177 lines
18 KiB
XML
177 lines
18 KiB
XML
<document id="E2543A0D12BD1E404F4B5CC584DF2441" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA72FFBFCBAFBBCCFC9DF839" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bellaspira amplicostata Fallon, 2016, new species" docType="treatment" docVersion="12" lastPageNumber="24" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="24" updateTime="1698184378282" updateUser="plazi">
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<mods:mods id="A5A58F69E2D21F2926D7BF9746365B6F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F4753BF7E7718106DBEBD7142B226BF5">
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<mods:title id="0F2F55FFB61002EA41E0B44B6024B318">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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</mods:titleInfo>
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<mods:name id="F12955E72F83058BECA93C130E93A290" type="personal">
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<mods:role id="0516911491EE994522F53692711E16B7">
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<mods:roleTerm id="DAB2916DD5DCF71ED41332CE1496FF0A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="57079E5EED7FB004E8B9B0590298D672">Fallon, Phillip J.</mods:namePart>
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<mods:typeOfResource id="CE20B89D945E43D84EFB7168DF03F2D4">text</mods:typeOfResource>
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<mods:titleInfo id="48A6E7DF11817B34AFE3F076C713C39E">
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<mods:title id="85FA97928A9203E28320760C888743B7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C0117743D921D9C3FABBEA20B2C56083">
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<mods:date id="2C05C1717C389D56A1F2E45B231B8455">2016</mods:date>
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<mods:detail id="7B4D245D657EF30AADF51E0049B41395" type="volume">
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<mods:number id="3C7397603D1E77D10B3D84F5655C6293">4090</mods:number>
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</mods:detail>
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<mods:detail id="D7FDA5A0AD78CA8135E912CD4E62F0CA" type="issue">
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<mods:number id="7F836575BE20DBC758BC645F86C0356A">1</mods:number>
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<mods:classification id="BD93B3B6F2A939CECFED01066D3FF72D">journal article</mods:classification>
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<mods:identifier id="EBC87D2DCCFC50F1E2D6CC7A5F7FD632" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
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<mods:identifier id="78392DF75A9EEEF719DCC035CA9AADAD" type="GBIF-Dataset">e6b35f5a-435e-4473-b29e-1e4d842f84b0</mods:identifier>
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<mods:identifier id="617E8FD05D0B51C79CBCC6C3A116568B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B7DF95A142D72728552665C3F30ECD31" type="Zenodo-Dep">263299</mods:identifier>
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<mods:identifier id="9F2649E8BBD6DE4799B7B96E53956BBA" type="ZooBank">203BAC25-B542-48FE-B5AD-EBA8C0285833</mods:identifier>
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</mods:mods>
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<treatment id="039F87C4FA72FFBFCBAFBBCCFC9DF839" ID-DOI="http://doi.org/10.5281/zenodo.6076249" ID-GBIF-Taxon="117911587" ID-Zenodo-Dep="6076249" LSID="urn:lsid:plazi:treatment:039F87C4FA72FFBFCBAFBBCCFC9DF839" httpUri="http://treatment.plazi.org/id/039F87C4FA72FFBFCBAFBBCCFC9DF839" lastPageNumber="24" pageId="23" pageNumber="24">
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<subSubSection id="C32C6559FA72FFBFCBAFBBCCFF1FFCB9" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8B8936D2FA72FFBFCBAFBBCCFDB9FD58" blockId="23.[151,593,727,787]" box="[151,593,727,754]" pageId="23" pageNumber="24">
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<heading id="D0C181BEFA72FFBFCBAFBBCCFDB9FD58" bold="true" box="[151,593,727,754]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B942EAC0FA72FFBFCBAFBBCCFDB9FD58" bold="true" box="[151,593,727,754]" pageId="23" pageNumber="24">
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<taxonomicName id="4C364D51FA72FFBFCBAFBBCCFE5AFD5B" ID-CoL="5WGLS" box="[151,434,727,753]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="amplicostata" status="sp. nov.">
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<emphasis id="B942EAC0FA72FFBFCBAFBBCCFE5AFD5B" bold="true" box="[151,434,727,753]" italics="true" pageId="23" pageNumber="24">Bellaspira amplicostata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27157BBFA72FFBFCAF9BBC3FDB9FD58" box="[449,593,728,754]" pageId="23" pageNumber="24" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936D2FA72FFBFCBAFBBE1FF1FFCB9" blockId="23.[151,593,727,787]" box="[151,247,762,787]" pageId="23" pageNumber="24">
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(
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<figureCitation id="130D2A57FA72FFBFCBA7BBE1FF06FCB9" box="[159,238,762,787]" captionStart="PLATE 4" captionStartId="24.[151,234,1935,1957]" captionTargetBox="[211,1368,193,1894]" captionTargetId="figure@24.[211,1368,193,1894]" captionTargetPageId="24" captionText="PLATE 4. Bellaspira amplicostata, new species. Upper panel. Fig. 1: holotype, NE of Contoy Light, Contoy I., Quintana Roo, Mexico (UF 449328), ventral, lateral & dorsal views. Lower panel. Figs. 2 – 11: paratypes, all from the original type lot (UF 449327)." httpUri="https://zenodo.org/record/263303/files/figure.png" pageId="23" pageNumber="24">Plate 4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA72FFBFCBAFBA58FCD4FC41" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8B8936D2FA72FFBFCBAFBA58FE96FC09" blockId="23.[151,1437,834,1939]" pageId="23" pageNumber="24">
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<emphasis id="B942EAC0FA72FFBFCBAFBA58FEADFCF1" bold="true" box="[151,325,834,859]" pageId="23" pageNumber="24">
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<typeStatus id="548D8870FA72FFBFCBAFBA58FF3AFCF0" box="[151,210,835,858]" pageId="23" pageNumber="24">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="548D8870FA72FFBFCA75BA59FE5DFCF1" box="[333,437,834,859]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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7.4 x
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<quantity id="4CCE9B37FA72FFBFCAC5BA58FDBCFCF0" box="[509,596,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="23" pageNumber="24" unit="mm" value="3.5">3.5 mm</quantity>
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(UF 449328); 11
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<typeStatus id="548D8870FA72FFBFC818BA58FC66FCF0" box="[800,910,835,858]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
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, 6.6 x 3.0, 7.3 x 3.2, 8.0 x 3.2, 6.9 x 3.0, 7.0 x 3.2, 7.1 x 3.0, 7.2 x 3.2, 6.6 x 3.0, 7.1 x 3.2, 6.2 x 2.9 & 7.5 x 3.3, all from the
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<typeStatus id="548D8870FA72FFBFCF10BA7CFBB0FCD4" box="[1064,1112,871,894]" pageId="23" pageNumber="24">type</typeStatus>
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locality (UF 449327). Riley Black!
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<date id="FF881012FA72FFBFCBD0BA90FE92FC09" box="[232,378,907,931]" pageId="23" pageNumber="24" value="1967-04-20">20 Apr 1967</date>
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.
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</paragraph>
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<paragraph id="8B8936D2FA72FFBFCBFFBAB4FBA3FC6C" blockId="23.[151,1437,834,1939]" box="[199,1099,942,967]" pageId="23" pageNumber="24">
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<emphasis id="B942EAC0FA72FFBFCBFFBAB4FE8DFC6D" bold="true" box="[199,357,942,967]" pageId="23" pageNumber="24">
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<typeStatus id="548D8870FA72FFBFCBFFBAB4FEEAFC6C" box="[199,258,943,966]" pageId="23" pageNumber="24">Type</typeStatus>
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locality.
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</emphasis>
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NE of Contoy Light, Contoy I., Quintana Roo,
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<collectingCountry id="F3217642FA72FFBFC845BAB5FC3DFC6D" box="[893,981,942,967]" name="Mexico" pageId="23" pageNumber="24">Mexico</collectingCountry>
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, in
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<specimenCount id="9D30FD5BFA72FFBFC8C5BAB4FBA3FC6C" box="[1021,1099,943,967]" pageId="23" pageNumber="24" type="generic">
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<quantity id="4CCE9B37FA72FFBFC8C5BAB4FBAFFC6C" box="[1021,1095,943,967]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="23" pageNumber="24" unit="m" value="183.0">183 m</quantity>
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.
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</specimenCount>
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</paragraph>
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<paragraph id="8B8936D2FA72FFBFCBFFBAC8FCD4FC41" blockId="23.[151,1437,834,1939]" box="[199,828,978,1003]" pageId="23" pageNumber="24">
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<emphasis id="B942EAC0FA72FFBFCBFFBAC8FE43FC41" bold="true" box="[199,427,978,1003]" pageId="23" pageNumber="24">Range and habitat.</emphasis>
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Known only from the
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<typeStatus id="548D8870FA72FFBFC993BAC8FD33FC40" box="[683,731,979,1002]" pageId="23" pageNumber="24">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA72FFBFCBFFBAEDFB7DFA15" pageId="23" pageNumber="24" type="description">
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<paragraph id="8B8936D2FA72FFBFCBFFBAEDFB7DFA15" blockId="23.[151,1437,834,1939]" pageId="23" pageNumber="24">
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<emphasis id="B942EAC0FA72FFBFCBFFBAEDFE74FBA5" bold="true" box="[199,412,1014,1039]" pageId="23" pageNumber="24">
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Description.
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<emphasis id="B942EAC0FA72FFBFCA58BAEDFE74FBA5" bold="true" box="[352,412,1014,1039]" italics="true" pageId="23" pageNumber="24">Shell</emphasis>
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</emphasis>
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very small (to
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<quantity id="4CCE9B37FA72FFBFC971BAECFD4BFBA5" box="[585,675,1014,1039]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="24" unit="mm" value="7.5">7.5 mm</quantity>
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), stoutly fusiform, truncated anteriorly, glossy, with up to 7 convex whorls, suture impressed—undulates over preceding whorl’s ribs; heavily ribbed. Last whorl large, approximately 64% of total shell length.
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<emphasis id="B942EAC0FA72FFBFCA8FBD25FDD4FBFD" bold="true" box="[439,572,1086,1111]" italics="true" pageId="23" pageNumber="24">Protoconch</emphasis>
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of 2 whorls, the first partially immersed in the second giving the shell apex a bluntly round shape.
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<emphasis id="B942EAC0FA72FFBFCAB6BD79FDABFBD1" bold="true" box="[398,579,1122,1147]" italics="true" pageId="23" pageNumber="24">Axial sculpture</emphasis>
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of broad, round ribs that extend from suture-to-suture, evanesce on body whorl at anterior fasciole, intercostal space narrower than ribs; 7–8 on penultimate and 4–6 on body whorl to varix. Growth striae present, heaviest on rib sides.
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<emphasis id="B942EAC0FA72FFBFC9BFBDB1FD2AFB68" bold="true" box="[647,706,1194,1219]" italics="true" pageId="23" pageNumber="24">Vari x</emphasis>
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consists of a slightly thickened last rib within ⅓-turn of the edge of the outer lip.
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<emphasis id="B942EAC0FA72FFBFCA73BDD5FDECFB4D" bold="true" box="[331,516,1230,1255]" italics="true" pageId="23" pageNumber="24">Spiral sculpture</emphasis>
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of very fine, weak incised lines over the entire teleoconch.
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<emphasis id="B942EAC0FA72FFBFCF98BDD5FB06FB4D" bold="true" box="[1184,1262,1230,1255]" italics="true" pageId="23" pageNumber="24">Sulcus</emphasis>
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absent, there is little change in the ribs in the sulcal area.
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<emphasis id="B942EAC0FA72FFBFC957BDE9FD3FFAA1" bold="true" box="[623,727,1266,1291]" italics="true" pageId="23" pageNumber="24">Outer lip</emphasis>
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thin, and forms a continuous low arc to the end of the anterior canal. No evidence of a stromboid notch present.
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<emphasis id="B942EAC0FA72FFBFC9F8BC0DFCD4FA85" bold="true" box="[704,828,1302,1327]" italics="true" pageId="23" pageNumber="24">Anal sinus</emphasis>
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a wide but shallow indentation of outer lip edge when shell is viewed laterally; an inverted V-shape at suture when shell viewed ventrally; callus present on both the parietal wall and inside of outer lip of sinus.
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<emphasis id="B942EAC0FA72FFBFC9B4BC44FD1BFADD" bold="true" box="[652,755,1374,1399]" italics="true" pageId="23" pageNumber="24">Inner lip</emphasis>
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recumbent, margined, broader along columella, narrower on parietal wall.
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<emphasis id="B942EAC0FA72FFBFCA0BBC99FE36FA31" bold="true" box="[307,478,1410,1435]" italics="true" pageId="23" pageNumber="24">Anterior canal</emphasis>
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short, open, curved to the left viewed ventrally; end with a shallow notch. Anterior fasciole not swollen.
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<emphasis id="B942EAC0FA72FFBFCABDBCBDFE2EFA15" bold="true" box="[389,454,1446,1471]" italics="true" pageId="23" pageNumber="24">Color</emphasis>
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off white with apex and intercostal space a light creamy brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA72FFBFCBFFBCD0FB6EF8E1" pageId="23" pageNumber="24" type="discussion">
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<paragraph id="8B8936D2FA72FFBFCBFFBCD0FB6EF8E1" blockId="23.[151,1437,834,1939]" pageId="23" pageNumber="24">
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<emphasis id="B942EAC0FA72FFBFCBFFBCD0FE2BFA48" bold="true" box="[199,451,1483,1506]" pageId="23" pageNumber="24">
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Remarks.
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<emphasis id="B942EAC0FA72FFBFCA7DBCD0FE2BFA48" box="[325,451,1483,1506]" italics="true" pageId="23" pageNumber="24">Taxonomy.</emphasis>
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</emphasis>
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<taxonomicName id="4C364D51FA72FFBFCAF6BCD1FD37FA49" box="[462,735,1482,1507]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="amplicostata">Bellaspira amplicostata</taxonomicName>
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exhibits the characteristics of the genus with axial ribs that extend from suture-to-suture (in this case without any change in the sulcal area), with a varix consisting of a slightly expanded rib, with an anal sinus typical of the genus, and with surface microsculpture of fine spiral incised lines. The incised lines in this species are feeble, even obsolete on some portions of the shell surface, perhaps due to wear. Also different are the heavy growth striae that appear on the sides of the ribs, but are reduced in the intercostal space. Striae are generally weaker in other species of
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<taxonomicName id="4C364D51FA72FFBFC847BF65FC1BF93D" box="[895,1011,1662,1687]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA72FFBFC847BF65FC1BF93D" box="[895,1011,1662,1687]" italics="true" pageId="23" pageNumber="24">Bellaspira</emphasis>
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</taxonomicName>
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, the spiral lines stronger.
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<emphasis id="B942EAC0FA72FFBFCE26BF65FA74F93D" bold="true" box="[1310,1436,1662,1687]" italics="true" pageId="23" pageNumber="24">Variability.</emphasis>
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The average length of 12 examined specimens is
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<quantity id="4CCE9B37FA72FFBFC9F8BFB8FCCCF910" box="[704,804,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="7.08" pageId="23" pageNumber="24" unit="mm" value="7.08">7.08 mm</quantity>
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(6.2–8.0 mm); their average width to length
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<taxonomicName id="4C364D51FA72FFBFCE24BFB9FAA7F911" box="[1308,1359,1698,1723]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="ratio">ratio</taxonomicName>
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(W/L) is 0.445. No variation in color is observed in the single
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<typeStatus id="548D8870FA72FFBFC9C5BFDCFCC5F974" box="[765,813,1735,1758]" pageId="23" pageNumber="24">type</typeStatus>
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lot of
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<specimenCount id="9D30FD5BFA72FFBFC84DBFDCFBE7F975" box="[885,1039,1734,1759]" pageId="23" pageNumber="24" type="generic">12 specimens</specimenCount>
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.
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<emphasis id="B942EAC0FA72FFBFCF21BFDDFB56F975" bold="true" box="[1049,1214,1734,1759]" italics="true" pageId="23" pageNumber="24">Identification.</emphasis>
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This shell is unique with its uniformly but narrowly spaced wide ribs. From
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<taxonomicName id="4C364D51FA72FFBFC82CBFF0FBBEF8A9" authority="Dall, 1889" authorityName="Dall" authorityYear="1889" box="[788,1110,1770,1795]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="pentagonalis">
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<emphasis id="B942EAC0FA72FFBFC82CBFF0FC2DF8A9" box="[788,965,1770,1795]" italics="true" pageId="23" pageNumber="24">B. pentagonalis</emphasis>
|
||
(Dall, 1889)
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</taxonomicName>
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it differs in having 7–8 ribs, not 5 on the penultimate and whorl, and in coloration—the darker color is between the ribs, not on rib crests. From
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<taxonomicName id="4C364D51FA72FFBFCBAFBE28FED9F8E0" box="[151,305,1843,1866]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="aurantiaca" status="sp. nov.">
|
||
<emphasis id="B942EAC0FA72FFBFCBAFBE28FED9F8E0" box="[151,305,1843,1866]" italics="true" pageId="23" pageNumber="24">B. aurantiaca</emphasis>
|
||
</taxonomicName>
|
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,
|
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<taxonomicNameLabel id="A27157BBFA72FFBFCA07BE28FE2EF8E1" box="[319,454,1842,1867]" pageId="23" pageNumber="24" rank="species">new species</taxonomicNameLabel>
|
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it differs in possessing broader ribs, and a different coloration.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C32C6559FA72FFBFCBFFBE4CFC9DF839" pageId="23" pageNumber="24" type="etymology">
|
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<paragraph id="8B8936D2FA72FFBFCBFFBE4CFC9DF839" blockId="23.[151,1437,834,1939]" pageId="23" pageNumber="24">
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<emphasis id="B942EAC0FA72FFBFCBFFBE4CFEA6F8C4" bold="true" box="[199,334,1879,1902]" pageId="23" pageNumber="24">Etymology.</emphasis>
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The Large-ribbed
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<taxonomicName id="4C364D51FA72FFBFC917BE4DFD4BF8C5" box="[559,675,1878,1903]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA72FFBFC917BE4DFD4BF8C5" box="[559,675,1878,1903]" italics="true" pageId="23" pageNumber="24">Bellaspira</emphasis>
|
||
</taxonomicName>
|
||
, from the Latin adjectives
|
||
<emphasis id="B942EAC0FA72FFBFC8D3BE4DFBD4F8C5" box="[1003,1084,1878,1903]" italics="true" pageId="23" pageNumber="24">amplus</emphasis>
|
||
(large) and
|
||
<emphasis id="B942EAC0FA72FFBFCFEBBE4CFAC6F8C4" box="[1235,1326,1879,1902]" italics="true" pageId="23" pageNumber="24">costatus</emphasis>
|
||
(ribbed). Named after an outstanding feature of this species—its large ribs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |