313 lines
37 KiB
XML
313 lines
37 KiB
XML
<document id="0CDF359B8D2C12F725596A261BA02193" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA32FFF0CBAFB983FDE6FC62" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Clathrodrillia Dall 1918" docType="treatment" docVersion="12" lastPageNumber="89" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="88" updateTime="1698184378282" updateUser="plazi">
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<mods:titleInfo id="AC7649F049E3E606D4047C6204020C89">
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<mods:title id="674143341BB2EA45A962CF9C6C862FE2">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:namePart id="15B794E8C23EB62CE0C7A82FD3C3D3E8">Fallon, Phillip J.</mods:namePart>
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<mods:title id="2583972F50901D080E3825C9495EE0F4">Zootaxa</mods:title>
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<mods:part id="91D86D09D049E0B3F19843A9B1750AFB">
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<mods:date id="D9E203EBCB6265E3B89D36D5D9695B44">2016</mods:date>
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<mods:number id="22DA633A191C0BF43A7F315383F74972">4090</mods:number>
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<mods:classification id="44A88EA78B9E6374475935FFC82B4FBD">journal article</mods:classification>
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<mods:identifier id="BDB8279A3045441619ADE5A26817AF57" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
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<treatment id="039F87C4FA32FFF0CBAFB983FDE6FC62" ID-DOI="http://doi.org/10.5281/zenodo.6076334" ID-GBIF-Taxon="117911724" ID-Zenodo-Dep="6076334" LSID="urn:lsid:plazi:treatment:039F87C4FA32FFF0CBAFB983FDE6FC62" httpUri="http://treatment.plazi.org/id/039F87C4FA32FFF0CBAFB983FDE6FC62" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88">
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<subSubSection id="C32C6559FA32FFFFCBAFB983FB6BFF5E" pageId="87" pageNumber="88" type="nomenclature">
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<paragraph id="8B8936D2FA32FFFFCBAFB983FDF1FF18" blockId="87.[151,537,151,178]" box="[151,537,151,178]" pageId="87" pageNumber="88">
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<heading id="D0C181BEFA32FFFFCBAFB983FDF1FF18" bold="true" box="[151,537,151,178]" fontSize="11" level="1" pageId="87" pageNumber="88" reason="1">
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<emphasis id="B942EAC0FA32FFFFCBAFB983FDF1FF18" bold="true" box="[151,537,151,178]" pageId="87" pageNumber="88">
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Genus
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<taxonomicName id="4C364D51FA32FFFFCBD6B98CFDF1FF18" ID-CoL="7NZQQ" authority="Dall, 1918" authorityName="Dall" authorityYear="1918" box="[238,537,151,178]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCBD6B98CFE7DFF1B" bold="true" box="[238,405,151,177]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
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Dall, 1918
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936D2FA32FFFFCBAFB9C4FB6BFF5E" blockId="87.[151,1155,222,244]" box="[151,1155,222,244]" pageId="87" pageNumber="88">
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<typeStatus id="548D8870FA32FFFFCBAFB9C4FF21FF5E" box="[151,201,223,244]" pageId="87" pageNumber="88">Type</typeStatus>
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species by original designation:
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<taxonomicName id="4C364D51FA32FFFFC92DB9C4FCC7FF5E" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[533,815,222,244]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="gibbosus">
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<emphasis id="B942EAC0FA32FFFFC92DB9C4FD5FFF5E" box="[533,695,222,244]" italics="true" pageId="87" pageNumber="88">Murex gibbosus</emphasis>
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Born, 1778
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</taxonomicName>
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(Recent,
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<typeStatus id="548D8870FA32FFFFC8A9B9C4FC55FF5E" box="[913,957,223,244]" pageId="87" pageNumber="88">type</typeStatus>
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locality unknown).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA32FFFFCBAFB83AFC11FAD8" pageId="87" pageNumber="88" type="diagnosis">
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<paragraph id="8B8936D2FA32FFFFCBAFB83AFB77FC34" blockId="87.[151,1437,289,1466]" pageId="87" pageNumber="88">
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<emphasis id="B942EAC0FA32FFFFCBAFB83AFEF9FE90" bold="true" box="[151,273,289,314]" pageId="87" pageNumber="88">Diagnosis.</emphasis>
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The original description of the genus was very brief: "for the generally brown or brownish clathrate species a few of which are found in nearly every fauna, and of which
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<taxonomicName id="4C364D51FA32FFFFC89EB85EFB3EFEF7" authority="Reeve" authorityName="Reeve" box="[934,1238,325,350]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="gibbosa">
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<emphasis id="B942EAC0FA32FFFFC89EB85EFB61FEF4" box="[934,1161,325,350]" italics="true" pageId="87" pageNumber="88">Pleurotoma gibbosa</emphasis>
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Reeve
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</taxonomicName>
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may be specified as a typical example…" (Dall, 1918: 317). Not long afterwards, in his seminal work on the gastropods of the Bowden Formation of
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<collectingCountry id="F3217642FA32FFFFCAA7B896FE11FE0C" box="[415,505,397,422]" name="Jamaica" pageId="87" pageNumber="88">Jamaica</collectingCountry>
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(Miocene), Woodring based his diagnosis of
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<taxonomicName id="4C364D51FA32FFFFCF37B896FB43FE0C" box="[1039,1195,397,422]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCF37B896FB43FE0C" box="[1039,1195,397,422]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
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</taxonomicName>
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on an illustration of
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<taxonomicName id="4C364D51FA32FFFFCBAFB8AAFDC4FE60" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,556,433,458]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="gibbosa">
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<emphasis id="B942EAC0FA32FFFFCBAFB8AAFE7CFE60" box="[151,404,433,458]" italics="true" pageId="87" pageNumber="88">Clathrodrillia gibbosa</emphasis>
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(Born, 1778)
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</taxonomicName>
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in Kiener (1840: pl. 16, fig. 2), and so was naturally limited by having only a single example of the genus, which he stated “seems to be a valid genus” (Woodring, 1928: 152). His description included two characteristics, a short, wide aperture and toothed outer lip that are true of
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<taxonomicName id="4C364D51FA32FFFFCFBDB8E2FAE8FDB8" box="[1157,1280,505,530]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="gibbosa">
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<emphasis id="B942EAC0FA32FFFFCFBDB8E2FAE8FDB8" box="[1157,1280,505,530]" italics="true" pageId="87" pageNumber="88">C. gibbosa</emphasis>
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</taxonomicName>
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but not of all species included in the genus today. Subsequent workers emphasized one or more characteristics of the genus but did not specify all those that set
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<taxonomicName id="4C364D51FA32FFFFC927BB5AFD53FDF0" box="[543,699,577,602]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFC927BB5AFD53FDF0" box="[543,699,577,602]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
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</taxonomicName>
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apart from other genera in
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<taxonomicName id="4C364D51FA32FFFFCF30BB5AFB9EFDF0" box="[1032,1142,577,602]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
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, e.g.: "The hump-backed variced body whorl is considered diagnostic of this genus." (Powell, 1966: 73). Some species in the genera
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<taxonomicName id="4C364D51FA32FFFFCBAFBB91FEF9FD0B" box="[151,273,650,673]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCBAFBB91FEF9FD0B" box="[151,273,650,673]" italics="true" pageId="87" pageNumber="88">Fenimorea</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C364D51FA32FFFFCA27BB92FE4BFD08" box="[287,419,649,674]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCA27BB92FE4BFD08" box="[287,419,649,674]" italics="true" pageId="87" pageNumber="88">Splendrillia</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C364D51FA32FFFFCA8ABB92FDCDFD08" box="[434,549,649,674]" class="Gastropoda" family="Drilliidae" genus="Neodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCA8ABB92FDCDFD08" box="[434,549,649,674]" italics="true" pageId="87" pageNumber="88">Neodrillia</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C364D51FA32FFFFC95CBB92FD31FD08" box="[612,729,649,674]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFC95CBB92FD31FD08" box="[612,729,649,674]" italics="true" pageId="87" pageNumber="88">Calliclava</emphasis>
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</taxonomicName>
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also possess this
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<typeStatus id="548D8870FA32FFFFC89ABB91FC3AFD0B" box="[930,978,650,673]" pageId="87" pageNumber="88">type</typeStatus>
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of varix. Or: "Its wider, shorter aperture than in
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<taxonomicName id="4C364D51FA32FFFFCBD4BBB5FE87FD6C" box="[236,367,686,710]" class="Gastropoda" family="Pseudomelatomidae" genus="Crassispira" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCBD4BBB5FE87FD6C" box="[236,367,686,710]" italics="true" pageId="87" pageNumber="88">Crassispira</emphasis>
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</taxonomicName>
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and particularly the definite stromboid notch and shallow spiral furrow of some width leading to the notch set it apart" (Fargo, 1953: 368), who seemed to have followed Woodring’s description, but a clear mischaracterization of
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<taxonomicName id="4C364D51FA32FFFFCAA2BBEEFDDEFCA4" box="[410,566,757,782]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCAA2BBEEFDDEFCA4" box="[410,566,757,782]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
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</taxonomicName>
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, which has a longer narrower aperture than
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<taxonomicName id="4C364D51FA32FFFFCF15BBEDFB58FCA4" box="[1069,1200,758,782]" class="Gastropoda" family="Pseudomelatomidae" genus="Crassispira" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCF15BBEDFB58FCA4" box="[1069,1200,758,782]" italics="true" pageId="87" pageNumber="88">Crassispira</emphasis>
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</taxonomicName>
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. Neither the form of the varix nor of the aperture alone is sufficient to separate
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<taxonomicName id="4C364D51FA32FFFFC802BA02FC3EFC98" box="[826,982,793,818]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFC802BA02FC3EFC98" box="[826,982,793,818]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
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</taxonomicName>
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from other genera in
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<taxonomicName id="4C364D51FA32FFFFCFEBBA02FAAAFC98" box="[1235,1346,793,818]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
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. Olsson (1964: 95–96) gave a broader diagnosis of
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<taxonomicName id="4C364D51FA32FFFFC9B0BA26FD04FCFC" box="[648,748,829,854]" class="Gastropoda" family="Turridae" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="subFamily" subFamily="Drillinae">Drillinae</taxonomicName>
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[sic], based on the model of
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<emphasis id="B942EAC0FA32FFFFCF79BA25FB2CFCFC" box="[1089,1220,829,854]" italics="true" pageId="87" pageNumber="88">
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<taxonomicName id="4C364D51FA32FFFFCF79BA25FB28FCFC" box="[1089,1216,829,854]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="gibbosa">C. gibbosa</taxonomicName>
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,
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</emphasis>
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and included such disparate genera as
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<taxonomicName id="4C364D51FA32FFFFCABCBA7AFDE3FCD0" box="[388,523,865,890]" class="Gastropoda" family="Pseudomelatomidae" genus="Hindsiclava" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCABCBA7AFDE3FCD0" box="[388,523,865,890]" italics="true" pageId="87" pageNumber="88">Hindsiclava</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C364D51FA32FFFFC974BA7AFD06FCD0" box="[588,750,865,890]" class="Gastropoda" family="Pseudomelatomidae" genus="Compsodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFC974BA7AFD06FCD0" box="[588,750,865,890]" italics="true" pageId="87" pageNumber="88">Compsodrillia</emphasis>
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</taxonomicName>
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. This accommodation required the diagnosis to include a “ridge-like spiral cord at the suture”, a feature not present in the family as it is known today.
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</paragraph>
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<paragraph id="8B8936D2FA32FFFFCBFFBAB2FC11FAD8" blockId="87.[151,1437,289,1466]" pageId="87" pageNumber="88">
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The following diagnosis is a more complete diagnosis of the genus. Shell small to large (
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<quantity id="4CCE9B37FA32FFFFCF80BAB2FADEFC68" box="[1208,1334,937,962]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="6.0" metricValueMin="1.3" pageId="87" pageNumber="88" unit="mm" value="36.5" valueMax="60.0" valueMin="13.0">13–60 mm</quantity>
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), slender with a high, acutely pointed spire composed of flat to convex whorls having a concave anal sulcus below the suture. Aperture narrowly oval, only slightly wider at its widest than the anal sinus and canal at opposite ends. Anterior canal well demarcated and of short to moderate length, and a weak to strong anterior fasciole with or without a weak false umbilicus, or chink. Protoconch of 2–3 smooth whorls. Sculpture of numerous ridged axial ribs (crests narrow, not round) that extend from suture-to-suture but are greatly reduced and recurved in the sulcus giving the shell a shouldered or turreted outline in most species; ribs either straight or slightly opisthocline. Varix hump-like in most, positioned ¼- to ⅓-turn from the outer lip. Strong spiral sculpture of incised grooves that create the appearance of bands, cords or threads that override or cut the axials. Outer lip thin, flattened from varix to lip edge, strengthened by irregular axial folds, with a stromboid notch anteriorly, and anal sinus posteriorly; edge of lip may be scalloped or “toothed” by spirals, or not. Inner lip margined, recumbent or erect, usually relatively wide; with a callus on the parietal wall at the anal sinus. Anal sinus deep, U-shaped, adjacent to the suture, offset from shell axis by parietal lobe and projected laterally so that it appears spout-like.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA32FFFFCBFFBC65FDAAF98F" pageId="87" pageNumber="88" type="key">
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<paragraph id="8B8936D2FA32FFFFCBFFBC65FDADFA10" blockId="87.[151,1437,289,1466]" pageId="87" pageNumber="88">
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<emphasis id="B942EAC0FA32FFFFCBFFBC65FE5BFA3C" bold="true" box="[199,435,1405,1430]" pageId="87" pageNumber="88">Key characteristics.</emphasis>
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The presence of the following characteristics is diagnostic of
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<taxonomicName id="4C364D51FA32FFFFCF5CBC66FAE8FA3C" ID-CoL="7NZQQ" box="[1124,1280,1405,1430]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA32FFFFCF5CBC66FAE8FA3C" box="[1124,1280,1405,1430]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
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</taxonomicName>
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||
and separates the genus from all others in
|
||
<taxonomicName id="4C364D51FA32FFFFCAE9BCBAFDA9FA10" ID-CoL="7NHDF" box="[465,577,1441,1466]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="8B8936D2FA32FFFFCBAFBCF1FDAAF98F" blockId="87.[151,1436,1513,1754]" pageId="87" pageNumber="88">1. Strong spiral sculpture of grooves or cords; grooves may create the appearance of cords when close together, or broader bands when further apart;</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C6559FA32FFF0CBAFBF29FDA1FC92" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="description">
|
||
<paragraph id="8B8936D2FA32FFFFCBAFBF29FEFCF9C4" blockId="87.[151,1436,1513,1754]" pageId="87" pageNumber="88">2. Axial ribs that extend from suture-to-suture, and to the anterior fasciole on the last whorl; reduced in the sulcus;</paragraph>
|
||
<paragraph id="8B8936D2FA32FFFFCBAFBF61FBFCF938" blockId="87.[151,1436,1513,1754]" box="[151,1044,1657,1682]" pageId="87" pageNumber="88">3. Aperture narrow, only slightly wider than anal sinus and anterior canal; and</paragraph>
|
||
<paragraph id="8B8936D2FA32FFFFCBAFBF85FBB1F970" blockId="87.[151,1436,1513,1754]" pageId="87" pageNumber="88">4. Varix hump-like (low and broad, shallow water species), or narrow and high (deep water species), positioned about ¼- to ⅓-turn from the edge of the outer lip, usually with a dark color patch.</paragraph>
|
||
<paragraph id="8B8936D2FA32FFF0CBFFBE11FE05FE7A" blockId="87.[151,1436,1801,2006]" lastBlockId="88.[151,1437,151,968]" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88">
|
||
<emphasis id="B942EAC0FA32FFFFCBFFBE11FE2EF88B" bold="true" box="[199,454,1802,1825]" pageId="87" pageNumber="88">Nomenclatural notes.</emphasis>
|
||
<taxonomicName id="4C364D51FA32FFFFCAF5BE12FD81F888" box="[461,617,1801,1826]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA32FFFFCAF5BE12FD81F888" box="[461,617,1801,1826]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
is treated as a subgenus of
|
||
<taxonomicName id="4C364D51FA32FFFFC8A7BE12FB87F888" authority="Gray, 1838" authorityName="Gray" authorityYear="1838" box="[927,1135,1801,1826]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA32FFFFC8A7BE12FC03F888" box="[927,1003,1801,1826]" italics="true" pageId="87" pageNumber="88">Drillia</emphasis>
|
||
Gray, 1838
|
||
</taxonomicName>
|
||
by some workers (Abbott, 1974, Tippett, 1995, Tippett, 2006); others have dropped Dall’s name altogether (Harasewych & Moretzsohn, 2010: 573, as
|
||
<taxonomicName id="4C364D51FA32FFFFCA0BBE4AFE37F8C0" box="[307,479,1873,1898]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="gibbosa">
|
||
<emphasis id="B942EAC0FA32FFFFCA0BBE4AFE37F8C0" box="[307,479,1873,1898]" italics="true" pageId="87" pageNumber="88">Drillia gibbosa</emphasis>
|
||
</taxonomicName>
|
||
). With recent general acceptance of the elevation of
|
||
<taxonomicName id="4C364D51FA32FFFFCF15BE4AFB70F8C0" box="[1069,1176,1873,1898]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
|
||
to family level, it is no longer necessary to tie TWA
|
||
<taxonomicName id="4C364D51FA32FFFFCADFBE6EFD6BF824" box="[487,643,1909,1934]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA32FFFFCADFBE6EFD6BF824" box="[487,643,1909,1934]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
to the West African and Indo-Pacific genus
|
||
<taxonomicName id="4C364D51FA32FFFFCFBCBE6EFB38F824" box="[1156,1232,1909,1934]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA32FFFFCFBCBE6EFB38F824" box="[1156,1232,1909,1934]" italics="true" pageId="87" pageNumber="88">Drillia</emphasis>
|
||
</taxonomicName>
|
||
at the subgeneric level, given their morphological differences, but perhaps instead at the subfamily level if supported by molecular or other data in the future. While some TWA
|
||
<taxonomicName id="4C364D51FA32FFFFC9A0BEA6FCDCF87C" box="[664,820,1981,2006]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA32FFFFC9A0BEA6FCDCF87C" box="[664,820,1981,2006]" italics="true" pageId="87" pageNumber="88">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
appear to be similar to W African and Indo-Pacific
|
||
<taxonomicName id="4C364D51FA3DFFF0CBAFB98CFF0BFF1A" box="[151,227,151,176]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CBAFB98CFF0BFF1A" box="[151,227,151,176]" italics="true" pageId="88" pageNumber="89">Drillia</emphasis>
|
||
</taxonomicName>
|
||
, there are important differences that merit their separation on morphological grounds at the level of genus. The
|
||
<typeStatus id="548D8870FA3DFFF0CBF1B9A6FF11FF7E" box="[201,249,189,212]" pageId="88" pageNumber="89">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C364D51FA3DFFF0CA26B9A7FE82FF7F" box="[286,362,188,213]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CA26B9A7FE82FF7F" box="[286,362,188,213]" italics="true" pageId="88" pageNumber="89">Drillia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C364D51FA3DFFF0CA42B9A7FD20FF7F" authority="Gray, 1838" authorityName="Gray" authorityYear="1838" box="[378,712,188,213]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="species" species="umbilicata">
|
||
<emphasis id="B942EAC0FA3DFFF0CA42B9A7FDADFF7F" box="[378,581,188,213]" italics="true" pageId="88" pageNumber="89">Drillia umbilicata</emphasis>
|
||
Gray, 1838
|
||
</taxonomicName>
|
||
) has broad swollen ribs, relatively few in number, while the
|
||
<typeStatus id="548D8870FA3DFFF0CE54B9A6FA74FF7E" box="[1388,1436,189,212]" pageId="88" pageNumber="89">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C364D51FA3DFFF0CA29B9C4FE45FF52" box="[273,429,223,248]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CA29B9C4FE45FF52" box="[273,429,223,248]" italics="true" pageId="88" pageNumber="89">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
has numerous narrow ribs. The anal sulcus is on the shoulder slope of
|
||
<taxonomicName id="4C364D51FA3DFFF0CFDBB9C4FAC7FF52" box="[1251,1327,223,248]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CFDBB9C4FAC7FF52" box="[1251,1327,223,248]" italics="true" pageId="88" pageNumber="89">Drillia</emphasis>
|
||
</taxonomicName>
|
||
and only slightly impacts the ribs, but the sulcus forms a broader furrow, usually concave, just below the suture of
|
||
<taxonomicName id="4C364D51FA3DFFF0CBAFB83CFEDBFEEA" box="[151,307,295,320]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CBAFB83CFEDBFEEA" box="[151,307,295,320]" italics="true" pageId="88" pageNumber="89">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
where there is a marked reduction in the ribs.
|
||
<taxonomicName id="4C364D51FA3DFFF0C87EB83CFC7AFEEA" box="[838,914,295,320]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0C87EB83CFC7AFEEA" box="[838,914,295,320]" italics="true" pageId="88" pageNumber="89">Drillia</emphasis>
|
||
</taxonomicName>
|
||
has an anterior fasciole that is usually strong, with a false umbilicus; the fasciole of
|
||
<taxonomicName id="4C364D51FA3DFFF0C95DB857FCE9FECF" box="[613,769,332,357]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0C95DB857FCE9FECF" box="[613,769,332,357]" italics="true" pageId="88" pageNumber="89">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
is generally weak, and the false umbilicus only weakly developed at most. The spiral ornamentation consists of fine threads in
|
||
<taxonomicName id="4C364D51FA3DFFF0C8E5B874FBC1FE22" box="[989,1065,367,392]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0C8E5B874FBC1FE22" box="[989,1065,367,392]" italics="true" pageId="88" pageNumber="89">Drillia</emphasis>
|
||
</taxonomicName>
|
||
, but in
|
||
<taxonomicName id="4C364D51FA3DFFF0CFB2B874FACEFE22" box="[1162,1318,367,392]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CFB2B874FACEFE22" box="[1162,1318,367,392]" italics="true" pageId="88" pageNumber="89">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
the spiral sculpture consists of grooves that produce the appearance of cords, or wider bands. These differences merit separation at the generic level.
|
||
</paragraph>
|
||
<paragraph id="8B8936D2FA3DFFF0CBFFB8C7FCADFDCA" blockId="88.[151,1437,151,968]" pageId="88" pageNumber="89">
|
||
<emphasis id="B942EAC0FA3DFFF0CBFFB8C7FE96FE5E" bold="true" box="[199,382,476,501]" pageId="88" pageNumber="89">Similar genera.</emphasis>
|
||
<taxonomicName id="4C364D51FA3DFFF0CABDB8C7FD53FE5E" authority="Woodring, 1928" authorityName="Woodring" authorityYear="1928" box="[389,699,476,501]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CABDB8C7FDE8FE5F" box="[389,512,476,501]" italics="true" pageId="88" pageNumber="89">Agladrillia</emphasis>
|
||
Woodring, 1928
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C364D51FA3DFFF0C9C9B8C7FBE7FE5E" authority="McLean, 1971" authorityName="McLean" authorityYear="1971" box="[753,1039,476,501]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0C9C9B8C7FC8EFE5F" box="[753,870,476,501]" italics="true" pageId="88" pageNumber="89">Calliclava</emphasis>
|
||
McLean, 1971
|
||
</taxonomicName>
|
||
both have strong spiral elements in addition to numerous ribs and narrow apertures so may be confused with
|
||
<taxonomicName id="4C364D51FA3DFFF0C8C6B8E4FB72FDB2" box="[1022,1178,511,536]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0C8C6B8E4FB72FDB2" box="[1022,1178,511,536]" italics="true" pageId="88" pageNumber="89">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C364D51FA3DFFF0CF94B8E4FACFFDB2" box="[1196,1319,511,536]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CF94B8E4FACFFDB2" box="[1196,1319,511,536]" italics="true" pageId="88" pageNumber="89">Agladrillia</emphasis>
|
||
</taxonomicName>
|
||
differs in possessing a varix located ½-turn from the edge of the outer lip, and lacks ribs on the dorsal surface of the last whorl.
|
||
<taxonomicName id="4C364D51FA3DFFF0CBDEBB5CFEB3FDCA" box="[230,347,583,608]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CBDEBB5CFEB3FDCA" box="[230,347,583,608]" italics="true" pageId="88" pageNumber="89">Calliclava</emphasis>
|
||
</taxonomicName>
|
||
differs in possessing a carinate protoconch.
|
||
</paragraph>
|
||
<paragraph id="8B8936D2FA3DFFF0CBFFBB76FAAAFD45" blockId="88.[151,1437,151,968]" pageId="88" pageNumber="89">
|
||
<taxonomicName id="4C364D51FA3DFFF0CBFFBB76FE0FFD2E" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[199,487,620,645]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CBFFBB76FEA9FD2E" box="[199,321,621,644]" italics="true" pageId="88" pageNumber="89">Fenimorea</emphasis>
|
||
Bartsch, 1934
|
||
</taxonomicName>
|
||
is similar in possessing hump-like varices (in shallow water species, and high and narrow ones in deep water species), axial cords that run from suture-to-suture that are heavily modified in the sulcus, and, in some, heavy spiral grooves that resemble those of
|
||
<taxonomicName id="4C364D51FA3DFFF0C8B1BBAFFBCDFD67" box="[905,1061,692,717]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0C8B1BBAFFBCDFD67" box="[905,1061,692,717]" italics="true" pageId="88" pageNumber="89">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C364D51FA3DFFF0CF0DBBAFFB39FD67" box="[1077,1233,692,717]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CF0DBBAFFB39FD67" box="[1077,1233,692,717]" italics="true" pageId="88" pageNumber="89">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
differs in usually possessing turreted whorls, stronger spiral sculpture, and all lack the surface microsculpture of
|
||
<taxonomicName id="4C364D51FA3DFFF0CFF9BBC3FAD3FD45" box="[1217,1339,728,751]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CFF9BBC3FAD3FD45" box="[1217,1339,728,751]" italics="true" pageId="88" pageNumber="89">Fenimorea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8936D2FA3DFFF0CBFFBBE7FDA1FC92" blockId="88.[151,1437,151,968]" pageId="88" pageNumber="89">
|
||
<taxonomicName id="4C364D51FA3DFFF0CBFFBBE7FE36FCBE" authority="Bartsch, 1943" authorityName="Bartsch" authorityYear="1943" box="[199,478,764,789]" class="Gastropoda" family="Drilliidae" genus="Neodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CBFFBBE7FED2FCBF" box="[199,314,764,789]" italics="true" pageId="88" pageNumber="89">Neodrillia</emphasis>
|
||
Bartsch, 1943
|
||
</taxonomicName>
|
||
is similar but is readily distinguished by the raised coarse spiral threads on the shell surface and shells that are not turreted.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C6559FA3DFFF0CBFFBA5FFDE6FC62" pageId="88" pageNumber="89" type="distribution">
|
||
<paragraph id="8B8936D2FA3DFFF0CBFFBA5FFDE6FC62" blockId="88.[151,1437,151,968]" pageId="88" pageNumber="89">
|
||
<emphasis id="B942EAC0FA3DFFF0CBFFBA5FFE88FCF7" bold="true" box="[199,352,836,861]" pageId="88" pageNumber="89">Distribution.</emphasis>
|
||
<taxonomicName id="4C364D51FA3DFFF0CA50BA5FFDECFCF7" box="[360,516,836,861]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="88" pageNumber="89" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA3DFFF0CA50BA5FFDECFCF7" box="[360,516,836,861]" italics="true" pageId="88" pageNumber="89">Clathrodrillia</emphasis>
|
||
</taxonomicName>
|
||
is widely distributed in the TWA, though most species are found in the southern Caribbean, in both shallow and deep water (>
|
||
<quantity id="4CCE9B37FA3DFFF0C9A9BA7CFD24FC2A" box="[657,716,871,896]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="88" pageNumber="89" unit="m" value="50.0">50 m</quantity>
|
||
). Three species are found in the subtropical region of the Gulf of
|
||
<collectingCountry id="F3217642FA3DFFF0CBAFBA97FF07FC0F" box="[151,239,908,933]" name="Mexico" pageId="88" pageNumber="89">Mexico</collectingCountry>
|
||
, Florida, and the
|
||
<collectingCountry id="F3217642FA3DFFF0CA87BA97FDF3FC0F" box="[447,539,908,933]" name="Bahamas" pageId="88" pageNumber="89">Bahama</collectingCountry>
|
||
Is., but appear to be restricted to deeper water. Approximate localities of all species are illustrated in
|
||
<figureCitation id="130D2A57FA3DFFF0CA94BAB4FDE2FC62" box="[428,522,943,968]" captionStart="PLATE 50" captionStartId="112.[151,234,1744,1766]" captionTargetBox="[268,1296,195,1707]" captionTargetId="figure@112.[268,1296,195,1707]" captionTargetPageId="112" captionText="PLATE 50. Clathrodrillia spp. map & lineup. Fig. 1. Map of approximate known localities. Figs. 2 – 7: 10 mm scale bar. Fig. 2. Holotype, 31.0 x 11.3 mm, C. rubrofasciata, new species (MZSP 122061). Fig. 3. Holotype, 27.0 x 9.6 mm, C. garciai, new species (MNHN IM- 2013 - 55984). Fig. 4. Holotype, 16.6 x 6.0 mm, C. wolfei (Tippett, 1995) (USNM 880070). Fig. 5. Holotype, 14.7 x 5.8 mm, C. guadeloupensis, new species (MNHN IM- 2013 - 7812). Fig. 6. Lectotype, 12.6 x 4.7 mm, C. tryoni (Dall, 1889) (USNM 87449). Fig. 7. Holotype, 10.5 x 4.0 mm, C. parva, new species (USNM 1291333). Figs. 8 – 12: 20 mm scale bar: Fig. 8. Non-type, 48.9 x 17.3 mm C. gibbosa (Born, 1778) (author’s coll.). Fig. 9. Holotype, 44.4 x 17.9 mm, C. colombiana, new species (UF 470280). Fig. 10. Holotype, 39.2 x 15.8 mm, C. petuchi (Tippett, 1995) (USNM 880071). Fig. 11. Holotype, 37.1 x 13.7 mm, C. marissae, new species (ANSP 369004). Fig. 12. Non-type, 32.2 x 12.3 mm, C. dautzenbergi (Tippett, 1995) (author’s coll.)." httpUri="https://zenodo.org/record/263350/files/figure.png" pageId="88" pageNumber="89">Plate 50</figureCitation>
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</treatment>
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