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<document id="EFB53231124D83E912B5B2FDFB1E91C3" ID="10.11646/zootaxa.4139.1.11" ID-DOI="10.11646/zootaxa.4139.1.11" ID-GBIF-Dataset="4cea37f6-aa68-4c87-b4fa-09f010e6be90" ID-ISSN="1175-5326" ID-Zenodo-Dep="263488" ID-ZooBank="72E28441-7726-4F68-A874-C8EED2DC6174" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468910859359" checkinUser="plazi" docAuthor="Araya, Juan Francisco, Catalán, Ricardo &amp; Aliaga, Juan Antonio" docDate="2016" docId="039F87AAFFE8FFCAFF0BFF09D940FE84" docLanguage="en" docName="zootaxa.4139.1.11.pdf" docOrigin="Zootaxa 4139 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Astyris atacamensis Araya, Catalán &amp; Aliaga, 2016, new species" docType="treatment" docUuid="A12E0BE2-377D-4A1D-A42E-0EFA4157919E" docUuidSource="ZooBank" docVersion="8" masterDocId="FFA6FFD2FFE9FFC9FF9CFF9EDD50FFBD" masterDocTitle="A new deep-water Astyris species (Buccinoidea: Columbellidae) from the southeastern Pacific" updateTime="1698365291106" updateUser="plazi">
<mods:mods id="E499826127A962F19691EB68E7963D53" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9450819AD0BC8769A8EC8DCB854871D6">
<mods:title id="8029A3922C377796A301AD56C5627CE2">A new deep-water Astyris species (Buccinoidea: Columbellidae) from the southeastern Pacific</mods:title>
</mods:titleInfo>
<mods:name id="CE30CEC631D195E268CD61777CAA0B05" type="personal">
<mods:role id="553CA189AF3F9F83D10F80F17F53295C">
<mods:roleTerm id="66056B7720BA4F44708673DFD61D82D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC04988BDF51B1F2D4C73B10A1A17C85">Araya, Juan Francisco</mods:namePart>
</mods:name>
<mods:name id="5E04FF3E5E98282CAC40833933FAE765" type="personal">
<mods:role id="3BAF7F661A83079CB24E1AE4042D16F8">
<mods:roleTerm id="8290A3CD656C4BD277706A1162064FCD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A3484A37D9ACD5247029AFE3C44AA32">Catalán, Ricardo</mods:namePart>
</mods:name>
<mods:name id="B479FD4F30C6038A7D05FB88DE4EB80D" type="personal">
<mods:role id="84386441DB716CB797EABF0D45D276FD">
<mods:roleTerm id="79C657C4C11A54FA335DB0D993F75A25">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA0CDEF5918BDC21E27F976729128142">Aliaga, Juan Antonio</mods:namePart>
</mods:name>
<mods:typeOfResource id="A22112606ED9C7E6911E01212D0584F0">text</mods:typeOfResource>
<mods:relatedItem id="EEB8BBE7EEDDE69FACC7F351A1AC9A3D" type="host">
<mods:titleInfo id="0BC5E1EE622C84ABD7D47C6EA01431EE">
<mods:title id="461DFDFC21D3BD1C51C27B0F0C48A218">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D611B33BC4743C559171030DDF05DAEE">
<mods:date id="9F2D359CE27961B1C72EAAB81F95A1E7">2016</mods:date>
<mods:detail id="8EA7367B80738B611D91B21FCBA0509D" type="volume">
<mods:number id="873018034DE9BE13999D66CA35C49F08">4139</mods:number>
</mods:detail>
<mods:detail id="261D5CC0A62BC81A6BA45E72D72992F8" type="issue">
<mods:number id="7328C50FA7E176225BE223872B7948EA">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="B9465066C7756DA4BEDAB69EF1E8ECA4">journal volume</mods:classification>
<mods:identifier id="5C3E6498F793C7AB06CED810192DE233" type="DOI">10.11646/zootaxa.4139.1.11</mods:identifier>
<mods:identifier id="7C0EB193E24A94A498F46564AFA7D22B" type="GBIF-Dataset">4cea37f6-aa68-4c87-b4fa-09f010e6be90</mods:identifier>
<mods:identifier id="9C480449B4A6083C0641364D6C050653" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="52EB1FABADB92A8D28DBCD56C6264618" type="Zenodo-Dep">263488</mods:identifier>
<mods:identifier id="3769DEFCF96E9224F095E08D231E2C6E" type="ZooBank">72E28441-7726-4F68-A874-C8EED2DC6174</mods:identifier>
</mods:mods>
<treatment id="039F87AAFFE8FFCAFF0BFF09D940FE84" ID-DOI="http://doi.org/10.5281/zenodo.6085404" ID-GBIF-Taxon="123198383" ID-Zenodo-Dep="6085404" LSID="urn:lsid:zoobank.org:act:A12E0BE2-377D-4A1D-A42E-0EFA4157919E" httpUri="http://treatment.plazi.org/id/039F87AAFFE8FFCAFF0BFF09D940FE84" lastPageId="3" pageId="1">
<subSubSection id="C32C6537FFE8FFC8FF0BFF09DD86FF6C" pageId="1" type="nomenclature">
<paragraph id="8B8936BCFFE8FFC8FF0BFF09DF4CFF0F" blockId="1.[151,1436,151,1028]" box="[151,540,151,178]" pageId="1">
<heading id="D0C181D0FFE8FFC8FF0BFF09DF4CFF0F" bold="true" box="[151,540,151,178]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="B942EAAEFFE8FFC8FF0BFF09DF4CFF0F" bold="true" box="[151,540,151,178]" pageId="1">
<taxonomicName id="4C364D3FFFE8FFC8FF0BFF09DCD4FF0C" ID-CoL="687FN" box="[151,388,151,177]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="1" phylum="Mollusca" rank="species" species="atacamensis" status="sp. nov.">
<emphasis id="B942EAAEFFE8FFC8FF0BFF09DCD4FF0C" bold="true" box="[151,388,151,177]" italics="true" pageId="1">Astyris atacamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157D5FFE8FFC8FE10FF06DF4CFF0F" box="[396,540,152,178]" pageId="1" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936BCFFE8FFC8FF0BFF24DD86FF6C" blockId="1.[151,1436,151,1028]" box="[151,214,186,209]" pageId="1">
<figureCitation id="130D2A39FFE8FFC8FF0BFF24DD86FF6C" box="[151,214,186,209]" captionStart="FIGURE 1" captionStartId="1.[151,244,1980,2002]" captionTargetBox="[151,1436,1071,1956]" captionTargetId="figure@1.[151,1436,1071,1956]" captionTargetPageId="1" captionText="FIGURE 1. Astyris atacamensis n. sp., off Huasco, Chile, 404 409 m depth. A E. Holotype SBMNH 456358. F I. Paratype SBMNH 452239." httpUri="https://zenodo.org/record/263489/files/figure.png" pageId="1">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32C6537FFE8FFC8FF0BFF43DF15FEC2" pageId="1" type="materials_examined">
<paragraph id="8B8936BCFFE8FFC8FF0BFF43DC18FE84" blockId="1.[151,1436,151,1028]" pageId="1">
<emphasis id="B942EAAEFFE8FFC8FF0BFF43DC13FF49" bold="true" box="[151,323,221,244]" pageId="1">
<typeStatus id="548D881EFFE8FFC8FF0BFF43DD9FFF49" box="[151,207,221,244]" pageId="1">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548D881EFFE8FFC8FECCFF43DCE7FF49" box="[336,439,221,244]" pageId="1" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED27AE79FFE8FFC8FE54FF43DF77FF49" box="[456,551,221,244]" country="USA" httpUri="http://grbio.org/cool/1vha-esv7" name="Santa Barbara Natural History Museum" pageId="1">SBNHM</collectionCode>
460092 (H: 10.6, D: 6.7 and LW:
<quantity id="4CCE9B59FFE8FFC8FC57FF43D978FF49" box="[971,1064,221,244]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="1" unit="mm" value="8.3">8.3 mm</quantity>
);
<typeStatus id="548D881EFFE8FFC8FBDDFF43D9FBFF49" box="[1089,1195,221,244]" pageId="1" type="paratype">paratypes</typeStatus>
:
<collectionCode id="ED27AE79FFE8FFC8FB20FF43D84BFF49" box="[1212,1307,221,244]" country="USA" httpUri="http://grbio.org/cool/1vha-esv7" name="Santa Barbara Natural History Museum" pageId="1">SBNHM</collectionCode>
460093 (
<specimenCount id="9D30FD35FFE8FFC8FA0CFF43DC50FEAA" pageId="1" type="generic">1 specimen</specimenCount>
),
<collectionCode id="ED27AE79FFE8FFC8FE8FFE9EDC3EFEAA" box="[275,366,256,279]" pageId="1">MPCCL</collectionCode>
0 1062016 (
<specimenCount id="9D30FD35FFE8FFC8FE73FE9EDF28FEAA" box="[495,632,256,279]" pageId="1" type="generic">4 specimens</specimenCount>
),
<collectionCode id="ED27AE79FFE8FFC8FD17FE9EDF88FEAA" box="[651,728,256,279]" country="Italy" httpUri="http://grbio.org/cool/70mh-t5r1" name="Universita di Cagliari" pageId="1">MZUC</collectionCode>
37641 (
<specimenCount id="9D30FD35FFE8FFC8FCAEFE9EDEEBFEAA" box="[818,955,256,279]" pageId="1" type="generic">4 specimens</specimenCount>
), MNHNCL 202493 (
<specimenCount id="9D30FD35FFE8FFC8FB35FE9ED862FEAA" box="[1193,1330,256,279]" pageId="1" type="generic">7 specimens</specimenCount>
); all from the
<typeStatus id="548D881EFFE8FFC8FF22FEBCDDBCFE84" box="[190,236,290,313]" pageId="1">type</typeStatus>
locality.
</paragraph>
<paragraph id="8B8936BCFFE8FFC8FF5BFEDBDF15FEC2" blockId="1.[151,1436,151,1028]" pageId="1">
<emphasis id="B942EAAEFFE8FFC8FF5BFEDBDC0EFEE1" bold="true" box="[199,350,325,348]" pageId="1">
<typeStatus id="548D881EFFE8FFC8FF5BFEDBDDAFFEE1" box="[199,255,325,348]" pageId="1">Type</typeStatus>
locality.
</emphasis>
<quantity id="4CCE9B59FFE8FFC8FEF9FEDBDCF8FEE1" box="[357,424,325,348]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="1" unit="km" value="20.0">20 km</quantity>
NW off Huasco (
<geoCoordinate id="EE02507BFFE8FFC8FDFBFEDBDFB9FEE1" box="[615,745,325,348]" direction="south" orientation="latitude" pageId="1" precision="15" value="-28.259167">28°14'93'' S</geoCoordinate>
;
<geoCoordinate id="EE02507BFFE8FFC8FD69FEDBDE2FFEE1" box="[757,895,325,348]" direction="west" orientation="longitude" pageId="1" precision="15" value="-71.29722">71°17'50'' W</geoCoordinate>
,
<quantity id="4CCE9B59FFE8FFC8FC16FEDBD956FEE1" box="[906,1030,325,348]" metricMagnitude="2" metricUnit="m" metricValue="4.0649999999999995" metricValueMax="4.09" metricValueMin="4.04" pageId="1" unit="m" value="406.5" valueMax="409.0" valueMin="404.0">404409 m</quantity>
),
<collectingRegion id="49F2F85EFFE8FFC8FB85FEDBD9BFFEE1" box="[1049,1263,325,348]" country="Chile" name="Atacama" pageId="1">Región de Atacama</collectingRegion>
, northern
<collectingCountry id="F321762CFFE8FFC8FAC7FEDBD8C8FEE1" box="[1371,1432,325,348]" name="Chile" pageId="1">Chile</collectingCountry>
. Leg. Ricardo Catalán,
<date id="FF88107CFFE8FFC8FE1BFEF6DF12FEC2" box="[391,578,360,383]" pageId="1" value="2000-02-03">February 3, 2000</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C6537FFE8FFC8FF5BFE14DF10FE79" pageId="1" type="etymology">
<paragraph id="8B8936BCFFE8FFC8FF5BFE14DF10FE79" blockId="1.[151,1436,151,1028]" pageId="1">
<emphasis id="B942EAAEFFE8FFC8FF5BFE14DC17FE1C" bold="true" box="[199,327,394,417]" pageId="1">Etymology.</emphasis>
The specific name of
<taxonomicName id="4C364D3FFFE8FFC8FDA7FE14DE40FE1C" box="[571,784,394,417]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="1" phylum="Mollusca" rank="species" species="atacamensis">
<emphasis id="B942EAAEFFE8FFC8FDA7FE14DE40FE1C" box="[571,784,394,417]" italics="true" pageId="1">Astyris atacamensis</emphasis>
</taxonomicName>
refers to presence of this species on the continental shelf of the
<collectingRegion id="49F2F85EFFE8FFC8FF22FE33DCC4FE79" box="[190,404,429,452]" country="Chile" name="Atacama" pageId="1">Región de Atacama</collectingRegion>
, northern
<collectingCountry id="F321762CFFE8FFC8FE62FE33DF6BFE79" box="[510,571,429,452]" name="Chile" pageId="1">Chile</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C6537FFE8FFC8FF5BFE4EDDBFFDF2" pageId="1" type="diagnosis">
<paragraph id="8B8936BCFFE8FFC8FF5BFE4EDDBFFDF2" blockId="1.[151,1436,151,1028]" pageId="1">
<emphasis id="B942EAAEFFE8FFC8FF5BFE4EDC6CFE5A" bold="true" box="[199,316,464,487]" pageId="1">Diagnosis.</emphasis>
Shell small (up to
<quantity id="4CCE9B59FFE8FFC8FD96FE4EDF39FE5A" box="[522,617,464,487]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="1" unit="mm" value="11.1">11.1 mm</quantity>
long), thick, stout, with 4 to 5 rounded and slightly angulated whorls, often decollate; with sculpture of fine growth lines; spiral sculpture of very fine spiral grooves most noticeable on base of shell. Aperture wide, oval; outer lip sharp, in mature specimens with minute internal lirae; siphonal canal broad and straight.
</paragraph>
</subSubSection>
<subSubSection id="C32C6537FFE8FFC8FF5BFDC4D984FBB9" pageId="1" type="description">
<paragraph id="8B8936BCFFE8FFC8FF5BFDC4D984FBB9" blockId="1.[151,1436,151,1028]" pageId="1">
<emphasis id="B942EAAEFFE8FFC8FF5BFDC4DC02FDCC" bold="true" box="[199,338,602,625]" pageId="1">Description.</emphasis>
Shell small (H
<quantity id="4CCE9B59FFE8FFC8FE67FDC4DFF2FDCC" box="[507,674,602,625]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" metricValueMax="11.1" metricValueMin="5.8" pageId="1" unit="mm" value="8.45" valueMax="11.1" valueMin="5.8">5.8 to 11.1 mm</quantity>
), thick, stout, with 4 to 5 weakly convex whorls; outline conic; sutures impressed. Measurements from fifty randomly chosen specimens: H 5.88.3
<quantity id="4CCE9B59FFE8FFC8FC4DFDE2D960FD2E" box="[977,1072,636,659]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="1" unit="mm" value="11.1">11.1 mm</quantity>
, D 3.55.3
<quantity id="4CCE9B59FFE8FFC8FB2EFDE2D855FD2E" box="[1202,1285,636,659]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" unit="mm" value="7.1">7.1 mm</quantity>
, and height of last whorl 4.56.5
<quantity id="4CCE9B59FFE8FFC8FEF1FD03DC91FD09" box="[365,449,669,692]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="1" unit="mm" value="8.7">8.7 mm</quantity>
(minimumaveragemaximum). Taking into account that most of the shells had the first whorls eroded, we could observe at most 5 whorls in the better preserved shells, with a bit over 4 whorls remaining in most specimens. Colour pale tan; earlier whorls slightly paler, showing indistinct spiral bands. Spire short, decollate, about two-fifths of total height; spire angle a bit less than 45°. Protoconch and early whorls eroded. Teleoconch smooth and slightly glossy, except for fine, irregular axial growth lines crossed by spiral sculpture of very fine, barely visible grooves; grooves cover almost entire exterior surface, coarsest and most noticeable on base. Last whorl large, about three-fourths of shell height, slightly convex, angulated near the base, concave below; maximum width at about one third from anterior end. Aperture prosocline, oval, slightly rectangular, posteriorly acuminate; aperture height slightly more than half of shell height. Parietal callus slightly thickened, concave. Outer lip sharp, simple, outline weakly convex, weakly concave over upper third, then swinging forwards weakly to suture; about 8 to 12 lirae inside outer lip; columella without denticles, callous. Siphonal canal short, very wide, anteriorly with rounded siphonal aperture.
</paragraph>
</subSubSection>
<caption id="DF496634FFE8FFC8FF0BF822DC11F852" httpUri="https://zenodo.org/record/263489/files/figure.png" pageId="1" targetBox="[151,1436,1071,1956]" targetPageId="1">
<paragraph id="8B8936BCFFE8FFC8FF0BF822DC11F852" blockId="1.[151,1436,1980,2031]" pageId="1">
<emphasis id="B942EAAEFFE8FFC8FF0BF822DC41F86F" bold="true" box="[151,273,1980,2002]" pageId="1">FIGURE 1.</emphasis>
<taxonomicName id="4C364D3FFFE8FFC8FE87F823DC8EF86F" box="[283,478,1981,2002]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="1" phylum="Mollusca" rank="species" species="atacamensis" status="sp. nov.">
<emphasis id="B942EAAEFFE8FFC8FE87F823DC8EF86F" box="[283,478,1981,2002]" italics="true" pageId="1">Astyris atacamensis</emphasis>
</taxonomicName>
<emphasis id="B942EAAEFFE8FFC8FE75F822DF74F86F" bold="true" box="[489,548,1980,2002]" pageId="1">
<taxonomicNameLabel id="A27157D5FFE8FFC8FE75F822DF74F86F" box="[489,548,1980,2002]" pageId="1" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, off Huasco, Chile, 404409 m depth.
<emphasis id="B942EAAEFFE8FFC8FC25F822DEBDF86F" bold="true" box="[953,1005,1980,2002]" pageId="1">AE.</emphasis>
Holotype SBMNH 456358.
<emphasis id="B942EAAEFFE8FFC8FA88F822D86DF86F" bold="true" box="[1300,1341,1980,2002]" pageId="1">FI.</emphasis>
Paratype SBMNH 452239.
</paragraph>
</caption>
<caption id="DF496634FFEBFFCBFF0BFB67D951FA96" httpUri="https://zenodo.org/record/263490/files/figure.png" pageId="2" targetBox="[150,1436,191,1251]" targetPageId="2">
<paragraph id="8B8936BCFFEBFFCBFF0BFB67D951FA96" blockId="2.[151,1436,1273,1323]" pageId="2">
<emphasis id="B942EAAEFFEBFFCBFF0BFB67DC5EFAB2" bold="true" box="[151,270,1273,1295]" pageId="2">FIGURE 2.</emphasis>
<taxonomicName id="4C364D3FFFEBFFCBFE8AFB67DF02FAB3" authority="Dall, 1890" authorityName="Dall" authorityYear="1890" box="[278,594,1273,1294]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFE8AFB67DC80FAB3" box="[278,464,1273,1294]" italics="true" pageId="2">Astyris permodesta</emphasis>
(Dall, 1890)
</taxonomicName>
. AE. Holotype, USNM 96526, north of Santa Rosa Isl., Channel Islands, California, USA, 276 fms [504.7 m], mud. FG. SBMNH 356555, Santa Monica Basin, off California.
</paragraph>
</caption>
<subSubSection id="C32C6537FFEBFFCBFF5BFAC9DEC2FA45" pageId="2" type="distribution">
<paragraph id="8B8936BCFFEBFFCBFF5BFAC9DEC2FA45" blockId="2.[151,1436,1367,2014]" pageId="2">
<emphasis id="B942EAAEFFEBFFCBFF5BFAC9DCB3FAD3" bold="true" box="[199,483,1367,1390]" pageId="2">Distribution and habitat.</emphasis>
Shells of this species (no live specimens) were found in greenish mud, from soft bottoms off Puerto Flamenco (
<geoCoordinate id="EE02507BFFEBFFCBFE1EFAE7DF54FA2D" box="[386,516,1401,1424]" direction="south" orientation="latitude" pageId="2" precision="15" value="-28.259167">28°14'93'' S</geoCoordinate>
;
<geoCoordinate id="EE02507BFFEBFFCBFD8DFAE7DFCDFA2D" box="[529,669,1401,1424]" direction="west" orientation="longitude" pageId="2" precision="15" value="-71.29722">71°17'50'' W</geoCoordinate>
),
<collectingRegion id="49F2F85EFFEBFFCBFD2DFAE7DED5FA2D" box="[689,901,1401,1424]" country="Chile" name="Atacama" pageId="2">Región de Atacama</collectingRegion>
to off Tongoy (
<geoCoordinate id="EE02507BFFEBFFCBFBB3FAE7D9FAFA2D" box="[1071,1194,1401,1424]" direction="south" orientation="latitude" pageId="2" precision="15" value="-30.193333">30°11'36''S</geoCoordinate>
;
<geoCoordinate id="EE02507BFFEBFFCBFB29FAE7D86AFA2D" box="[1205,1338,1401,1424]" direction="west" orientation="longitude" pageId="2" precision="15" value="-71.556946">71°33'25''W</geoCoordinate>
),
<collectingRegion id="49F2F85EFFEBFFCBFAD2FAE7DC7BFA0E" country="Chile" name="Coquimbo" pageId="2">Región de Coquimbo</collectingRegion>
,
<collectingCountry id="F321762CFFEBFFCBFEA4FA02DC24FA0E" box="[312,372,1436,1459]" name="Chile" pageId="2">Chile</collectingCountry>
, from depths of
<quantity id="4CCE9B59FFEBFFCBFDB4FA02DFEEFA0E" box="[552,702,1436,1459]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="4.7" metricValueMin="1.8" pageId="2" unit="m" value="325.0" valueMax="470.0" valueMin="180.0">
180 to
<specimenCount id="9D30FD35FFEBFFCBFDEAFA02DFEEFA0E" box="[630,702,1436,1459]" pageId="2" type="generic">470 m</specimenCount>
</quantity>
. A single specimen, recently collected from sediments of the Bahía Inglesa Formation (Miocene to Pliocene) at Quebrada Blanca (
<geoCoordinate id="EE02507BFFEBFFCBFCD7FA21DE86FA6B" box="[843,982,1471,1494]" direction="south" orientation="latitude" pageId="2" precision="15" value="-27.065277">27°0355” S</geoCoordinate>
;
<geoCoordinate id="EE02507BFFEBFFCBFC79FA21D92AFA6B" box="[997,1146,1471,1494]" direction="west" orientation="longitude" pageId="2" precision="15" value="-70.791664">70°4730” W</geoCoordinate>
), near Caldera,
<collectingCountry id="F321762CFFEBFFCBFAB6FA21D834FA6B" box="[1322,1380,1471,1494]" name="Chile" pageId="2">Chile</collectingCountry>
may represent the oldest record for this species (Nielsen et al., unpublished).
</paragraph>
</subSubSection>
<subSubSection id="C32C6537FFEBFFCAFF5BF99AD940FE84" lastPageId="3" pageId="2" type="discussion">
<paragraph id="8B8936BCFFEBFFCAFF5BF99ADD8CFF6C" blockId="2.[151,1436,1367,2014]" lastBlockId="3.[151,1436,151,313]" lastPageId="3" pageId="2">
<emphasis id="B942EAAEFFEBFFCBFF5BF99ADC65F9A6" bold="true" box="[199,309,1540,1563]" pageId="2">Remarks.</emphasis>
All of the specimens examined have the apex eroded to varying degrees, all of them lacking the protoconch and part of the earlier whorls. The definite generic assignment is thus still preliminary, but they agree partially with the description given by
<bibRefCitation id="EFA74B4DFFEBFFCBFDAAF9D7DF9AF9DD" author="Garcia" box="[566,714,1609,1632]" pageId="2" refString="Garcia, E. F. (2009) A new Astyris species (Gastropoda: Columbellidae) from the Gulf of Mexico, with notes on the genus. Novapex, 10 (1), 5 - 8" type="journal article" year="2009">Garcia (2009)</bibRefCitation>
for
<taxonomicName id="4C364D3FFFEBFFCBFD64F9D7DE12F9DD" box="[760,834,1609,1632]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="genus">
<emphasis id="B942EAAEFFEBFFCBFD64F9D7DE12F9DD" box="[760,834,1609,1632]" italics="true" pageId="2">Astyris</emphasis>
</taxonomicName>
. The distinctive broad shell shape, with a wide siphonal canal, is quite distinctive and different from all the other known columbellid species from the southeastern Pacific.
<taxonomicName id="4C364D3FFFEBFFCBFF0BF910DC3BF91B" box="[151,363,1678,1702]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="atacamensis" status="sp. nov.">
<emphasis id="B942EAAEFFEBFFCBFF0BF910DC3BF91B" box="[151,363,1678,1702]" italics="true" pageId="2">Astyris atacamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157D5FFEBFFCBFEEEF911DCFBF91B" box="[370,427,1679,1702]" pageId="2" rank="species">sp. n.</taxonomicNameLabel>
is similar in shell characters only to
<taxonomicName id="4C364D3FFFEBFFCBFCAEF910D9DBF91B" authority="Dall, 1890" authorityName="Dall" authorityYear="1890" box="[818,1163,1678,1702]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFCAEF910DEAFF91B" box="[818,1023,1678,1702]" italics="true" pageId="2">Astyris permodesta</emphasis>
(
<bibRefCitation id="EFA74B4DFFEBFFCBFB92F911D9D3F91B" author="Dall" box="[1038,1155,1679,1702]" pageId="2" refString="Dall, W. H. (1890) Scientific results of explorations by steamer Albatross No. VII. Preliminary report on the collection of mollusca and brachiopoda obtained in 1887 - ' 88. Proceedings of the United States National Museum, 12, 219 - 362. http: // dx. doi. org / 10.5479 / si. 00963801.12 - 773.219" type="journal article" year="1890">Dall, 1890</bibRefCitation>
)
</taxonomicName>
, a species described from deep water in the northeastern Pacific (
<bibRefCitation id="EFA74B4DFFEBFFCBFDA4F92FDFFCF975" author="Dall" box="[568,684,1713,1736]" pageId="2" refString="Dall, W. H. (1890) Scientific results of explorations by steamer Albatross No. VII. Preliminary report on the collection of mollusca and brachiopoda obtained in 1887 - ' 88. Proceedings of the United States National Museum, 12, 219 - 362. http: // dx. doi. org / 10.5479 / si. 00963801.12 - 773.219" type="journal article" year="1890">Dall, 1890</bibRefCitation>
, Thorsson 2002). The new species differs from
<taxonomicName id="4C364D3FFFEBFFCBFB37F92FD815F975" box="[1195,1349,1712,1736]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFB37F92FD9EBF975" box="[1195,1211,1713,1736]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFB55F92ED815F975" box="[1225,1349,1712,1736]" italics="true" pageId="2">permodesta</emphasis>
</taxonomicName>
(
<figureCitation id="130D2A39FFEBFFCBFAC8F92FD8C4F975" box="[1364,1428,1713,1736]" captionStart="FIGURE 2" captionStartId="2.[151,244,1273,1295]" captionTargetBox="[150,1436,191,1251]" captionTargetId="figure@2.[150,1436,191,1251]" captionTargetPageId="2" captionText="FIGURE 2. Astyris permodesta (Dall, 1890). A E. Holotype, USNM 96526, north of Santa Rosa Isl., Channel Islands, California, USA, 276 fms [504.7 m], mud. F G. SBMNH 356555, Santa Monica Basin, off California." httpUri="https://zenodo.org/record/263490/files/figure.png" pageId="2">Fig. 2</figureCitation>
) in shell shape (slightly angulated versus convex whorls), with a last whorl slightly concave in
<taxonomicName id="4C364D3FFFEBFFCBFB58F94AD83AF956" box="[1220,1386,1748,1771]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="atacamensis">
<emphasis id="B942EAAEFFEBFFCBFB58F94AD984F956" box="[1220,1236,1748,1771]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFB7AF94AD83AF956" box="[1254,1386,1748,1771]" italics="true" pageId="2">atacamensis</emphasis>
</taxonomicName>
and compressed below in
<taxonomicName id="4C364D3FFFEBFFCBFE0DF968DF61F8B3" box="[401,561,1782,1806]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFE0DF968DCF1F8B0" box="[401,417,1782,1805]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFE29F968DF61F8B3" box="[437,561,1782,1806]" italics="true" pageId="2">permodesta</emphasis>
</taxonomicName>
(producing a differentiated/protruding siphonal canal in this species); in the thickness of shell and lip (thin in
<taxonomicName id="4C364D3FFFEBFFCBFE65F887DFC2F88D" box="[505,658,1816,1840]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFE65F887DF59F88D" box="[505,521,1817,1840]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFD8AF886DFC2F88D" box="[534,658,1816,1840]" italics="true" pageId="2">permodesta</emphasis>
</taxonomicName>
and thick in
<taxonomicName id="4C364D3FFFEBFFCBFC81F887DEEDF88D" box="[797,957,1817,1840]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="atacamensis">
<emphasis id="B942EAAEFFEBFFCBFC81F887DE7DF88D" box="[797,813,1817,1840]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFCA6F887DEEDF88D" box="[826,957,1817,1840]" italics="true" pageId="2">atacamensis</emphasis>
</taxonomicName>
; most visible at the upper end of outer lip), a slightly different aperture outline (product of the more convex last whorl in
<taxonomicName id="4C364D3FFFEBFFCBFC5CF8A2D90AF8EE" box="[960,1114,1851,1875]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFC5CF8A2DE80F8EE" box="[960,976,1852,1875]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFC42F8A5D90AF8EE" box="[990,1114,1851,1875]" italics="true" pageId="2">permodesta</emphasis>
</taxonomicName>
, slightly angulated in the new species), a thick columella (not callous in
<taxonomicName id="4C364D3FFFEBFFCBFDCBF8C0DFA0F8CB" box="[599,752,1886,1910]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFDCBF8C0DF37F8C8" box="[599,615,1886,1909]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFDE8F8C0DFA0F8CB" box="[628,752,1886,1910]" italics="true" pageId="2">permodesta</emphasis>
</taxonomicName>
), in the presence of lirae in the interior border of the outer lip in the new species (absent in
<taxonomicName id="4C364D3FFFEBFFCBFE28F81FDF1EF825" box="[436,590,1920,1944]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFE28F81FDC94F825" box="[436,452,1921,1944]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFE4EF81EDF1EF825" box="[466,590,1920,1944]" italics="true" pageId="2">permodesta</emphasis>
</taxonomicName>
) and, particularly, in having a broader and less constricted siphonal canal, with a more pronounced columellar fold in the new species (absent in
<taxonomicName id="4C364D3FFFEBFFCBFCF8F83AD950F806" box="[868,1024,1955,1979]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">
<emphasis id="B942EAAEFFEBFFCBFCF8F83ADE24F806" box="[868,884,1956,1979]" italics="true" pageId="2">A</emphasis>
.
<emphasis id="B942EAAEFFEBFFCBFC18F83DD950F806" box="[900,1024,1955,1979]" italics="true" pageId="2">permodesta</emphasis>
</taxonomicName>
). Furthermore, the habitat of the new species also seems to differ from that of
<emphasis id="B942EAAEFFEBFFCBFDD8F858DE48F863" box="[580,792,1990,2014]" italics="true" pageId="2">
<taxonomicName id="4C364D3FFFEBFFCBFDD8F858DE43F863" box="[580,787,1990,2014]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" phylum="Mollusca" rank="species" species="permodesta">Astyris permodesta</taxonomicName>
,
</emphasis>
which is unusual in that it has been described from reducing environments, recorded along with species of large vesicomyid bivalves of the genus
<taxonomicName id="4C364D3FFFEAFFCAFBB0FF09DD88FF6C" authority="Dall, 1891 (McLean, 1996)" authorityName="Dall, 1891 (McLean" authorityYear="1996" class="Bivalvia" family="Vesicomyidae" genus="Calyptogena" kingdom="Animalia" order="Veneroida" pageId="3" phylum="Mollusca" rank="genus">
<emphasis id="B942EAAEFFEAFFCAFBB0FF09D9E4FF12" box="[1068,1204,151,175]" italics="true" pageId="3">Calyptogena</emphasis>
Dall, 1891 (
<bibRefCitation id="EFA74B4DFFEAFFCAFAA7FF06DD81FF6C" author="McLean" pageId="3" refString="McLean, J. H., Gosliner, T. M., Scott, P. H., Blake, J. A. &amp; Lissner, A. L. (1996) Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 9. The Mollusca. Part 2: The Gastropoda. Santa Barbara Museum of Natural History, Santa Barbara, 219 pp." type="book" year="1996">McLean, 1996</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8936BCFFEAFFCAFF5BFF43D940FE84" blockId="3.[151,1436,151,313]" pageId="3">
This new species thus represents the first record in
<collectingCountry id="F321762CFFEAFFCAFC8AFF43DE00FF49" box="[790,848,221,244]" name="Chile" pageId="3">Chile</collectingCountry>
of the genus
<taxonomicName id="4C364D3FFFEAFFCAFC68FF43D96EFF49" box="[1012,1086,221,244]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="3" phylum="Mollusca" rank="genus">
<emphasis id="B942EAAEFFEAFFCAFC68FF43D96EFF49" box="[1012,1086,221,244]" italics="true" pageId="3">Astyris</emphasis>
</taxonomicName>
, previously unknown from the southeastern Pacific. Further sediment sampling in the area will probably reveal more molluscan species to be discovered and described, particularly micromollusks, as described by
<bibRefCitation id="EFA74B4DFFEAFFCAFC93FEBCD95CFE84" author="Araya" box="[783,1036,290,313]" pageId="3" refString="Araya, J. F. &amp; Geiger, D. L. (2013) Cornisepta guzmani n. sp.: first species of genus confirmed from Pacific cold seep environments off central Chile (Gastropoda: Vetigastropoda: Fissurellidae). Nautilus, 127, 115 - 118." type="journal article" year="2013">Araya &amp; Geiger (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>