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<document id="007227494BA494A24AEF1D2E966CAD6F" ID-CLB-Dataset="51606" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FB36FEFDCBAFBB2CFDCEFDB2" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Syntomodrillia portoricana" docType="treatment" docVersion="14" lastPageNumber="342" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="340" updateTime="1698186879226" updateUser="ExternalLinkService">
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<mods:title id="6E415A31E3269911066469AA3F7EC1C5">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:namePart id="FCD534D145167E39ECC4122A381E1154">Fallon, Phillip J.</mods:namePart>
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<mods:date id="8D1E86E8CA2015162BA70AAFF9141522">2016</mods:date>
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<mods:identifier id="374FADCB87F48622213C59F513607E39" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
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<treatment id="039F87C4FB36FEFDCBAFBB2CFDCEFDB2" ID-DOI="http://doi.org/10.5281/zenodo.6076672" ID-GBIF-Taxon="117911667" ID-Zenodo-Dep="6076672" LSID="urn:lsid:plazi:treatment:039F87C4FB36FEFDCBAFBB2CFDCEFDB2" httpUri="http://treatment.plazi.org/id/039F87C4FB36FEFDCBAFBB2CFDCEFDB2" lastPageId="341" lastPageNumber="342" pageId="339" pageNumber="340">
<subSubSection id="C32C6559FB36FEFBCBAFBB2CFD93FD59" pageId="339" pageNumber="340" type="nomenclature">
<paragraph id="8B8936D2FB36FEFBCBAFBB2CFB8BFDF8" blockId="339.[151,1123,567,627]" box="[151,1123,567,594]" pageId="339" pageNumber="340">
<heading id="D0C181BEFB36FEFBCBAFBB2CFB8BFDF8" bold="true" box="[151,1123,567,594]" fontSize="11" level="1" pageId="339" pageNumber="340" reason="1">
<emphasis id="B942EAC0FB36FEFBCBAFBB2CFB8BFDF8" bold="true" box="[151,1123,567,594]" pageId="339" pageNumber="340">
<emphasis id="B942EAC0FB36FEFBCBAFBB2CFE0EFDFB" bold="true" box="[151,486,567,593]" italics="true" pageId="339" pageNumber="340">
<taxonomicName id="4C364D51FB36FEFBCBAFBB2CFE08FDFB" ID-CoL="53Z9V" box="[151,480,567,593]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="portoricana">Syntomodrillia portoricana</taxonomicName>
,
</emphasis>
new name for
<taxonomicName id="4C364D51FB36FEFBC998BB2CFB8BFDF8" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[672,1123,567,594]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="tantula">
<emphasis id="B942EAC0FB36FEFBC998BB2CFC58FDFB" bold="true" box="[672,944,567,593]" italics="true" pageId="339" pageNumber="340">Syntomodrillia tantula</emphasis>
Bartsch, 1934
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936D2FB36FEFBCBAFBB41FEFBFDD9" blockId="339.[151,1123,567,627]" box="[151,275,602,627]" pageId="339" pageNumber="340">
(
<figureCitation id="130D2A57FB36FEFBCBA7BB41FEE2FDD9" box="[159,266,602,627]" captionStart="PLATE 176" captionStartId="340.[151,234,1593,1615]" captionTargetBox="[151,1436,195,1549]" captionTargetId="figure@340.[151,1436,195,1569]" captionTargetPageId="340" captionText="PLATE 176. Syntomodrillia portoricana, new name. Upper panel. Figs. 1 3: holotype, off N coast of Puerto Rico (USNM 430164). Fig. 1: ventral, lateral &amp; dorsal views; Fig. 2: enlarged view of protoconch (not to scale); and Fig. 3: apical view, V = varix, L = edge of outer lip. Lower panel. Figs. 4, 7 8: Jobos, Puerto Rico (authors coll.): Figs. 4, 7: ventral view; Fig. 8: enlarged view of protoconch (not to scale); Figs. 5 6: Jobos, Puerto Rico (USNM 900153); Figs. 9, 11: off Mayaguez, Puerto Rico (ANSP 253140); Fig. 10: Mayaguez, Puerto Rico (ANSP 314085)." httpUri="https://zenodo.org/record/263476/files/figure.png" pageId="339" pageNumber="340">Plate 176</figureCitation>
)
</paragraph>
<paragraph id="8B8936D2FB36FEFBCBAFBBBBFD93FD59" blockId="339.[151,1435,672,816]" pageId="339" pageNumber="340">
<emphasis id="B942EAC0FB36FEFBCBAFBBBBFE5CFD1C" box="[151,436,672,694]" italics="true" pageId="339" pageNumber="340">
<taxonomicName id="4C364D51FB36FEFBCBAFBBBBFE58FD1C" box="[151,432,672,694]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="portoricana">Syntomodrillia portoricana</taxonomicName>
,
</emphasis>
new name for
<taxonomicName id="4C364D51FB36FEFBC974BBBBFC04FD1C" authority="Bartsch, 1934: 28" authorityName="Bartsch" authorityPageNumber="28" authorityYear="1934" box="[588,1004,672,694]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="tantula">
<emphasis id="B942EAC0FB36FEFBC974BBBBFCD9FD1C" box="[588,817,672,694]" italics="true" pageId="339" pageNumber="340">Syntomodrillia tantula</emphasis>
Bartsch, 1934: 28
</taxonomicName>
29, pl. 8, figs. 2, 3, 6, non
<taxonomicName id="4C364D51FB36FEFBCE3DBBBBFEE7FD7E" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="tantula">
<emphasis id="B942EAC0FB36FEFBCE3DBBBBFEE7FD7E" italics="true" pageId="339" pageNumber="340">Syntomodrillia tantula</emphasis>
</taxonomicName>
(Conrad, 1848a:284; 1848b: 115). Conrads name is a senior secondary homonym (ICZN 2000 Article 53.3) of a fossil of the Oligocene of Vicksburg, Miss.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB36FEFBCBAFBBE7FC5FFC60" pageId="339" pageNumber="340" type="description">
<paragraph id="8B8936D2FB36FEFBCBAFBBE7FB4CFC9A" blockId="339.[151,1435,672,816]" pageId="339" pageNumber="340">
<taxonomicName id="4C364D51FB36FEFBCBAFBBE7FD5AFCB8" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[151,690,764,786]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="tantula" subGenus="Syntomodrillia">
<emphasis id="B942EAC0FB36FEFBCBAFBBE7FDE2FCB8" box="[151,522,764,786]" italics="true" pageId="339" pageNumber="340">Splendrillia (Syntomodrillia) tantula</emphasis>
(Bartsch, 1934)
</taxonomicName>
: Powell (1966: 84); Abbott (1974: 275, species 3112).
<taxonomicName id="4C364D51FB36FEFBCBAFBA01FDECFC9A" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[151,516,794,816]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="tantula">
<emphasis id="B942EAC0FB36FEFBCBAFBA01FEB5FC9A" box="[151,349,794,816]" italics="true" pageId="339" pageNumber="340">Splendrillia tantula</emphasis>
(Bartsch, 1934)
</taxonomicName>
: Rios (1975: 134, pl. 40, fig. 606); Williams (2005: species 1585).
</paragraph>
<paragraph id="8B8936D2FB36FEFBCBAFBA42FD3AFCC4" blockId="339.[151,1436,856,970]" box="[151,722,856,878]" pageId="339" pageNumber="340">
<emphasis id="B942EAC0FB36FEFBCBAFBA42FD3AFCC4" bold="true" box="[151,722,856,878]" pageId="339" pageNumber="340">Not this species, or unconfirmed extralimital reports.</emphasis>
</paragraph>
<paragraph id="8B8936D2FB36FEFBCBAFBA6DFC5FFC60" blockId="339.[151,1436,856,970]" pageId="339" pageNumber="340">
<taxonomicName id="4C364D51FB36FEFBCBAFBA6DFD7BFC26" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[151,659,886,908]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="tantula" subGenus="Splendrillia">
<emphasis id="B942EAC0FB36FEFBCBAFBA6DFE04FC26" box="[151,492,886,908]" italics="true" pageId="339" pageNumber="340">Splendrillia (Splendrillia) tantula</emphasis>
(Bartsch, 1934)
</taxonomicName>
: Rios (1975: 134, pl. 40, fig. 606 [
<collectingCountry id="F3217642FB36FEFBC8D5BA6DFBD9FC26" box="[1005,1073,886,908]" name="Brazil" pageId="339" pageNumber="340">Brazil</collectingCountry>
: Amapá]).
<taxonomicName id="4C364D51FB36FEFBCBAFBA8EFD5DFC01" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[151,693,917,939]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="339" pageNumber="340" phylum="Mollusca" rank="species" species="tantula" subGenus="Syntomodrillia">
<emphasis id="B942EAC0FB36FEFBCBAFBA8EFDE4FC01" box="[151,524,917,939]" italics="true" pageId="339" pageNumber="340">Splendrillia (Syntomodrillia) tantula</emphasis>
(Bartsch, 1934)
</taxonomicName>
: Rios (1985: 140; pl. 48, fig. 646 [
<collectingCountry id="F3217642FB36FEFBCF20BA8EFBB4FC01" box="[1048,1116,917,939]" name="Brazil" pageId="339" pageNumber="340">Brazil</collectingCountry>
: Amapá]); Rios (1994: 163; pl. 54, fig. 738 [
<collectingCountry id="F3217642FB36FEFBCA70BAAFFE64FC60" box="[328,396,948,970]" name="Brazil" pageId="339" pageNumber="340">Brazil</collectingCountry>
: Amapá]); Rios (2009: 315, fig. 802 [
<collectingCountry id="F3217642FB36FEFBC830BAAFFCA4FC60" box="[776,844,948,970]" name="Brazil" pageId="339" pageNumber="340">Brazil</collectingCountry>
: Amapá]).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB36FEFBCBAFBAE9FC21FAE1" pageId="339" pageNumber="340" type="materials_examined">
<paragraph id="8B8936D2FB36FEFBCBAFBAE9FD29FB87" blockId="339.[151,1437,1009,2034]" pageId="339" pageNumber="340">
<emphasis id="B942EAC0FB36FEFBCBAFBAE9FEAEFBA0" bold="true" box="[151,326,1009,1034]" pageId="339" pageNumber="340">
<typeStatus id="548D8870FB36FEFBCBAFBAE9FF3AFBA3" box="[151,210,1010,1033]" pageId="339" pageNumber="340">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548D8870FB36FEFBCA75BAEAFE5DFBA0" box="[333,437,1009,1034]" pageId="339" pageNumber="340" type="holotype">Holotype</typeStatus>
7.0 x
<quantity id="4CCE9B37FB36FEFBCAC7BAE9FDBFFBA3" box="[511,599,1010,1034]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="339" pageNumber="340" unit="mm" value="2.5">2.5 mm</quantity>
(7.1 x 2.8 according to Bartsch) (USNM 430164). The
<typeStatus id="548D8870FB36FEFBCFEABAE9FADAFBA3" box="[1234,1330,1010,1033]" pageId="339" pageNumber="340" type="paratype">paratype</typeStatus>
from the
<typeStatus id="548D8870FB36FEFBCBAFBD0DFF2FFB87" box="[151,199,1046,1069]" pageId="339" pageNumber="340">type</typeStatus>
locality was not examined (USNM 430854).
</paragraph>
<paragraph id="8B8936D2FB36FEFBCBFFBD22FB72FBDF" blockId="339.[151,1437,1009,2034]" pageId="339" pageNumber="340">
<emphasis id="B942EAC0FB36FEFBCBFFBD22FE8EFBFB" bold="true" box="[199,358,1080,1105]" pageId="339" pageNumber="340">
<typeStatus id="548D8870FB36FEFBCBFFBD22FEEAFBFA" box="[199,258,1081,1104]" pageId="339" pageNumber="340">Type</typeStatus>
locality.
</emphasis>
R/V
<emphasis id="B942EAC0FB36FEFBCA9ABD23FDEDFBFB" box="[418,517,1080,1105]" italics="true" pageId="339" pageNumber="340">Caroline</emphasis>
Sta. 16, off the N coast of
<collectingCountry id="F3217642FB36FEFBC80BBD22FC51FBFB" box="[819,953,1080,1105]" name="Puerto Rico" pageId="339" pageNumber="340">Puerto Rico</collectingCountry>
near Morro Castle light and Punta Salinas, between
<geoCoordinate id="EE025015FB36FEFBCBC3BD46FE6AFBDF" box="[251,386,1117,1141]" direction="north" orientation="latitude" pageId="339" pageNumber="340" precision="15" value="18.494444">18°29'40&quot;N</geoCoordinate>
,
<geoCoordinate id="EE025015FB36FEFBCAB5BD46FDFFFBDF" box="[397,535,1117,1141]" direction="west" orientation="longitude" pageId="339" pageNumber="340" precision="15" value="-66.14166">66°08'30&quot;W</geoCoordinate>
and
<geoCoordinate id="EE025015FB36FEFBC976BD46FD3BFBDF" box="[590,723,1117,1141]" direction="north" orientation="latitude" pageId="339" pageNumber="340" precision="15" value="18.516666">18°31'00&quot;N</geoCoordinate>
,
<geoCoordinate id="EE025015FB36FEFBC9E7BD46FC81FBDF" box="[735,873,1117,1141]" direction="west" orientation="longitude" pageId="339" pageNumber="340" precision="15" value="-66.17083">66°10'15&quot;W</geoCoordinate>
, in 3895 fms [
<quantity id="4CCE9B37FB36FEFBCF2EBD46FB67FBDF" box="[1046,1167,1117,1141]" metricMagnitude="2" metricUnit="m" metricValue="1.215" metricValueMax="1.7399999999999998" metricValueMin="0.69" pageId="339" pageNumber="340" unit="m" value="121.5" valueMax="174.0" valueMin="69.0">69174 m</quantity>
].
</paragraph>
<paragraph id="8B8936D2FB36FEFBCBFFBD9BFE05FA82" blockId="339.[151,1437,1009,2034]" pageId="339" pageNumber="340">
<emphasis id="B942EAC0FB36FEFBCBFFBD9BFE15FB33" bold="true" box="[199,509,1152,1177]" pageId="339" pageNumber="340">Other material examined.</emphasis>
An additional eight specimens were examined: 2 spec., 7.8 x 2.9 &amp; 7.4 x
<quantity id="4CCE9B37FB36FEFBCE07BD9BFA70FB32" box="[1343,1432,1152,1176]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="339" pageNumber="340" unit="mm" value="2.7">2.7 mm</quantity>
, in
<quantity id="4CCE9B37FB36FEFBCB8CBDBEFF0BFB16" box="[180,227,1189,1212]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="339" pageNumber="340" unit="m" value="5.0">5 m</quantity>
, Boca de Inferno, Jobos,
<collectingCountry id="F3217642FB36FEFBC93BBDBEFD62FB17" box="[515,650,1188,1213]" name="Puerto Rico" pageId="339" pageNumber="340">Puerto Rico</collectingCountry>
(USNM 900153); 2 spec., 6.9 x 2.7 &amp; 7.2 x
<quantity id="4CCE9B37FB36FEFBCFB1BDBFFB08FB16" box="[1161,1248,1188,1212]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="339" pageNumber="340" unit="mm" value="2.5">2.5 mm</quantity>
in
<quantity id="4CCE9B37FB36FEFBCE3DBDBEFADCFB16" box="[1285,1332,1189,1212]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="339" pageNumber="340" unit="m" value="5.0">5 m</quantity>
, Boca de Inferno, Jobos,
<collectingCountry id="F3217642FB36FEFBCA71BDD3FE39FB4A" box="[329,465,1223,1248]" name="Puerto Rico" pageId="339" pageNumber="340">Puerto Rico</collectingCountry>
(authors coll.); 2 spec., 8.6 x 3.1 &amp; 8.3 x
<quantity id="4CCE9B37FB36FEFBC8FDBDD3FBC8FB75" box="[965,1056,1224,1248]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="339" pageNumber="340" unit="mm" value="2.9">2.9 mm</quantity>
, off Mayaguez,
<collectingCountry id="F3217642FB36FEFBCFE4BDD3FA8DFB4A" box="[1244,1381,1223,1248]" name="Puerto Rico" pageId="339" pageNumber="340">Puerto Rico</collectingCountry>
, G.L Warmke! 1952 (ANSP 253140); 1 spec., 9.4 x
<quantity id="4CCE9B37FB36FEFBC991BDF7FCEAFAA9" box="[681,770,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="339" pageNumber="340" unit="mm" value="3.2">3.2 mm</quantity>
, in
<quantity id="4CCE9B37FB36FEFBC814BDF7FC80FAA9" box="[812,872,1260,1284]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="339" pageNumber="340" unit="m" value="12.0">12 m</quantity>
, off docks, Mayaguez
<collectingCountry id="F3217642FB36FEFBCF51BDF7FB18FAAE" box="[1129,1264,1259,1284]" name="Puerto Rico" pageId="339" pageNumber="340">Puerto Rico</collectingCountry>
, G.L. Warmke!
<date id="FF881012FB36FEFBCBAFBC14FEC1FA8D" box="[151,297,1295,1319]" pageId="339" pageNumber="340" value="1952-11-16">16 Nov 1952</date>
(ANSP 314085).
</paragraph>
<paragraph id="8B8936D2FB36FEFBCBFFBC2FFC21FAE1" blockId="339.[151,1437,1009,2034]" box="[199,969,1331,1356]" pageId="339" pageNumber="340">
<emphasis id="B942EAC0FB36FEFBCBFFBC2FFE43FAE6" bold="true" box="[199,427,1331,1356]" pageId="339" pageNumber="340">Range and habitat.</emphasis>
<collectingCountry id="F3217642FB36FEFBCA8ABC2FFDDFFAE6" box="[434,567,1331,1356]" name="Puerto Rico" pageId="339" pageNumber="340">Puerto Rico</collectingCountry>
(N coast). Reported from
<quantity id="4CCE9B37FB36FEFBC866BC28FC2CFAE1" box="[862,964,1331,1355]" metricMagnitude="1" metricUnit="m" metricValue="6.5" metricValueMax="12.5" metricValueMin="0.5" pageId="339" pageNumber="340" unit="m" value="65.0" valueMax="125.0" valueMin="5.0">
5
<specimenCount id="9D30FD5BFB36FEFBC843BC28FC2CFAE1" box="[891,964,1331,1355]" pageId="339" pageNumber="340" type="generic">125 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB36FEFBCBFFBC4DFC12F858" pageId="339" pageNumber="340" type="description">
<paragraph id="8B8936D2FB36FEFBCBFFBC4DFC12F858" blockId="339.[151,1437,1009,2034]" pageId="339" pageNumber="340">
<emphasis id="B942EAC0FB36FEFBCBFFBC4DFE73FAC5" bold="true" box="[199,411,1366,1391]" pageId="339" pageNumber="340">
Description.
<emphasis id="B942EAC0FB36FEFBCA58BC4DFE73FAC5" bold="true" box="[352,411,1366,1391]" italics="true" pageId="339" pageNumber="340">Shell</emphasis>
</emphasis>
very small (to
<quantity id="4CCE9B37FB36FEFBC97FBC4CFD76FAC4" box="[583,670,1367,1391]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="339" pageNumber="340" unit="mm" value="9.4">9.4 mm</quantity>
total length), narrowly fusiform, glossy, whorls slightly convex, rib peripheries below mid-whorl; whorls to about 9¾, the last approximately 59% of total length. Sculptured with numerous ribs; aperture oval and short.
<emphasis id="B942EAC0FB36FEFBC95EBC85FD04FA1D" bold="true" box="[614,748,1438,1463]" italics="true" pageId="339" pageNumber="340">Protoconch</emphasis>
of approximately 1½ round, rapidly expanding, translucent whorls, tip of first partially impressed in second; protoconch globular, but not exceeding diameter of first teleoconch whorl.
<emphasis id="B942EAC0FB36FEFBCA51BCFEFDF2FA54" bold="true" box="[361,538,1509,1534]" italics="true" pageId="339" pageNumber="340">Axial sculpture</emphasis>
of broad ribs suture-to-suture on spire whorls, to fasciole on last. Ribs slightly opisthocline, recurved in sulcus; 811 on penultimate, 510 on last whorl to varix. Ribs about as wide as intercostal space; rib crests ridged.
<emphasis id="B942EAC0FB36FEFBCAF9BF36FE14F9EE" bold="true" box="[449,508,1581,1606]" italics="true" pageId="339" pageNumber="340">Vari x</emphasis>
heavy, straight, not oblique like adjacent ribs, located approximately ⅓-turn from edge of outer lip.
<emphasis id="B942EAC0FB36FEFBCA5ABF4BFDF3F9C3" bold="true" box="[354,539,1616,1641]" italics="true" pageId="339" pageNumber="340">Spiral sculpture</emphasis>
of deep grooves appearing as threads between the ribs (obsolete on rib crests), absent in sulcus but present on all whorls; threads become stronger on shell base then become ridges just before and on anterior fasciole, 910 on fasciole.
<emphasis id="B942EAC0FB36FEFBC9BCBF83FD3AF91B" bold="true" box="[644,722,1688,1713]" italics="true" pageId="339" pageNumber="340">Sulcus</emphasis>
absent; ribs and growth striae are curved just below suture to reflect the outline of anal sinus, spiral grooves absent.
<emphasis id="B942EAC0FB36FEFBC9C1BFA7FC88F97F" bold="true" box="[761,864,1724,1749]" italics="true" pageId="339" pageNumber="340">Outer lip</emphasis>
thin, translucent, flattened outward from the varix, with one or more strengthening axial folds; edge of lip from anal sinus to anterior canal forms a low arc; slightly indented at stromboid notch and flexed outward along canal.
<emphasis id="B942EAC0FB36FEFBC87EBE18FC2AF8B6" bold="true" box="[838,962,1795,1820]" italics="true" pageId="339" pageNumber="340">Anal sinus</emphasis>
a deep U-shaped notch on whorl shoulder near suture, sides parallel; outer lip and sinus inner margin flared out; sinus angled away from shell axis.
<emphasis id="B942EAC0FB36FEFBCE0EBE33FA73F8EA" bold="true" box="[1334,1435,1831,1856]" italics="true" pageId="339" pageNumber="340">Inner lip</emphasis>
recumbent, moderately wide, margined, thickest anteriorly, thin on parietal wall; developed into an elongate lobe posteriorly along one side of the anal sinus on the roof of the aperture.
<emphasis id="B942EAC0FB36FEFBC88ABE75FBB3F82D" bold="true" box="[946,1115,1902,1927]" italics="true" pageId="339" pageNumber="340">Anterior canal</emphasis>
moderately long, open, bent slightly to the right at its end when viewed ventrally; unnotched; columella straight; fasciole not swollen.
<emphasis id="B942EAC0FB36FEFBCE63BE89FA74F801" bold="true" box="[1371,1436,1938,1963]" italics="true" pageId="339" pageNumber="340">Color</emphasis>
pale wax yellow (
<typeStatus id="548D8870FB36FEFBCA59BEADFE21F865" box="[353,457,1974,1999]" pageId="339" pageNumber="340" type="holotype">holotype</typeStatus>
); what are possibly fresher specimens are a golden brown, the shell base and outer lip paler. A darker golden brown patch is present on the ventral side of the varix.
</paragraph>
</subSubSection>
<caption id="DF49665AFB31FEFCCBAFBF22FC9BF960" httpUri="https://zenodo.org/record/263476/files/figure.png" pageId="340" pageNumber="341" targetBox="[151,1436,195,1549]" targetPageId="340">
<paragraph id="8B8936D2FB31FEFCCBAFBF22FC9BF960" blockId="340.[151,1436,1593,1738]" pageId="340" pageNumber="341">
<emphasis id="B942EAC0FB31FEFCCBAFBF22FEF6F9E5" bold="true" box="[151,286,1593,1615]" pageId="340" pageNumber="341">PLATE 176.</emphasis>
<taxonomicName id="4C364D51FB31FEFCCA1EBF22FDD5F9E5" box="[294,573,1593,1615]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="340" pageNumber="341" phylum="Mollusca" rank="species" species="portoricana">
<emphasis id="B942EAC0FB31FEFCCA1EBF22FDD5F9E5" box="[294,573,1593,1615]" italics="true" pageId="340" pageNumber="341">Syntomodrillia portoricana</emphasis>
</taxonomicName>
, new name.
<emphasis id="B942EAC0FB31FEFCC9FBBF21FCB9F9E5" bold="true" box="[707,849,1594,1615]" pageId="340" pageNumber="341">Upper panel.</emphasis>
Figs. 13: holotype, off N coast of Puerto Rico (USNM 430164). Fig. 1: ventral, lateral &amp; dorsal views; Fig. 2: enlarged view of protoconch (not to scale); and Fig. 3: apical view, V = varix, L = edge of outer lip.
<emphasis id="B942EAC0FB31FEFCCAFBBF6CFDA4F926" bold="true" box="[451,588,1655,1676]" pageId="340" pageNumber="341">Lower panel</emphasis>
. Figs. 4, 78: Jobos, Puerto Rico (authors coll.): Figs. 4, 7: ventral view; Fig. 8: enlarged view of protoconch (not to scale); Figs. 56: Jobos, Puerto Rico (USNM 900153); Figs. 9, 11: off Mayaguez, Puerto Rico (ANSP 253140); Fig. 10: Mayaguez, Puerto Rico (ANSP 314085).
</paragraph>
</caption>
<subSubSection id="C32C6559FB31FEFDCBFFBFECFC27FE7A" lastPageId="341" lastPageNumber="342" pageId="340" pageNumber="341" type="discussion">
<paragraph id="8B8936D2FB31FEFCCBFFBFECFA75F84D" blockId="340.[151,1437,1782,2023]" pageId="340" pageNumber="341">
<emphasis id="B942EAC0FB31FEFCCBFFBFECFE01F8A5" bold="true" box="[199,489,1782,1807]" pageId="340" pageNumber="341">
Remarks.
<emphasis id="B942EAC0FB31FEFCCA7CBFEDFE01F8A5" bold="true" box="[324,489,1782,1807]" italics="true" pageId="340" pageNumber="341">Nomenclature</emphasis>
</emphasis>
. An entirely different species,
<taxonomicName id="4C364D51FB31FEFCC874BFEDFA74F8A5" authority="Conrad, 1848a: 284" authorityName="Conrad" authorityPageNumber="284" authorityYear="1848" box="[844,1436,1782,1807]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="340" pageNumber="341" phylum="Mollusca" rank="species" species="tantula" subGenus="Clavatula">
<emphasis id="B942EAC0FB31FEFCC874BFEDFB59F8A5" box="[844,1201,1782,1807]" italics="true" pageId="340" pageNumber="341">Pleurotoma (Clavatula) tantula</emphasis>
Conrad, 1848a: 284
</taxonomicName>
(Oligocene, Vicksburg, Mississippi fossil) was placed in
<taxonomicName id="4C364D51FB31FEFCC82BBE01FC52F899" box="[787,954,1818,1843]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="340" pageNumber="341" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB31FEFCC82BBE01FC52F899" box="[787,954,1818,1843]" italics="true" pageId="340" pageNumber="341">Syntomodrillia</emphasis>
</taxonomicName>
by Mansfield (1940: 206, pl. 27, fig. 15), a combination recently used by MacNeil &amp; Dockery (1984: 201, pl. 36, figs. 1315, 18, 19).
<collectingCountry id="F3217642FB31FEFCCFFEBE24FB0DF8FD" box="[1222,1253,1855,1879]" name="American Samoa" pageId="340" pageNumber="341">As</collectingCountry>
such, Bartschs species becomes a junior secondary homonym, as Conrads name has priority. To the best of this authors knowledge, there are no available synonyms for Bartschs species, so the new name is proposed herein.
<emphasis id="B942EAC0FB31FEFCCE25BE9CFE23F868" italics="true" pageId="340" pageNumber="341">
<emphasis id="B942EAC0FB31FEFCCE25BE9CFA71F834" bold="true" box="[1309,1433,1927,1950]" italics="true" pageId="340" pageNumber="341">Taxonomy.</emphasis>
<taxonomicName id="4C364D51FB31FEFCCBAFBEB1FE23F868" box="[151,459,1962,1987]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="340" pageNumber="341" phylum="Mollusca" rank="species" species="portoricana">Syntomodrillia portoricana</taxonomicName>
</emphasis>
has all of the characteristics described by Woodring for the genus: small, glossy, axial ribs numerous, from suture-to-suture, microscopic threads between the ribs, and a moderately long anterior canal.
</paragraph>
<paragraph id="8B8936D2FB30FEFDCBAFB98CFC27FE7A" blockId="341.[151,1437,151,536]" pageId="341" pageNumber="342">
<emphasis id="B942EAC0FB30FEFDCBAFB98CFEFDFF1A" bold="true" box="[151,277,151,176]" italics="true" pageId="341" pageNumber="342">Variability.</emphasis>
The average total length of eight specimens is
<quantity id="4CCE9B37FB30FEFDC819B983FC6CFF05" box="[801,900,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="7.83" pageId="341" pageNumber="342" unit="mm" value="7.83">7.83 mm</quantity>
(
<quantity id="4CCE9B37FB30FEFDC8AAB98CFBF6FF1A" box="[914,1054,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="8.15" metricValueMax="9.4" metricValueMin="6.9" pageId="341" pageNumber="342" unit="mm" value="8.15" valueMax="9.4" valueMin="6.9">6.99.4 mm</quantity>
); their average W/
<taxonomicName id="4C364D51FB30FEFDCFD4B98CFADEFF1A" box="[1260,1334,151,176]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="ratio">L ratio</taxonomicName>
is 0.360.
<emphasis id="B942EAC0FB30FEFDCBAFB9A7FED3FF7F" bold="true" box="[151,315,188,213]" italics="true" pageId="341" pageNumber="342">Identification.</emphasis>
Some of the small, glossy, spindle-shaped species in the genus
<taxonomicName id="4C364D51FB30FEFDCF20B9A7FB56FF7F" box="[1048,1214,188,213]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB30FEFDCF20B9A7FB56FF7F" box="[1048,1214,188,213]" italics="true" pageId="341" pageNumber="342">Syntomodrillia</emphasis>
</taxonomicName>
may be difficult to distinguish.
<taxonomicName id="4C364D51FB30FEFDCA18B9C4FDBCFF5D" box="[288,596,223,248]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="portoricana">
<emphasis id="B942EAC0FB30FEFDCA18B9C4FDBCFF5D" box="[288,596,223,248]" italics="true" pageId="341" pageNumber="342">Syntomodrillia portoricana</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C364D51FB30FEFDC9D0B9FBFBFDFF52" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[744,1045,223,248]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="carolinae">
<emphasis id="B942EAC0FB30FEFDC9D0B9FBFC86FF52" box="[744,878,223,248]" italics="true" pageId="341" pageNumber="342">S. carolinae</emphasis>
Bartsch, 1934
</taxonomicName>
,
<taxonomicName id="4C364D51FB30FEFDCF19B9FBFA8FFF52" authority="(E.A. Smith, 1888)" baseAuthorityName="E. A. Smith" baseAuthorityYear="1888" box="[1057,1383,223,248]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="cookei">
<emphasis id="B942EAC0FB30FEFDCF19B9FBFB6EFF52" box="[1057,1158,223,248]" italics="true" pageId="341" pageNumber="342">S. cookei</emphasis>
(E.A. Smith, 1888)
</taxonomicName>
, and
<taxonomicName id="4C364D51FB30FEFDCBAFB81FFEFCFEB7" box="[151,276,260,285]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="hesperia" status="sp. nov.">
<emphasis id="B942EAC0FB30FEFDCBAFB81FFEFCFEB7" box="[151,276,260,285]" italics="true" pageId="341" pageNumber="342">S. hesperia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFB30FEFDCA1BB81EFE47FEB7" box="[291,431,260,285]" pageId="341" pageNumber="342" rank="species">new species</taxonomicNameLabel>
. From
<taxonomicName id="4C364D51FB30FEFDCAC7B81FFD6FFEB7" box="[511,647,260,285]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="carolinae">
<emphasis id="B942EAC0FB30FEFDCAC7B81FFD6FFEB7" box="[511,647,260,285]" italics="true" pageId="341" pageNumber="342">S. carolinae</emphasis>
</taxonomicName>
it differs in having whorl peripheries below mid-whorl, not generally mid-whorl, a less translucent shell, fewer and broader ribs (811 versus 1014 on the penultimate whorl), an anterior canal bent to the right when viewed ventrally, not straight, and more spiral threads on the anterior canal (9 10 versus 35). From
<taxonomicName id="4C364D51FB30FEFDCAA9B86BFE1FFE22" box="[401,503,367,392]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="cookei">
<emphasis id="B942EAC0FB30FEFDCAA9B86BFE1FFE22" box="[401,503,367,392]" italics="true" pageId="341" pageNumber="342">S. cookei</emphasis>
</taxonomicName>
it differs in having broader ribs that are not as oblique, a mostly opaque shell, not translucent as in
<taxonomicName id="4C364D51FB30FEFDCA5EB88FFE3AFE07" box="[358,466,404,429]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="cookei">
<emphasis id="B942EAC0FB30FEFDCA5EB88FFE3AFE07" box="[358,466,404,429]" italics="true" pageId="341" pageNumber="342">S. cookei</emphasis>
</taxonomicName>
, and a different coloration. From
<taxonomicName id="4C364D51FB30FEFDC84BB88FFC1EFE07" box="[883,1014,404,429]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="species" species="hesperia">
<emphasis id="B942EAC0FB30FEFDC84BB88FFC1EFE07" box="[883,1014,404,429]" italics="true" pageId="341" pageNumber="342">S. hesperia</emphasis>
</taxonomicName>
it differs in being stockier, having opisthocline, not straight ribs, and in lacking a diffuse white central band.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB30FEFDCBFFB8C6FDCEFDB2" pageId="341" pageNumber="342" type="etymology">
<paragraph id="8B8936D2FB30FEFDCBFFB8C6FDCEFDB2" blockId="341.[151,1437,151,536]" pageId="341" pageNumber="342">
<emphasis id="B942EAC0FB30FEFDCBFFB8C6FEA6FE5E" bold="true" box="[199,334,477,500]" pageId="341" pageNumber="342">Etymology.</emphasis>
The Puerto Rican
<taxonomicName id="4C364D51FB30FEFDC91BB8C7FD21FE5F" box="[547,713,476,501]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="341" pageNumber="342" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB30FEFDC91BB8C7FD21FE5F" box="[547,713,476,501]" italics="true" pageId="341" pageNumber="342">Syntomodrillia</emphasis>
</taxonomicName>
. The name recognizes the country of the
<typeStatus id="548D8870FB30FEFDCF98B8C6FB38FE5E" box="[1184,1232,477,500]" pageId="341" pageNumber="342">type</typeStatus>
locality where all specimens have been found to date.
</paragraph>
</subSubSection>
</treatment>
</document>