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<document id="BC3E766A19CBBF5A18752DCE933B753F" ID-CLB-Dataset="30081" ID-DOI="10.11646/zootaxa.4420.1.1" ID-GBIF-Dataset="a3ea0a39-5d5d-4f18-a4b2-4c3aa667f06f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1247134" ID-ZooBank="70B25E60-2637-4D35-8837-14A0796D82B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1526382456942" checkinUser="plazi" docAuthor="Alipanah, Helen, Khodadad, Masoumeh, Rajaei, Hossein &amp; Haseli, Mohammad" docDate="2018" docId="039F87FDFFB30978FF6AFB9AFCE6F84B" docLanguage="en" docName="zootaxa.4420.1.1.pdf.imf" docOrigin="Zootaxa 4420 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Evergestis boursini Amsel 1939" docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="FFA6FF85FFB5097EFFFDFFBBFF9BFFD2" masterDocTitle="Taxonomic study of the genus Evergestis Hübner, 1825 (Lepidoptera: Crambidae: Glaphyriinae) in Iran with description of a new species" masterLastPageNumber="33" masterPageNumber="1" pageNumber="7" updateTime="1698544382926" updateUser="ExternalLinkService">
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<mods:title id="1A6A91A20E5F06427F509590A8FCF55E">Taxonomic study of the genus Evergestis Hübner, 1825 (Lepidoptera: Crambidae: Glaphyriinae) in Iran with description of a new species</mods:title>
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<mods:namePart id="DE50D4AD1269815B88A2948B1F24F6CA">Alipanah, Helen</mods:namePart>
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<mods:namePart id="658A9DE2F7A771D14400AFD2508555BB">Khodadad, Masoumeh</mods:namePart>
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<mods:namePart id="43C1280D20D9EE160BB9B3E57F3A3DD4">Rajaei, Hossein</mods:namePart>
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<mods:namePart id="A4C06BFB0D978677D87AA7B612A8AC4F">Haseli, Mohammad</mods:namePart>
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<treatment id="039F87FDFFB30978FF6AFB9AFCE6F84B" ID-DOI="http://doi.org/10.5281/zenodo.5966952" ID-GBIF-Taxon="143970632" ID-Zenodo-Dep="5966952" LSID="urn:lsid:plazi:treatment:039F87FDFFB30978FF6AFB9AFCE6F84B" httpUri="http://treatment.plazi.org/id/039F87FDFFB30978FF6AFB9AFCE6F84B" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection id="C32C6560FFB30978FF6AFB9AFD85FBE9" box="[151,542,1057,1083]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8B8936EBFFB30978FF6AFB9AFD85FBE9" blockId="6.[151,542,1057,1083]" box="[151,542,1057,1083]" pageId="6" pageNumber="7">
<heading id="D0C18187FFB30978FF6AFB9AFD85FBE9" bold="true" box="[151,542,1057,1083]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="4C364D68FFB30978FF6AFB9AFD85FBE9" ID-CoL="3DHBM" authority="Amsel, 1939" authorityName="Amsel" authorityYear="1939" box="[151,542,1057,1083]" class="Insecta" family="Crambidae" genus="Evergestis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="boursini">
<emphasis id="B942EAF9FFB30978FF6AFB9AFD85FBE9" bold="true" box="[151,542,1057,1083]" pageId="6" pageNumber="7">
<emphasis id="B942EAF9FFB30978FF6AFB9AFEE4FBE9" bold="true" box="[151,383,1057,1083]" italics="true" pageId="6" pageNumber="7">Evergestis boursini</emphasis>
<bibRefCitation id="EFA74B1AFFB30978FE7BFB9AFD85FBE9" box="[390,542,1057,1083]" pageId="6" pageNumber="7" refId="ref18931" refString="Amsel, H. G. (1939) Beitrag zur Kleinschmetterlings-Fauna der Balkanlander und Kleinasiens. Deutsche entomologische Zeitschrift Iris, Dresden, 52 (1938) (3 / 4), 152 - 159, pl. 4." type="journal article">Amsel, 1939</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32C6560FFB30978FF6AFBD3FE04FB77" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8B8936EBFFB30978FF6AFBD3FE04FB77" blockId="6.[151,1437,1128,1945]" pageId="6" pageNumber="7">
<emphasis id="B942EAF9FFB30978FF6AFBD3FE7EFB53" bold="true" box="[151,485,1128,1153]" pageId="6" pageNumber="7">
Material examined.
<collectionCode id="ED27AE2EFFB30978FE75FBD3FE7BFB53" box="[392,480,1128,1153]" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" pageId="6" pageNumber="7">SMNK</collectionCode>
:
</emphasis>
<typeStatus id="548D8849FFB30978FE10FBD2FDC9FB53" box="[493,594,1129,1153]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
,
<specimenCount id="9D30FD62FFB30978FDA3FBD2FDEEFB52" box="[606,629,1129,1152]" pageId="6" pageNumber="7" type="male"></specimenCount>
, Asia min.,
<collectingRegion id="49F2F809FFB30978FD03FBD3FCA6FB53" box="[766,829,1128,1153]" country="Turkey" name="Sivas" pageId="6" pageNumber="7">Sivas</collectingRegion>
, Zukowsky,
<date id="FF88102BFFB30978FC31FBD3FBDDFB53" box="[972,1094,1128,1153]" pageId="6" pageNumber="7" value="1936-09-29">29.ix.1936</date>
, leg. H. Amsel (ex. coll. H. G. Amsel) (GS: HA-2047)
</paragraph>
</subSubSection>
<subSubSection id="C32C6560FFB30978FF3AFB0BFD06FA73" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8B8936EBFFB30978FF3AFB0BFD06FA73" blockId="6.[151,1437,1128,1945]" pageId="6" pageNumber="7">
<emphasis id="B942EAF9FFB30978FF3AFB0BFEA0FB1B" bold="true" box="[199,315,1200,1225]" pageId="6" pageNumber="7">Remarks.</emphasis>
This species was described by
<bibRefCitation id="EFA74B1AFFB30978FD5AFB0BFCDFFB1A" box="[679,836,1200,1225]" pageId="6" pageNumber="7" refId="ref18931" refString="Amsel, H. G. (1939) Beitrag zur Kleinschmetterlings-Fauna der Balkanlander und Kleinasiens. Deutsche entomologische Zeitschrift Iris, Dresden, 52 (1938) (3 / 4), 152 - 159, pl. 4." type="journal article">Amsel (1939)</bibRefCitation>
based on three male specimens collected in
<collectingCountry id="F321767BFFB30978FABAFB0BFA03FB1B" box="[1351,1432,1200,1225]" name="Turkey" pageId="6" pageNumber="7">Turkey</collectingCountry>
,
<collectingRegion id="49F2F809FFB30978FF6AFB6FFF4FFB3F" box="[151,212,1236,1261]" country="Turkey" name="Sivas" pageId="6" pageNumber="7">Sivas</collectingRegion>
(two males including the
<typeStatus id="548D8849FFB30978FDFCFB6FFDF2FB3F" box="[513,617,1236,1261]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
in
<date id="FF88102BFFB30978FD6CFB6FFD68FB3F" box="[657,755,1236,1261]" pageId="6" pageNumber="7" value="1936-10-04">4.x.1936</date>
and one male in
<date id="FF88102BFFB30978FC41FB6FFBA3FB3F" box="[956,1080,1236,1261]" pageId="6" pageNumber="7" value="1936-09-29">29.ix.1936</date>
).
<bibRefCitation id="EFA74B1AFFB30978FBB0FB6FFB71FB3F" box="[1101,1258,1236,1261]" pageId="6" pageNumber="7" refId="ref19135" refString="Amsel, H. G. (1961) Die Microlepidopteren der Brandt'schen Iran-Ausbeute. 5. Teil. Arkiv for Zoologie (N. S.), Stockholm (ser. 2) 13 (17), 323 - 445, pls. 1 - 9." type="journal article">Amsel (1961)</bibRefCitation>
later recoreded this species from
<collectingCountry id="F321767BFFB30978FE9DFB42FE16FAC2" box="[352,397,1273,1296]" name="Iran" pageId="6" pageNumber="7">Iran</collectingCountry>
based on eight males and one female collected in Karaj (Nesa) in
<date id="FF88102BFFB30978FB7AFB42FA80FAC3" box="[1159,1307,1272,1297]" pageId="6" pageNumber="7" value="1936-08">August 1936</date>
. In spite of collecting a female specimen, no description for female was presented by
<bibRefCitation id="EFA74B1AFFB30978FC20FAA7FBE7FAE7" box="[989,1148,1308,1333]" pageId="6" pageNumber="7" refId="ref19135" refString="Amsel, H. G. (1961) Die Microlepidopteren der Brandt'schen Iran-Ausbeute. 5. Teil. Arkiv for Zoologie (N. S.), Stockholm (ser. 2) 13 (17), 323 - 445, pls. 1 - 9." type="journal article">Amsel (1961)</bibRefCitation>
. None of the nine Iranian specimens collected by Amsel were available to study, but one male
<typeStatus id="548D8849FFB30978FC66FAFAFC66FA8B" box="[923,1021,1345,1369]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
(see material examined) was studied and dissected (
<figureCitation id="130D2A6EFFB30978FEC1FADFFE0CFAAF" box="[316,407,1380,1405]" captionStart="FIGURE 4" captionStartId="7.[151,250,1532,1554]" captionTargetBox="[170,1421,204,1497]" captionTargetId="figure@7.[151,1436,193,1510]" captionTargetPageId="7" captionText="FIGURE 4. Evergestis boursini, Paratype (♂). A) Adult male, upperside. BF) Male genitalia. B: main body and phallus. C: posterior end including uncus and gnathos in ventral view. D: posterior end of uncus in lateral view. E: posterior end of gnathos in laterl view, F: distal end of phallus." httpUri="https://zenodo.org/record/1247142/files/figure.png" pageId="6" pageNumber="7">Fig. 4A</figureCitation>
). Due to the paucity of genital information in the description provided by
<bibRefCitation id="EFA74B1AFFB30978FB2CFADFFAF5FAAF" box="[1233,1390,1380,1405]" pageId="6" pageNumber="7" refId="ref18931" refString="Amsel, H. G. (1939) Beitrag zur Kleinschmetterlings-Fauna der Balkanlander und Kleinasiens. Deutsche entomologische Zeitschrift Iris, Dresden, 52 (1938) (3 / 4), 152 - 159, pl. 4." type="journal article">Amsel (1939)</bibRefCitation>
, the male genitalia is here redescribed and figured.
</paragraph>
</subSubSection>
<subSubSection id="C32C6560FFB30978FF3AFA17FD4DF987" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8B8936EBFFB30978FF3AFA17FD4DF987" blockId="6.[151,1437,1128,1945]" pageId="6" pageNumber="7">
<emphasis id="B942EAF9FFB30978FF3AFA17FEDAFA17" bold="true" box="[199,321,1452,1477]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
According to
<bibRefCitation id="EFA74B1AFFB30978FE1DFA17FDE7FA17" box="[480,636,1452,1477]" pageId="6" pageNumber="7" refId="ref18931" refString="Amsel, H. G. (1939) Beitrag zur Kleinschmetterlings-Fauna der Balkanlander und Kleinasiens. Deutsche entomologische Zeitschrift Iris, Dresden, 52 (1938) (3 / 4), 152 - 159, pl. 4." type="journal article">Amsel (1939)</bibRefCitation>
, this species is close to
<taxonomicName id="4C364D68FFB30978FC7EFA16FB04FA17" authority="(Staudinger)" authorityName="Staudinger" box="[899,1183,1452,1477]" class="Insecta" family="Crambidae" genus="Evergestis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="serratalis">
<emphasis id="B942EAF9FFB30978FC7EFA16FB96FA16" box="[899,1037,1452,1476]" italics="true" pageId="6" pageNumber="7">E. serratalis</emphasis>
(Staudinger)
</taxonomicName>
based on the forewing pattern. However, examination of the male genitalia revealed that it is more close to
<taxonomicName id="4C364D68FFB30978FB95FA6AFA07FA3B" authority="(Staudinger)" authorityName="Staudinger" box="[1128,1436,1488,1513]" class="Insecta" family="Crambidae" genus="Evergestis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="spiniferalis">
<emphasis id="B942EAF9FFB30978FB95FA6AFA9CFA3A" box="[1128,1287,1488,1512]" italics="true" pageId="6" pageNumber="7">E. spiniferalis</emphasis>
(Staudinger)
</taxonomicName>
than
<taxonomicName id="4C364D68FFB30978FF2CFA4EFEC4F9DE" box="[209,351,1524,1548]" class="Insecta" family="Crambidae" genus="Evergestis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="serratalis">
<emphasis id="B942EAF9FFB30978FF2CFA4EFEC4F9DE" box="[209,351,1524,1548]" italics="true" pageId="6" pageNumber="7">E. serratalis</emphasis>
</taxonomicName>
, especially in the shape of valva.
<taxonomicName id="4C364D68FFB30978FD0CFA4EFC57F9DE" authorityName="Amsel" authorityYear="1939" box="[753,972,1524,1548]" class="Insecta" family="Crambidae" genus="Evergestis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="boursini">
<emphasis id="B942EAF9FFB30978FD0CFA4EFC57F9DE" box="[753,972,1524,1548]" italics="true" pageId="6" pageNumber="7">Evergestis boursini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C364D68FFB30978FBF4FA4EFB32F9DE" box="[1033,1193,1524,1548]" class="Insecta" family="Crambidae" genus="Evergestis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="spiniferalis">
<emphasis id="B942EAF9FFB30978FBF4FA4EFB32F9DE" box="[1033,1193,1524,1548]" italics="true" pageId="6" pageNumber="7">E. spiniferalis</emphasis>
</taxonomicName>
, have two groups of cornuti, but in
<taxonomicName id="4C364D68FFB30978FEBBF9A2FE7DF9E2" box="[326,486,1560,1584]" class="Insecta" family="Crambidae" genus="Evergestis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="spiniferalis">
<emphasis id="B942EAF9FFB30978FEBBF9A2FE7DF9E2" box="[326,486,1560,1584]" italics="true" pageId="6" pageNumber="7">E. spiniferalis</emphasis>
</taxonomicName>
they are formed a small group and a large group, while in
<taxonomicName id="4C364D68FFB30978FB59F9A2FAB8F9E2" authorityName="Amsel" authorityYear="1939" box="[1188,1315,1560,1584]" class="Insecta" family="Crambidae" genus="Evergestis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="boursini">
<emphasis id="B942EAF9FFB30978FB59F9A2FAB8F9E2" box="[1188,1315,1560,1584]" italics="true" pageId="6" pageNumber="7">E. boursini</emphasis>
</taxonomicName>
these two groups are in the same size with different densities.
</paragraph>
</subSubSection>
<subSubSection id="C32C6560FFB30978FF3AF9DBFC97F8A7" pageId="6" pageNumber="7" type="description">
<paragraph id="8B8936EBFFB30978FF3AF9DBFC97F8A7" blockId="6.[151,1437,1128,1945]" pageId="6" pageNumber="7">
<emphasis id="B942EAF9FFB30978FF3AF9DBFEEFF9AB" bold="true" box="[199,372,1632,1657]" pageId="6" pageNumber="7">Redescription.</emphasis>
Male genitalia (
<figureCitation id="130D2A6EFFB30978FDCBF9DBFD2EF9AA" box="[566,693,1632,1657]" captionStart="FIGURE 4" captionStartId="7.[151,250,1532,1554]" captionTargetBox="[170,1421,204,1497]" captionTargetId="figure@7.[151,1436,193,1510]" captionTargetPageId="7" captionText="FIGURE 4. Evergestis boursini, Paratype (♂). A) Adult male, upperside. BF) Male genitalia. B: main body and phallus. C: posterior end including uncus and gnathos in ventral view. D: posterior end of uncus in lateral view. E: posterior end of gnathos in laterl view, F: distal end of phallus." httpUri="https://zenodo.org/record/1247142/files/figure.png" pageId="6" pageNumber="7">Figs 4BF</figureCitation>
). Uncus wide at base, narrowed and elongated posteriorly, with blunt tip and sparse short thin setae laterally, without elongated scales distally (
<figureCitation id="130D2A6EFFB30978FBDCF93FFB3FF94F" box="[1057,1188,1668,1693]" captionStart="FIGURE 4" captionStartId="7.[151,250,1532,1554]" captionTargetBox="[170,1421,204,1497]" captionTargetId="figure@7.[151,1436,193,1510]" captionTargetPageId="7" captionText="FIGURE 4. Evergestis boursini, Paratype (♂). A) Adult male, upperside. BF) Male genitalia. B: main body and phallus. C: posterior end including uncus and gnathos in ventral view. D: posterior end of uncus in lateral view. E: posterior end of gnathos in laterl view, F: distal end of phallus." httpUri="https://zenodo.org/record/1247142/files/figure.png" pageId="6" pageNumber="7">Figs 4BD</figureCitation>
); gnathos tapered and narrowed distally, with almost pointed tip, its length slightly more than three-fourths the length of uncus, with a row of small teeth (including 1112 teeth) on its ventral surface extended from the apex to halfway along the shaft (
<figureCitation id="130D2A6EFFB30978FF62F94BFEDAF8DA" box="[159,321,1776,1801]" captionStart="FIGURE 4" captionStartId="7.[151,250,1532,1554]" captionTargetBox="[170,1421,204,1497]" captionTargetId="figure@7.[151,1436,193,1510]" captionTargetPageId="7" captionText="FIGURE 4. Evergestis boursini, Paratype (♂). A) Adult male, upperside. BF) Male genitalia. B: main body and phallus. C: posterior end including uncus and gnathos in ventral view. D: posterior end of uncus in lateral view. E: posterior end of gnathos in laterl view, F: distal end of phallus." httpUri="https://zenodo.org/record/1247142/files/figure.png" pageId="6" pageNumber="7">Figs 4B, C, E</figureCitation>
); valvae slightly narrowed distally, with obliquely rounded apex and basally thickened costa (left valva was damged before dissection), without clasper; juxta elongated oval; distal end of phallus slightly less than two-thirds of its proximal end, with two clusters of cornuti (one cluster dense and another one with less density) and paired brush-like pads at distal portion (
<figureCitation id="130D2A6EFFB30978FD78F8E7FD64F8A7" box="[645,767,1884,1909]" captionStart="FIGURE 4" captionStartId="7.[151,250,1532,1554]" captionTargetBox="[170,1421,204,1497]" captionTargetId="figure@7.[151,1436,193,1510]" captionTargetPageId="7" captionText="FIGURE 4. Evergestis boursini, Paratype (♂). A) Adult male, upperside. BF) Male genitalia. B: main body and phallus. C: posterior end including uncus and gnathos in ventral view. D: posterior end of uncus in lateral view. E: posterior end of gnathos in laterl view, F: distal end of phallus." httpUri="https://zenodo.org/record/1247142/files/figure.png" pageId="6" pageNumber="7">Figs 4B, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C6560FFB30978FF3AF83BFCE6F84B" box="[199,893,1920,1945]" pageId="6" pageNumber="7" type="distribution">
<paragraph id="8B8936EBFFB30978FF3AF83BFCE6F84B" blockId="6.[151,1437,1128,1945]" box="[199,893,1920,1945]" pageId="6" pageNumber="7">
<emphasis id="B942EAF9FFB30978FF3AF83BFEFBF84B" bold="true" box="[199,352,1920,1945]" pageId="6" pageNumber="7">Distribution.</emphasis>
<collectingCountry id="F321767BFFB30978FE9AF83BFE23F84B" box="[359,440,1920,1945]" name="Turkey" pageId="6" pageNumber="7">Turkey</collectingCountry>
,
<collectingCountry id="F321767BFFB30978FE3EF83AFE6BF84A" box="[451,496,1921,1944]" name="Iran" pageId="6" pageNumber="7">Iran</collectingCountry>
(Karaj, Nesa) (
<bibRefCitation id="EFA74B1AFFB30978FD61F83BFCB2F84A" box="[668,809,1920,1945]" pageId="6" pageNumber="7" refId="ref18931" refString="Amsel, H. G. (1939) Beitrag zur Kleinschmetterlings-Fauna der Balkanlander und Kleinasiens. Deutsche entomologische Zeitschrift Iris, Dresden, 52 (1938) (3 / 4), 152 - 159, pl. 4." type="journal article">Amsel 1939</bibRefCitation>
,
<bibRefCitation id="EFA74B1AFFB30978FCC8F83BFCEAF84B" box="[821,881,1920,1945]" pageId="6" pageNumber="7" refId="ref19135" refString="Amsel, H. G. (1961) Die Microlepidopteren der Brandt'schen Iran-Ausbeute. 5. Teil. Arkiv for Zoologie (N. S.), Stockholm (ser. 2) 13 (17), 323 - 445, pls. 1 - 9." type="journal article">1961</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>