaclerda</emphasis>
</taxonomicName>
(4 spp. from Asia). Currently, therefore, it includes five genera and 61 species (
<bibRefCitation id="EF8A26830AEB46962EDFF885FC29D062" author="Garcia Morales" box="[692,994,1803,1829]" etAl="et al." firstAuthor="Garcia Morales" pageId="196" pageNumber="197" refId="ref201635" refString="Garcia Morales, M., Denno, B. D., Miller, D. R., Miller, G. L., Ben-Dov, Y. &amp; Hardy, N. B. (2019) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: https: // data. nal. usda. gov / dataset / scalenet-scaleinsects-coccoidea-database (accessed 12 August 2019)" type="url" year="2019">
García Morales
<emphasis id="B96F87600AEB46962F03F882FC6BD062" box="[872,928,1803,1829]" italics="true" pageId="196" pageNumber="197">et al.</emphasis>
2019
</bibRefCitation>
). The family
<taxonomicName id="4C1B20F10AEB4696281FF885FB20D062" authorityName="Cockerell" authorityYear="1905" box="[1140,1259,1803,1829]" class="Insecta" family="Aclerdidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">Aclerdidae</taxonomicName>
is generally accepted as being either sister to
<taxonomicName id="4C1B20F10AEB46962D9DF8A1FD50D00E" authorityName="Silvestri" authorityYear="1939" box="[502,667,1839,1865]" class="Insecta" family="Micrococcidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">Micrococcidae</taxonomicName>
, with the micrococcids plus aclerdids sister to the coccids (
<bibRefCitation id="EF8A26830AEB46962953F8A1FE8BD02A" author="Miller, D. R. &amp; Williams, D. J." pageId="196" pageNumber="197" pagination="199 - 247" refId="ref209066" refString="Miller, D. R. &amp; Williams, D. J. (1995) Systematic revision of the Family Micrococcidae (Homoptera: Coccoidea), with a discussion of its relationships, and a description of a gynandromorph. Bollettino del Laboratorio di Entomologia Agraria ' Filippo Silvestri'. Portici, 50, 199 - 247. [1993]" type="journal article" year="1995">Miller &amp; Williams, 1995</bibRefCitation>
) or sister to
<taxonomicName id="4C1B20F10AEB46962DA2F8DDFDE5D02A" box="[457,558,1875,1901]" class="Insecta" family="Coccidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">Coccidae</taxonomicName>
(
<bibRefCitation id="EF8A26830AEB46962E56F8DDFC80D02A" author="Miller &amp; Hodgson" box="[573,843,1875,1901]" firstAuthor="Miller" pageId="196" pageNumber="197" pagination="1 - 452" refId="ref208664" refString="Miller, D. R. &amp; Hodgson, C. J. (1997) 1.1. 3.7 Phylogeny. In: Soft Scale Insects: Their Biology, Natural Enemies and Control. Vol. 7 A. Elsevier Amsterdam, New York, pp. 1 - 452. https: // doi. org / 10.1016 / S 1572 - 4379 (97) 80053 - X" type="book chapter" year="1997">Miller &amp; Hodgson 1997</bibRefCitation>
;
<bibRefCitation id="EF8A26830AEB46962F3DF8DDFC31D02A" author="Hodgson, C. J." box="[854,1018,1875,1901]" pageId="196" pageNumber="197" pagination="129 - 137" refId="ref203564" refString="Hodgson, C. J. (2002) Preliminary phylogeny of some non-margarodid Coccoidea (Hemiptera) based on adult male characters. Bollettino di Zoologia Agraria e di Bachicoltura, Milano, 33 (3), 129 - 137. [2001]" type="journal article" year="2002">Hodgson 2002</bibRefCitation>
;
<bibRefCitation id="EF8A26830AEB4696286DF8DDFADAD02A" author="Hodgson &amp; Millar" box="[1030,1297,1875,1901]" firstAuthor="Hodgson" pageId="196" pageNumber="197" pagination="469 - 517" refId="ref204361" refString="Hodgson, C. J. &amp; Millar, I. M. (2002) A new subfamily, two new genera and three new species of Aclerdidae (Hemiptera: Coccoidea) from southern Africa, with a phylogenetic analysis of relationships. Systematic Entomology, 27 (4), 469 - 517. https: // doi. org / 10.1046 / j. 1365 - 3113.2002.00189. x" type="journal article" year="2002">Hodgson &amp; Millar 2002</bibRefCitation>
;
<bibRefCitation id="EF8A26830AEB46962976F8DDFED6D0D6" author="Hodgson &amp; Hardy" firstAuthor="Hodgson" pageId="196" pageNumber="197" pagination="794 - 804" refId="ref203953" refString="Hodgson, C. J. &amp; Hardy, N. B. (2013) The phylogeny of the superfamily Coccoidea (Hemiptera: Sternorrhyncha) based on the morphology of extant and extinct macropterous males. Systematic Entomology, 38, 794 - 804. https: // doi. org / 10.1111 / syen. 12030" type="journal article" year="2013">Hodgson &amp; Hardy 2013</bibRefCitation>
;
<bibRefCitation id="EF8A26830AEB46962D43F8F9FDD0D0D5" author="Vea &amp; Grimaldi" box="[296,539,1911,1938]" firstAuthor="Vea" pageId="196" pageNumber="197" pagination="23487" refId="ref212464" refString="Vea, I. M. &amp; Grimaldi, D. A. (2016) Putting scales into evolutionary time: the divergence of major scale insect lineages (Hemiptera) predates the radiation of modern angiosperm hosts. Scientific Reports, 6, 23487. https: // doi. org / 10.1038 / srep 23487" type="journal article" year="2016">Vea &amp; Grimaldi 2016</bibRefCitation>
). The only molecular study that has included a member of the Acleridae (
<taxonomicName id="4C1B20F10AEB4696292CF8F9FEA7D0F2" authority="Kuwana" authorityName="Kuwana" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="takahashii">
<emphasis id="B96F87600AEB4696292CF8F9FEC2D0F2" italics="true" pageId="196" pageNumber="197">Aclerda takahashii</emphasis>
Kuwana
</taxonomicName>
) is that of T. Kondo &amp; L.G Cook (unpublished data based on a Bayesian phylogenetic analysis of the family
<taxonomicName id="4C1B20F10AEB46962D7BF831FEBDD09E" box="[272,374,1983,2009]" class="Insecta" family="Coccidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">Coccidae</taxonomicName>
using DNA sequences based on 18S, 28S and partial CO1 data), who found that the
<taxonomicName id="4C1B20F10AEB4696294FF831FA56D09E" authorityName="Cockerell" authorityYear="1905" box="[1316,1437,1983,2009]" class="Insecta" family="Aclerdidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">Aclerdidae</taxonomicName>
fell within the
<taxonomicName id="4C1B20F10AEB46962D53F86DFE69D0BA" box="[312,418,2019,2045]" class="Insecta" family="Coccidae" kingdom="Animalia" order="Hemiptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="family">Coccidae</taxonomicName>
.
</paragraph>
<caption id="DF640BFA0AEA46972CFCF817FC09D092" ID-DOI="http://doi.org/10.5281/zenodo.3774332" ID-Zenodo-Dep="3774332" httpUri="https://zenodo.org/record/3774332/files/figure.png" pageId="197" pageNumber="198" startId="197.[151,250,1945,1970]" targetBox="[174,1412,192,1917]" targetPageId="197">
<paragraph id="8BA45B720AEA46972CFCF817FC09D092" blockId="197.[151,1437,1945,2006]" pageId="197" pageNumber="198">
<emphasis id="B96F87600AEA46972CFCF817FED4D0F5" bold="true" box="[151,287,1945,1970]" pageId="197" pageNumber="198">FIGURE 77.</emphasis>
<taxonomicName id="4C1B20F10AEA46972D4EF814FCE4D0F5" authority="Bytinski-Salz. Apterous" authorityName="Bytinski-Salz. Apterous" box="[293,815,1945,1970]" class="Insecta" family="Micrococcidae" genus="Micrococcus" kingdom="Animalia" order="Hemiptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="species" species="bodenheimeri">
<emphasis id="B96F87600AEA46972D4EF814FDFCD0F5" box="[293,567,1946,1970]" italics="true" pageId="197" pageNumber="198">Micrococcus bodenheimeri</emphasis>
Bytinski-Salz. Apterous
</taxonomicName>
adult male. (
<taxonomicName id="4C1B20F10AEA46972FDAF817FB80D0F5" authorityName="Silvestri" authorityYear="1939" box="[945,1099,1945,1970]" class="Insecta" family="Micrococcidae" kingdom="Animalia" order="Hemiptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="family">Micrococcidae</taxonomicName>
) (After
<bibRefCitation id="EF8A26830AEA469728F0F817FA5AD0F5" author="Miller, D. R. &amp; Williams, D. J." box="[1179,1425,1945,1970]" pageId="197" pageNumber="198" pagination="199 - 247" refId="ref209066" refString="Miller, D. R. &amp; Williams, D. J. (1995) Systematic revision of the Family Micrococcidae (Homoptera: Coccoidea), with a discussion of its relationships, and a description of a gynandromorph. Bollettino del Laboratorio di Entomologia Agraria ' Filippo Silvestri'. Portici, 50, 199 - 247. [1993]" type="journal article" year="1995">Miller &amp; Williams 1995</bibRefCitation>
). Where B=small bilocular pores, C=discoidal pores and D=large bilocular pores.
</paragraph>
</caption>
<paragraph id="8BA45B720AE946942CACFF17FE80D604" blockId="198.[151,1437,153,395]" pageId="198" pageNumber="199">
The only detailed descriptions of male
<taxonomicName id="4C1B20F10AE946942E19FF17FD22D7F4" authorityName="Cockerell" authorityYear="1905" box="[626,745,153,179]" class="Insecta" family="Aclerdidae" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="family">Aclerdidae</taxonomicName>
are those of
<bibRefCitation id="EF8A26830AE946942F13FF17FBF4D7F3" author="Nada" box="[888,1087,153,180]" etAl="et al." firstAuthor="Nada" pageId="198" pageNumber="199" pagination="133 - 140" refId="ref209825" refString="Nada, S. M. A., Afifi, S. A. &amp; Amin, A. H. (1976) Taxonomic status of family Aclerdidae according to the adult males (Homoptera: Coccoidea). Bulletin de la Societe Entomologique d'Egypte, 60, 133 - 140." type="journal article" year="1976">
Nada
<emphasis id="B96F87600AE946942FD2FF14FC23D7F4" box="[953,1000,153,179]" italics="true" pageId="198" pageNumber="199">et al</emphasis>
. (1976)
</bibRefCitation>
of
<taxonomicName id="4C1B20F10AE94694280FFF17FE3ED790" authority="McConnell, and Hodgson and Millar (2002)" authorityName="McConnell, and Hodgson and Millar" authorityYear="2002" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="arundinariae">
<emphasis id="B96F87600AE94694280FFF17FAD8D7F4" box="[1124,1299,153,179]" italics="true" pageId="198" pageNumber="199">A. arundinariae</emphasis>
McConnell, and
<bibRefCitation id="EF8A26830AE946942CACFF33FE3ED790" author="Hodgson and Millar" box="[199,501,189,215]" firstAuthor="Hodgson" pageId="198" pageNumber="199" pagination="469 - 517" refId="ref204361" refString="Hodgson, C. J. &amp; Millar, I. M. (2002) A new subfamily, two new genera and three new species of Aclerdidae (Hemiptera: Coccoidea) from southern Africa, with a phylogenetic analysis of relationships. Systematic Entomology, 27 (4), 469 - 517. https: // doi. org / 10.1046 / j. 1365 - 3113.2002.00189. x" type="journal article" year="2002">Hodgson and Millar (2002)</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C1B20F10AE946942E6DFF33FC3FD790" authority="Hodgson &amp; Millar" authorityName="Hodgson &amp; Millar. Macropterous" authorityYear="2002" box="[518,1012,189,215]" class="Insecta" family="Aclerdidae" genus="Lecanaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="macropoda">
<emphasis id="B96F87600AE946942E6DFF33FCD4D790" box="[518,799,189,215]" italics="true" pageId="198" pageNumber="199">Lecanaclerda macropoda</emphasis>
Hodgson &amp; Millar
</taxonomicName>
,
<taxonomicName id="4C1B20F10AE94694286BFF33FF09D7BC" authority="McConnell" authorityName="McConnell" authorityYear="1954" class="Insecta" family="Aclerdidae" genus="Rhodesaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="combreticola">
<emphasis id="B96F87600AE94694286BFF33FAF2D790" box="[1024,1337,189,215]" italics="true" pageId="198" pageNumber="199">Rhodesaclerda combreticola</emphasis>
McConnell
</taxonomicName>
,
<taxonomicName id="4C1B20F10AE946942CA6FF6CFDDED7BC" authority="Hodgson &amp; Millar" authorityName="Hodgson &amp; Millar. Macropterous" authorityYear="2002" box="[205,533,225,251]" class="Insecta" family="Aclerdidae" genus="Rhodesaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="insleyae">
<emphasis id="B96F87600AE946942CA6FF6CFE8FD7BC" box="[205,324,225,251]" italics="true" pageId="198" pageNumber="199">R. insleyae</emphasis>
Hodgson &amp; Millar
</taxonomicName>
and
<taxonomicName id="4C1B20F10AE946942E21FF6FFBEAD7BB" authority="Hodgson &amp; Millar" authorityName="Hodgson &amp; Millar" box="[586,1057,225,252]" class="Insecta" family="Aclerdidae" genus="Kwazulaclerda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="loranthi">
<emphasis id="B96F87600AE946942E21FF6FFC86D7BC" box="[586,845,225,251]" italics="true" pageId="198" pageNumber="199">Kwazulaclerda loranthi</emphasis>
Hodgson &amp; Millar
</taxonomicName>
), although McConnell does briefly describe those of
<taxonomicName id="4C1B20F10AE946942D3DFE8BFD0DD658" authority="McConnell" authorityName="McConnell, and Hodgson and Millar" authorityYear="2002" box="[342,710,261,287]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="arundinariae">
<emphasis id="B96F87600AE946942D3DFE8BFD89D658" box="[342,578,261,287]" italics="true" pageId="198" pageNumber="199">Aclerda arundinariae</emphasis>
McConnell
</taxonomicName>
,
<taxonomicName id="4C1B20F10AE946942EBBFE8BFC18D658" authority="McConnell" authorityName="McConnell" authorityYear="1954" box="[720,979,261,287]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="simplicis">
<emphasis id="B96F87600AE946942EBBFE8BFC84D658" box="[720,847,261,287]" italics="true" pageId="198" pageNumber="199">A. simplicis</emphasis>
McConnell
</taxonomicName>
,
<taxonomicName id="4C1B20F10AE946942FB5FE8BFAA5D658" authority="Kuwana" authorityName="Kuwana" box="[990,1390,261,287]" class="Insecta" family="Aclerdidae" genus="Nipponaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="biwakoensis">
<emphasis id="B96F87600AE946942FB5FE8BFAC6D658" box="[990,1293,261,287]" italics="true" pageId="198" pageNumber="199">Nipponaclerda biwakoensis</emphasis>
Kuwana
</taxonomicName>
and
<taxonomicName id="4C1B20F10AE946942CFCFEA4FE8FD604" authorityName="McConnell" authorityYear="1954" box="[151,324,297,323]" class="Insecta" family="Aclerdidae" genus="Rhodesaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="combreticola">
<emphasis id="B96F87600AE946942CFCFEA4FE8FD604" box="[151,324,297,323]" italics="true" pageId="198" pageNumber="199">R. combreticola</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA45B720AE946942CACFEC3FAE1D6CC" blockId="198.[151,1437,153,395]" pageId="198" pageNumber="199">
Below are described the adult males of 6 further species of
<taxonomicName id="4C1B20F10AE946942F34FEC3FC7FD620" authorityName="Signoret" authorityYear="1874" box="[863,948,333,359]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AE946942F34FEC3FC7FD620" box="[863,948,333,359]" italics="true" pageId="198" pageNumber="199">Aclerda</emphasis>
</taxonomicName>
. In addition, a figure of
<taxonomicName id="4C1B20F10AE9469428A3FEC3FE7ED6CC" authority="Hodgson &amp; Millar" authorityName="Hodgson &amp; Millar. Macropterous" authorityYear="2002" class="Insecta" family="Aclerdidae" genus="Lecanaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="macropoda">
<emphasis id="B96F87600AE9469428A3FEC3FF2ED6CC" italics="true" pageId="198" pageNumber="199">Lecanaclerda macropoda</emphasis>
Hodgson &amp; Millar
</taxonomicName>
is included (
<figureCitation id="132047F70AE946942E2AFEFFFD58D6CC" box="[577,659,369,395]" captionStart="FIGURE 84" captionStartId="216.[151,250,1945,1970]" captionTargetBox="[171,1420,189,1920]" captionTargetId="figure@216.[161,1426,181,1920]" captionTargetPageId="216" captionText="FIGURE 84. Lecanaclerda macropoda Hodgson &amp; Millar. Macropterous male. (Aclerdidae. Aclerdinae). Where A=cranial apophysis, D=simple pore, K=apex of antennal capitate seta and R=dermal reticulations on ocular sclerite." figureDoi="http://doi.org/10.5281/zenodo.3774346" httpUri="https://zenodo.org/record/3774346/files/figure.png" pageId="198" pageNumber="199">Fig. 84</figureCitation>
). This is followed by a brief review of the Rhodesaclerdinae.
</paragraph>
</subSubSection>
<subSubSection id="C30108F90AE946942CFCFE53FBABD6B0" box="[151,1120,477,503]" pageId="198" pageNumber="199" type="nomenclature">
<paragraph id="8BA45B720AE946942CFCFE53FBABD6B0" blockId="198.[151,1120,477,503]" box="[151,1120,477,503]" pageId="198" pageNumber="199">
<emphasis id="B96F87600AE946942CFCFE53FBABD6B0" bold="true" box="[151,1120,477,503]" pageId="198" pageNumber="199">
Key to subfamilies and genera of
<taxonomicName id="4C1B20F10AE946942E5CFE53FD0BD6B0" ID-CoL="5XP" ID-ENA="249417" authorityName="Cockerell" authorityYear="1905" box="[567,704,477,503]" class="Insecta" family="Aclerdidae" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="family">Aclerdidae</taxonomicName>
based on adult male morphology
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30108F90AE946942CFCFDACFA57D4B7" pageId="198" pageNumber="199" type="key">
<key id="FF8A0C430AE946942CFCFDACFA57D4B7" pageId="198" pageNumber="199">
<keyStep id="30EF43D70AE946942CFCFDACFA57D5C8" pageId="198" pageNumber="199">
<paragraph id="8BA45B720AE946942CFCFDACFA57D513" blockId="198.[151,1436,546,656]" pageId="198" pageNumber="199">
<keyLead id="30EAF8470AE946942CFCFDACFA57D513" pageId="198" pageNumber="199">
1. Antennae 1 or 10 segmented. Sclerotised ridges absent from around dorsal simple eyes. Abdomen with hs setae only. Claws without a denticle …
<taxonomicName id="4C1B20F10AE946942DDBFDB1FDD2D512" box="[432,537,575,597]" pageId="198" pageNumber="199">Aclerdinae</taxonomicName>
Cockerell................................................................. 2
</keyLead>
</paragraph>
<paragraph id="8BA45B720AE946942CFCFDD3FA57D5C8" blockId="198.[151,1436,546,656]" pageId="198" pageNumber="199">
<keyLead id="30EAF8470AE946942CFCFDD3FA57D5C8" pageId="198" pageNumber="199">- Antennae 7 or 8 segmented. Sclerotised ridges present around dorsal simple eyes. Abdomen with both hs and fs setae. Claws with a denticle … Rhodesaclerdinae Hodgson &amp; Millar....................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF43D70AE946942CFCFD3AFA57D443" pageId="198" pageNumber="199">
<paragraph id="8BA45B720AE946942CFCFD3AFA57D5A0" blockId="198.[151,1436,692,773]" pageId="198" pageNumber="199">
<keyLead id="30EAF8470AE946942CFCFD3AFA57D5A0" pageId="198" pageNumber="199">
2. Adult male apterous. Legs extremely reduced to 4 ring-like segments. Penial sheath very short, only slightly longer than broad..................................................................
<taxonomicName id="4C1B20F10AE94694282AFD5CFA89D5AF" ID-CoL="64B7" authority="McConnell" authorityName="McConnell" authorityYear="1954" box="[1089,1346,722,744]" class="Insecta" family="Aclerdidae" genus="Nipponaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AE94694282AFD5CFB1BD5A0" box="[1089,1232,722,743]" italics="true" pageId="198" pageNumber="199">Nipponaclerda</emphasis>
McConnell
</taxonomicName>
(
<figureCitation id="132047F70AE946942924FD5CFA5ED5A0" box="[1359,1429,721,744]" captionStart="FIGURE 85" captionStartId="218.[151,250,1068,1093]" captionTargetBox="[369,1219,185,1038]" captionTargetId="figure@218.[363,1223,181,1042]" captionTargetPageId="218" captionText="FIGURE 85. Nipponaclerda biwakoensis (Kuwana). Apterous male. (Aclerdidae, Aclerdina) (after McConnell, 1954). Where H=antenna, I=anterior leg, J=posterior spiracle, K=metathoracic leg, and L=ventral view of penial sheath." figureDoi="http://doi.org/10.5281/zenodo.3774348" httpUri="https://zenodo.org/record/3774348/files/figure.png" pageId="198" pageNumber="199">Fig. 85</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BA45B720AE946942CFCFD61FA57D443" blockId="198.[151,1436,692,773]" box="[151,1436,751,773]" pageId="198" pageNumber="199">
<keyLead id="30EAF8470AE946942CFCFD61FA57D443" box="[151,1436,751,773]" pageId="198" pageNumber="199">- Adult male alate. Legs normally developed with 5 long segments. Penial sheath several times longer than basal width..... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF43D70AE946942CFCFCA4FA57D4DF" pageId="198" pageNumber="199">
<paragraph id="8BA45B720AE946942CFCFCA4FA57D41A" blockId="198.[151,1436,810,920]" pageId="198" pageNumber="199">
<keyLead id="30EAF8470AE946942CFCFCA4FA57D41A" pageId="198" pageNumber="199">
3. Caudal extension on abdominal segment VIII well developed. Pleural setae absent from abdominal segment VIII. Penial sheath not articulating with abdominal segment VIII..............................
<taxonomicName id="4C1B20F10AE946942863FCC9FA89D41A" ID-CoL="5BMT" authority="Hodgson &amp; Millar" authorityName="Hodgson &amp; Millar" authorityYear="2002" box="[1032,1346,839,861]" class="Insecta" family="Aclerdidae" genus="Lecanaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AE946942863FCC9FB40D41B" box="[1032,1163,839,860]" italics="true" pageId="198" pageNumber="199">Lecanaclerda</emphasis>
Hodgson &amp; Millar
</taxonomicName>
(
<figureCitation id="132047F70AE946942924FCC9FA5ED41A" box="[1359,1429,839,861]" captionStart="FIGURE 84" captionStartId="216.[151,250,1945,1970]" captionTargetBox="[171,1420,189,1920]" captionTargetId="figure@216.[161,1426,181,1920]" captionTargetPageId="216" captionText="FIGURE 84. Lecanaclerda macropoda Hodgson &amp; Millar. Macropterous male. (Aclerdidae. Aclerdinae). Where A=cranial apophysis, D=simple pore, K=apex of antennal capitate seta and R=dermal reticulations on ocular sclerite." figureDoi="http://doi.org/10.5281/zenodo.3774346" httpUri="https://zenodo.org/record/3774346/files/figure.png" pageId="198" pageNumber="199">Fig. 84</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BA45B720AE946942CFCFCEBFA57D4DF" blockId="198.[151,1436,810,920]" pageId="198" pageNumber="199">
<keyLead id="30EAF8470AE946942CFCFCEBFA57D4DF" pageId="198" pageNumber="199">
- Caudal extension on abdominal segment VIII at most a gentle curve. Pleural setae present on abdominal segment VIII. Penial sheath clearly articulating with abdominal segment VIII..........................................
<taxonomicName id="4C1B20F10AE946942891FC0CFA57D4DF" ID-CoL="MZX" authority="Signoret" authorityName="Signoret" authorityYear="1874" box="[1274,1436,898,920]" class="Insecta" family="Aclerdidae" genus="Aclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AE946942891FC0CFA8ED4D0" box="[1274,1349,898,919]" italics="true" pageId="198" pageNumber="199">Aclerda</emphasis>
Signoret
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BA45B720AE946942CFCFC32FA57D4B7" blockId="198.[151,1436,956,1008]" pageId="198" pageNumber="199">
<keyStep id="30EF43D70AE946942CFCFC32FA57D4B7" pageId="198" pageNumber="199">
<keyLead id="30EAF8470AE946942CFCFC32FA57D4B7" pageId="198" pageNumber="199">
4. Antennae 7 segmented. Lateropleurite broad. Membranous area on scutum quadrate....
<taxonomicName id="4C1B20F10AE94694282AFC32FA89D495" ID-CoL="793C" authority="McConnell" authorityName="McConnell" authorityYear="1954" box="[1089,1346,956,978]" class="Insecta" family="Aclerdidae" genus="Rhodesaclerda" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AE94694282AFC32FB1BD496" box="[1089,1232,956,977]" italics="true" pageId="198" pageNumber="199">Rhodesaclerda</emphasis>
McConnell
</taxonomicName>
(
<figureCitation id="132047F70AE946942924FC32FA5ED495" box="[1359,1429,956,978]" captionStart="FIGURE 86" captionStartId="219.[151,250,1846,1871]" captionTargetBox="[160,1423,191,1821]" captionTargetId="figure@219.[151,1436,181,1821]" captionTargetPageId="219" captionText="FIGURE 86. Rhodesaclerda insleyae Hodgson &amp; Millar. Macropterous male. (Aclerdidae, Rhodesaclerdinae).Where D=simple pore and R=dermal reticulations on ocular sclerite." figureDoi="http://doi.org/10.5281/zenodo.3774350" httpUri="https://zenodo.org/record/3774350/files/figure.png" pageId="198" pageNumber="199">Fig. 86</figureCitation>
) - Antennae 8 segmented. Lateropleurite narrow. Membranous area on scutum triangular...
<taxonomicName id="4C1B20F10AE94694283FFC54FA57D4B7" authority="Hodgson &amp; Millar" authorityName="Hodgson &amp; Millar" authorityYear="2002" box="[1108,1436,986,1008]" class="Insecta" family="Aclerdidae" genus="Kwazulaclerda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B96F87600AE94694283FFC54FB2FD4A8" box="[1108,1252,986,1007]" italics="true" pageId="198" pageNumber="199">Kwazulaclerda</emphasis>
Hodgson &amp; Millar
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
</key>
</subSubSection>
</treatment>
</document>
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