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<document id="C5529B9CBB9EF6ED21E0137C7DC3EC04" ID="10.11646/zootaxa.4169.2.1" ID-DOI="10.11646/zootaxa.4169.2.1" ID-GBIF-Dataset="d1fbe4b3-de10-4d72-a117-aa6bfedf210d" ID-ISSN="1175-5326" ID-Zenodo-Dep="262585" ID-ZooBank="B8B067BF-F6E6-4122-B884-AA385FF04421" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474351831659" checkinUser="plazi" docAuthor="Andrew Hamilton, K. G." docDate="2016" docId="03A5742DFFC6AE19FF76ADD4C8EBFA7B" docLanguage="en" docName="zootaxa.4169.2.1.pdf" docOrigin="Zootaxa 4169 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neaenus" docType="key" docVersion="7" lastPageNumber="220" masterDocId="FF9C0C55FFD5AE0AFFE1A955CD70FFD1" masterDocTitle="Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae" masterLastPageNumber="250" masterPageNumber="201" pageNumber="220" updateTime="1698386375821" updateUser="plazi">
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<mods:title id="D4E2E89F899D4F30EE5837181A0F67AC">Neotropical spittlebugs related to Neaenini (Hemiptera, Cercopidae) and the origins of subfamily Cercopinae</mods:title>
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<mods:namePart id="D57FDB28D8B53645C19D3A93AF0791F2">Andrew Hamilton, K. G.</mods:namePart>
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<mods:date id="A0B46AC84C25E11D39F9E6590ABC41FB">2016</mods:date>
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<heading id="D0FB7257FFC6AE19FF76ADD4CCA6FB4A" bold="true" box="[151,470,1153,1179]" fontSize="11" level="1" pageId="19" pageNumber="220" reason="1">
<emphasis id="B9781929FFC6AE19FF76ADD4CCA6FB4A" bold="true" box="[151,470,1153,1179]" pageId="19" pageNumber="220">
Key to species of
<taxonomicName id="4C0CBEB8FFC6AE19FE8CADD4CCA6FB4A" box="[365,470,1153,1179]" class="Insecta" family="Aphrophoridae" genus="Neaenus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="genus">
<emphasis id="B9781929FFC6AE19FE8CADD4CCA6FB4A" bold="true" box="[365,470,1153,1179]" italics="true" pageId="19" pageNumber="220">Neaenus</emphasis>
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<subSubSection id="C31696B0FFC6AE19FF76AD92C8EBFA7B" pageId="19" pageNumber="220" type="key">
<keyStep id="30F8DD9EFFC6AE19FF76AD92C8ECFB2B" pageId="19" pageNumber="220">
<paragraph id="8BB3C53BFFC6AE19FF76AD92C8EBFB0D" blockId="19.[151,1436,1223,1450]" box="[151,1435,1223,1245]" pageId="19" pageNumber="220">
<keyLead id="30FD660EFFC6AE19FF76AD92C8EBFB0D" box="[151,1435,1223,1245]" pageId="19" pageNumber="220">
1. Venation evident; tegmina mottled (
<figureCitation id="1337D9BEFFC6AE19FDD9AD92CFF1FB0D" box="[568,641,1223,1244]" captionStart="FIGURES 10 11" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[199,1406,299,1890]" captionTargetId="figure@20.[199,1406,281,1890]" captionTargetPageId="20" captionText="FIGURES 10 11. Neaenus Fowler, dorsolateral aspect (10) and female genitalia, ventral aspect (11). 10 A, N. varius Fowler; 10 B, N. hystricosus sp. nov.; 10 C, undescribed female from 15 mi east of El Palmito in Mexico; 10 D, N. natrix sp. nov.; 10 E F, 11 A, N. (Neaniskus) gilli sp. nov.; 10 G H, 11 B, N. (Neaniskus) zacki sp. nov.." httpUri="https://zenodo.org/record/262588/files/figure.png" pageId="19" pageNumber="220">Figs 10</figureCitation>
EH); length&gt;
<quantity id="4CF468DEFFC6AE19FCF3AD92CE38FB0D" box="[786,840,1223,1244]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="220" unit="mm" value="8.0">8 mm</quantity>
............................
<emphasis id="B9781929FFC6AE19FB48AD92C87CFB0D" box="[1193,1292,1223,1244]" italics="true" pageId="19" pageNumber="220">Neaniscus</emphasis>
<emphasis id="B9781929FFC6AE19FAF3AD92C807FB0C" bold="true" box="[1298,1399,1223,1245]" pageId="19" pageNumber="220">subg. nov.</emphasis>
...4
</keyLead>
</paragraph>
<paragraph id="8BB3C53BFFC6AE19FF76ADB0C8ECFB2B" blockId="19.[151,1436,1223,1450]" box="[151,1436,1253,1274]" pageId="19" pageNumber="220">
<keyLead id="30FD660EFFC6AE19FF76ADB0C8ECFB2B" box="[151,1436,1253,1274]" pageId="19" pageNumber="220">
- Venation obscure; tegmina with patches of color, or evenly colored (
<figureCitation id="1337D9BEFFC6AE19FC8EADB0CEC8FB2B" box="[879,952,1253,1274]" captionStart="FIGURES 10 11" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[199,1406,299,1890]" captionTargetId="figure@20.[199,1406,281,1890]" captionTargetPageId="20" captionText="FIGURES 10 11. Neaenus Fowler, dorsolateral aspect (10) and female genitalia, ventral aspect (11). 10 A, N. varius Fowler; 10 B, N. hystricosus sp. nov.; 10 C, undescribed female from 15 mi east of El Palmito in Mexico; 10 D, N. natrix sp. nov.; 10 E F, 11 A, N. (Neaniskus) gilli sp. nov.; 10 G H, 11 B, N. (Neaniskus) zacki sp. nov.." httpUri="https://zenodo.org/record/262588/files/figure.png" pageId="19" pageNumber="220">Figs 10</figureCitation>
AD); length &lt;
<quantity id="4CF468DEFFC6AE19FBAAADB0C9F2FB2B" box="[1099,1154,1253,1274]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="220" unit="mm" value="7.0">7 mm</quantity>
..... nominate subgenus...2
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<paragraph id="8BB3C53BFFC6AE19FF76AC57C8ECFAC6" blockId="19.[151,1436,1223,1450]" box="[151,1436,1282,1303]" pageId="19" pageNumber="220">
<keyLead id="30FD660EFFC6AE19FF76AC57C8ECFAC6" box="[151,1436,1282,1303]" pageId="19" pageNumber="220">
2. Tegmina usually with pale spots beyond base; aedeagus with paired apical processes......................
<taxonomicName id="4C0CBEB8FFC6AE19FAF1AC57C8ECFAC6" authority="Fowler" authorityName="Fowler" box="[1296,1436,1282,1303]" class="Insecta" family="Aphrophoridae" genus="Neaenus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="varius">
<emphasis id="B9781929FFC6AE19FAF1AC57C821FAC6" box="[1296,1361,1282,1303]" italics="true" pageId="19" pageNumber="220">varius</emphasis>
Fowler
</taxonomicName>
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</paragraph>
<paragraph id="8BB3C53BFFC6AE19FF76AC4AC8EBFAE5" blockId="19.[151,1436,1223,1450]" box="[151,1435,1311,1332]" pageId="19" pageNumber="220">
<keyLead id="30FD660EFFC6AE19FF76AC4AC8EBFAE5" box="[151,1435,1311,1332]" pageId="19" pageNumber="220">- Tegmina unmarked or with pale base; aedeagus without paired processes......................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F8DD9EFFC6AE19FF76AC68C8EBFAA1" pageId="19" pageNumber="220">
<paragraph id="8BB3C53BFFC6AE19FF76AC68C8EBFA83" blockId="19.[151,1436,1223,1450]" box="[151,1435,1340,1362]" pageId="19" pageNumber="220">
<keyLead id="30FD660EFFC6AE19FF76AC68C8EBFA83" box="[151,1435,1340,1362]" pageId="19" pageNumber="220">
3. Male subgenital plates tapered; style serpentine (
<figureCitation id="1337D9BEFFC6AE19FD55AC68CF8AFA83" box="[692,762,1341,1362]" captionStart="FIGURES 46 54" captionStartId="40.[151,264,1868,1890]" captionTargetBox="[167,1435,197,1840]" captionTargetId="figure@40.[167,1452,193,1848]" captionTargetPageId="40" captionText="FIGURES 46 54. Male genitalia of nominate genus of Neaenini (46 49) and nominate subgenus of Microsarganini (50 54): 46, Neaenus natrix; 47, N. hystricosus; 48, N. (Neaniskus) gilli; 49, Helioptera gilvum; 50, Microsargane vittata; 51, M. apicata; 52, M. ornatipennis; 53, M. mutans; 54, M. cristata. A, lateral aspect of genital capsule and style; B, same, of theca; C, same in caudal aspect (with detail in Microsargane showing tips of processes in widest aspect); D, ventral aspect of subgenital plates, smaller scale; E, style, dorsal aspect." httpUri="https://zenodo.org/record/262597/files/figure.png" pageId="19" pageNumber="220">Fig. 46</figureCitation>
A)...........................................
<taxonomicName id="4C0CBEB8FFC6AE19FAF1AC68C83EFA83" box="[1296,1358,1341,1362]" class="Insecta" family="Aphrophoridae" genus="Neaenus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="natrix" status="sp. nov.">
<emphasis id="B9781929FFC6AE19FAF1AC68C83EFA83" box="[1296,1358,1341,1362]" italics="true" pageId="19" pageNumber="220">natrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA452FFC6AE19FAB4AC69C8EBFA83" box="[1365,1435,1340,1362]" pageId="19" pageNumber="220" rank="species">
<emphasis id="B9781929FFC6AE19FAB4AC69C8EBFA83" bold="true" box="[1365,1435,1340,1362]" pageId="19" pageNumber="220">sp.nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BB3C53BFFC6AE19FF76AC0FC8EBFAA1" blockId="19.[151,1436,1223,1450]" box="[151,1435,1370,1392]" pageId="19" pageNumber="220">
<keyLead id="30FD660EFFC6AE19FF76AC0FC8EBFAA1" box="[151,1435,1370,1392]" pageId="19" pageNumber="220">
- Male subgenital plates with prominent lobe near base; style straight, apically furcate (
<figureCitation id="1337D9BEFFC6AE19FBF7AC0FC92BFABE" box="[1046,1115,1370,1391]" captionStart="FIGURES 46 54" captionStartId="40.[151,264,1868,1890]" captionTargetBox="[167,1435,197,1840]" captionTargetId="figure@40.[167,1452,193,1848]" captionTargetPageId="40" captionText="FIGURES 46 54. Male genitalia of nominate genus of Neaenini (46 49) and nominate subgenus of Microsarganini (50 54): 46, Neaenus natrix; 47, N. hystricosus; 48, N. (Neaniskus) gilli; 49, Helioptera gilvum; 50, Microsargane vittata; 51, M. apicata; 52, M. ornatipennis; 53, M. mutans; 54, M. cristata. A, lateral aspect of genital capsule and style; B, same, of theca; C, same in caudal aspect (with detail in Microsargane showing tips of processes in widest aspect); D, ventral aspect of subgenital plates, smaller scale; E, style, dorsal aspect." httpUri="https://zenodo.org/record/262597/files/figure.png" pageId="19" pageNumber="220">Fig. 47</figureCitation>
A)........
<taxonomicName id="4C0CBEB8FFC6AE19FB02AC0FC83FFABE" box="[1251,1359,1370,1391]" class="Insecta" family="Aphrophoridae" genus="Neaenus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="hystricosus" status="sp. nov.">
<emphasis id="B9781929FFC6AE19FB02AC0FC83FFABE" box="[1251,1359,1370,1391]" italics="true" pageId="19" pageNumber="220">hystricosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA452FFC6AE19FAB4AC0FC8EBFAA1" box="[1365,1435,1370,1392]" pageId="19" pageNumber="220" rank="species">
<emphasis id="B9781929FFC6AE19FAB4AC0FC8EBFAA1" bold="true" box="[1365,1435,1370,1392]" pageId="19" pageNumber="220">sp.nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F8DD9EFFC6AE19FF76AC22C8EBFA7B" pageId="19" pageNumber="220">
<paragraph id="8BB3C53BFFC6AE19FF76AC22C8EBFA5C" blockId="19.[151,1436,1223,1450]" box="[151,1435,1399,1421]" pageId="19" pageNumber="220">
<keyLead id="30FD660EFFC6AE19FF76AC22C8EBFA5C" box="[151,1435,1399,1421]" pageId="19" pageNumber="220">
4. Tegmina striped in brown and yellow (
<figureCitation id="1337D9BEFFC6AE19FDB5AC22CFE9FA5D" box="[596,665,1399,1420]" captionStart="FIGURES 10 11" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[199,1406,299,1890]" captionTargetId="figure@20.[199,1406,281,1890]" captionTargetPageId="20" captionText="FIGURES 10 11. Neaenus Fowler, dorsolateral aspect (10) and female genitalia, ventral aspect (11). 10 A, N. varius Fowler; 10 B, N. hystricosus sp. nov.; 10 C, undescribed female from 15 mi east of El Palmito in Mexico; 10 D, N. natrix sp. nov.; 10 E F, 11 A, N. (Neaniskus) gilli sp. nov.; 10 G H, 11 B, N. (Neaniskus) zacki sp. nov.." httpUri="https://zenodo.org/record/262588/files/figure.png" pageId="19" pageNumber="220">Fig. 10</figureCitation>
EF); female with base of ovipositor exposed (
<figureCitation id="1337D9BEFFC6AE19FBBAAC22C9EDFA5D" box="[1115,1181,1399,1420]" captionStart="FIGURES 10 11" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[199,1406,299,1890]" captionTargetId="figure@20.[199,1406,281,1890]" captionTargetPageId="20" captionText="FIGURES 10 11. Neaenus Fowler, dorsolateral aspect (10) and female genitalia, ventral aspect (11). 10 A, N. varius Fowler; 10 B, N. hystricosus sp. nov.; 10 C, undescribed female from 15 mi east of El Palmito in Mexico; 10 D, N. natrix sp. nov.; 10 E F, 11 A, N. (Neaniskus) gilli sp. nov.; 10 G H, 11 B, N. (Neaniskus) zacki sp. nov.." httpUri="https://zenodo.org/record/262588/files/figure.png" pageId="19" pageNumber="220">Fig. 11</figureCitation>
A)........
<taxonomicName id="4C0CBEB8FFC6AE19FAC9AC22C83FFA5D" box="[1320,1359,1399,1420]" class="Insecta" family="Aphrophoridae" genus="Neaenus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="gilli" status="sp. nov.">
<emphasis id="B9781929FFC6AE19FAC9AC22C83FFA5D" box="[1320,1359,1399,1420]" italics="true" pageId="19" pageNumber="220">gilli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA452FFC6AE19FAB4AC22C8EBFA5C" box="[1365,1435,1399,1421]" pageId="19" pageNumber="220" rank="species">
<emphasis id="B9781929FFC6AE19FAB4AC22C8EBFA5C" bold="true" box="[1365,1435,1399,1421]" pageId="19" pageNumber="220">sp.nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BB3C53BFFC6AE19FF76ACC0C8EBFA7B" blockId="19.[151,1436,1223,1450]" box="[151,1435,1428,1450]" pageId="19" pageNumber="220">
<keyLead id="30FD660EFFC6AE19FF76ACC0C8EBFA7B" box="[151,1435,1428,1450]" pageId="19" pageNumber="220">
- Tegmina mottled (
<figureCitation id="1337D9BEFFC6AE19FE71ACC0CCA9FA7B" box="[400,473,1429,1450]" captionStart="FIGURES 10 11" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[199,1406,299,1890]" captionTargetId="figure@20.[199,1406,281,1890]" captionTargetPageId="20" captionText="FIGURES 10 11. Neaenus Fowler, dorsolateral aspect (10) and female genitalia, ventral aspect (11). 10 A, N. varius Fowler; 10 B, N. hystricosus sp. nov.; 10 C, undescribed female from 15 mi east of El Palmito in Mexico; 10 D, N. natrix sp. nov.; 10 E F, 11 A, N. (Neaniskus) gilli sp. nov.; 10 G H, 11 B, N. (Neaniskus) zacki sp. nov.." httpUri="https://zenodo.org/record/262588/files/figure.png" pageId="19" pageNumber="220">Figs 10</figureCitation>
GH); female with base of ovipositor covered by shiny black knob (
<figureCitation id="1337D9BEFFC6AE19FB87ACC0C9DAFA7B" box="[1126,1194,1429,1450]" captionStart="FIGURES 10 11" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[199,1406,299,1890]" captionTargetId="figure@20.[199,1406,281,1890]" captionTargetPageId="20" captionText="FIGURES 10 11. Neaenus Fowler, dorsolateral aspect (10) and female genitalia, ventral aspect (11). 10 A, N. varius Fowler; 10 B, N. hystricosus sp. nov.; 10 C, undescribed female from 15 mi east of El Palmito in Mexico; 10 D, N. natrix sp. nov.; 10 E F, 11 A, N. (Neaniskus) gilli sp. nov.; 10 G H, 11 B, N. (Neaniskus) zacki sp. nov.." httpUri="https://zenodo.org/record/262588/files/figure.png" pageId="19" pageNumber="220">Fig. 11</figureCitation>
B)......
<taxonomicName id="4C0CBEB8FFC6AE19FAFFACC0C83FFA7B" box="[1310,1359,1429,1450]" class="Insecta" family="Aphrophoridae" genus="Neaenus" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="zacki" status="sp. nov.">
<emphasis id="B9781929FFC6AE19FAFFACC0C83FFA7B" box="[1310,1359,1429,1450]" italics="true" pageId="19" pageNumber="220">zacki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA452FFC6AE19FAB4ACC1C8EBFA7B" box="[1365,1435,1428,1450]" pageId="19" pageNumber="220" rank="species">
<emphasis id="B9781929FFC6AE19FAB4ACC1C8EBFA7B" bold="true" box="[1365,1435,1428,1450]" pageId="19" pageNumber="220">sp.nov.</emphasis>
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